,*(,**2) !"#$%&.' ()*+, - . /,01,23 4526 7 89 :;<
=>=?<4@AB<C.D%E FG.HI=JKLM
=NOPQR=#STUVWEX R=YZ[\]
EM^.%7 _`ab=cdef4gh iPQ=89=jdkRlmno4pq r6js 89tuvw =x4yQk6%7
=zZ {|$}~
zZ ! " "#$ %&
'()* ! +, (- . /0123456748!"(
9:;34 (1/(<=>?@
3A=B C34 "(
3"DEFG(=HI
ῌai JKLM1NK(YM
@=OPQIRSPQITUI(VB (WX Y.Z[ \] ^? _`a.=
῍9?= bcdF*./ Kefg*./
hij:g*./ k;hlmno*.+`!p q (/*ml) Rrstgju vwdxGy:
vwz k{ |}q+ml3ey~:i-./ml
,/+
῎9 B +l, p +l, k{ *..l, /? ,/+
4=
῏9 1(@3 vij:i 8
?T+0.]4vij:i. B-4 +(
,1+4=
ῐ? k0!]B+4!
1( 4vij:i,!a
,1+ +3k
vij:i4k
ῑ89tuvw :cimc F =
"p a"#4DEFG = 1(@="3DEFG/
. H2a.$¡
~k ῌ =B(vij:i
+0 %+0 ¢ ῍((=
q£@(q&¤.'¥((8).
k¢¦8*8,3 [,3§+>
? ῎(!(= ¤.'¥()>
¨¢ vij:i.©!,3a.
k{,ª#«>¬!,3[ k.
®-"(+?(0¯¢ ῏ k,!] vij:i°4 ±²®-"3.³k0¯`[
ῐ 31(@/¢"/
84"DEFG = +(´* 0µ(
¶1=" =DEFGp*.+l 2. :cimc·>³,3 F·>³ +/,3 ="·>³+3 ±T¸8$¡4 ¹ B"/¢]jgd¸¢
º» <5¼ +?C4
"= "DEFG( =½68
!"#$%&$' ( ) $*+,- ./
010#
2 34 54 6789 :;<=>(Acetobac- ter)?@&ABCDE$9 FG H IF :JK !L#M0+N$OP QR STU1V(
0W :;<=> :B(A. aceti) Uῌ ! XY ZU [9+\( !(S( U ]W X^"_U` #$ W 89 +a#"
\( b(Aspergillus oryzae) !%
cEdEeK 3 0fgUeK
&(Saccharomyces cerevisiae) !"h>i 'jk 89PF l() %0
*aWma+n *a, $o !(+p) +n$
%q+rW!L% st +!L% eK-. +uY v/ w . h>i-.xy + zw . {=- . |D.#}~@ 0WW *
ῌ )12WN-9 3 &`4WOn %
0W5 !6P07
! 0$%
) .$+0 8(.$+0.
\( !( 9 !( ^ῌ
+ LI: 89 ;Z)
% !<3f^\(^
a#"=+%$ QR\(=
>0+?9@9
A. acetiU
IFOPA ,fPkB
C#}~@#7 $#$.-.
2`+ ¡D¢$£9 QR .¤¥U` ¦E#!L F§G¨k+ ©ªcªEe o W « f¬!-a#&
HfI®¯°±² f¬J2` K³%
$ ´µ¶0·W6# G¨k JL¸WNN¹M0Ng. O0 PPWU£º%
l() »Q¼½/
0W R¢H v/ l +
,
+ !"#"$%&' ()*+ ,-
ῒΐ ῐῑ ῍ ῌ ῌ !"#$%
῍ & ῌ 'ῌ ()'*+,- ./0123)45ῌ 67#894:;
<&)4=> ῐῌ῍+ῑ῍
?@'ῌ ABCDEFGDῌ HIJ0
<& &K>)4ῌ L῏<MNO#FPBC DQRE'ῌ L῏<MSTUῌ V
<WXYZ῎[\]^O#&_`abcQd
^<)4=>῍efῌBgOhiEῐgῑ QR#jk 5ῌ Pl ῐmnopῑ qRrs tSuvwxy<-z<῍ 'ῌ M{<
u|l}3O~ῌ L῏<ῌ ~]ῌ
r O6r῍ D'ῌ ῌ EU ῌ 4 'a<n,<D ῍ EQRDῌ OV<rs
#O῍
ῐ ῒu¡6 ΐ ¢£¤¥¦ῑ