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0 6 1 1.1 9 1.2 10 2 1 2.1 2.1.1 14 2.1.2 15 2.1.3 16 2.1.4 21 2.1.5 24 2.1.6 25 2.1.7 27 2.1.8 31

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2.2.1 32 2.2.2 32 2.2.3 34 2.3 2.3.1 39 2.3.2 39 2.4 2.4.1 41 2.4.2 45 3 3.1 3.1.1 47 3.1.2 50 3.1.3 52

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3.1.6 salivary oxytocin; sOT 57 3.1.7 58 3.2 3.2.1 60 3.2.2 60 3.2.3 sOT 62 3.3 3.3.1 64 3.3.2 sOT 67 3.3.3 sOT 69 3.3.4 sOT 72 3.4 3.4.1 sOT 75 3.4.2 sOT 76 3.4.3 77

(5)

3.4.5 79

4 81

83 84

(6)

5

5

(7)

sOT sOT sOT

(8)
(9)

1.1 β 1 1 1 1 altruistic behavior Hamilton, 1963; Trivers, 1971; Nowak, & Sigmund, 1998

(10)

1.2 cost benefit β Trivers, 1971 Eisenberg Mussen 1989/ 1991 Batson 2010/ 2012 β Ridley Dawkins 1981 β

(11)

Fehr, & Zehnder, 2009

Sober Wilson 1998

1 β

Sober, & Wilson, 1998

(12)
(13)
(14)

2.1 2.1.1

β

Alexander, 1987; Nowak, & Sigmund, 1998

Kurzban, & Leary, 2001 1

Kato-Shimizu, Onishi, Kanazawa, & Hinobayashi, 2013

(15)

1

Rege, & Telle, 2004

Izuma, Saito, & Sadato, 2010

2.1.2

(16)

Izuma et al., 2010 × β 2.1.3 Dictator Game 2

(17)

Burnham 2003

2 ×

Burnham, 2003 Haley Fessler 2005

1-A

0

1 ×

Haley, & Fessler, 2005 3 1-B

(18)

Oda 2011 Haley Fessler 2005

Haley Fessler 2005

Oda, Niwa, Honma, & Hiraishi, 2011

1-C

Mifune, Hashimoto, & Yamagishi,

2010 β

(19)

1 ×

Nettle, Harper, Kidson, Stone, Penton-Voak, & Bateson, 2013

1-E

Bateson, Nettle, & Roberts, 2006 1-F

Ernest-Jones, Nettle, & Bateson, 2011

(20)

A Haley and Fessler 2005 B Rigdon et al. 2009 C Mifune et al. 2010 D Nettle et al. 2013 E Bateson et al. 2006 F Ernest-Jones et al. 2011 G Powell

D

E F

Eye (left) and control (right) images

G

Watching-eyes Configuration (left); Neutral Configuration (right)

B

C A

(21)

Oda 2011 β Mifiune 2010 β 2.1.4 Engelmann 2012 5 helping stealing

(22)

1

stealing helping

Engelmann, Herrmann, & Tomasello, 2012 Engelmann 2013 5

5

Engelmann, Over, Herrmann, &

Tomasello, 2013 5 β

(23)

Leimgruber, Shaw, Santos, & Olson, 2012

β Houser 2012 6-11 i.e., i.e., private public 9

(24)

1

9 higher-order

Theory of Mind 1

× Amodio & Frith, 2006; Frith &

Frith, 2008

2.1.5

Premack Woodruff 1978 1

Premack, & Woodruff, 1978

1

1 Wimmer Perner 1983

(25)

1

× 1

Perner, & Wimmer, 1985 A X Y 1

B

A B

Autistic Spectrum Disorder; ASD

ASD 1

(26)

Wellman 2001

× Wellman, Cross, & Watson, 2001

6−9

Perner & Wimmer, 1985; , 2002

1 1 Sullivan, Zaitchik,

& Tager-Flusberg, 1994 4 5 Coull, & Leekam, 2006

1 2002

6

,

(27)

2.1.7

Banerjee, 2002

1

(28)

1 2 2 A A A 1 0次 一次的表象 二次的表象 A A A A

(29)

5 Engelmann 2012; 2013 5

5

Ultimatum Game Güth, Schmittberger, & Schwarze, 1982 5

Takagishi, Kameshima, Schug, Koizumi, & Yamagishi, 2010; Takagishi, Koizumi, Fujii, Schug, Kameshima, & Yamagishi, 2014

2

(30)

β × β 1 5 5 Izuma 2011 ASD ASD

(31)

β Izuma, Matsumoto, Camerer, & Adolphs, 2011 ASD 1 ASD β 2.1.8 × 5 β 5 5 5

(32)

2.2 2.2.1 42 17 , 25 , 70.4 , = 3.7 2.2.2 10 3 23 1

(33)

Mifune et al., 2010

(34)

2.2.3 DIK Wimmer Perner 1983 Perner Wimmer 1985 4-1 4-2, 3 4-4, 5, 6 4-7 3 1 1

(35)

1 2 3 4 5 6 7 8

(36)

5-1 5-2 5-3 5-4 5-5 5-6

(37)

4-7, 8

(38)

5 1 2 3 4 5 6 7 8

(39)

2.3 2.3.1 6 7 F(2, 82) = 10.15, p < .001, ηp2 = .198 Bonferroni t(41) = 3.01, p = .013, d = 0.59 t(41) = 1.59, p = .356, d = 0.17 2.3.2 76 32 / 42 12% 5 / 41 1 1 41

(40)

6 * p < .05 0 5 10 15 20 25 30 35 40 0 1 2 3 4 5 6 7 8 9 10 F re qu en cy of S am pl e Direct Monitoring Indirect Monitoring Non-Monitoring 0 1 2 3 4 5 6 Direct

Monitoring MonitoringIndirect Non-Monitoring

T h e A m o u n t o f A llo ca tio n Condition

*

*

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2.4 2.4.1 5 1 5 5 12 5 / 41 1 1 5 β 5 Takagishi et al., 2010; Takagishi et al., 2014 1

(42)

1 × 5

1

1

Bateson et al., 2006; Haley, & Fessler, 2005; Mifune et al., 2010; Nettle et al., 2013 Oda et al., 2011

β

(43)

4-5 1 1 8-9 1 8 1 1 1

(44)

8 5 β 8 β Age 3 4 5 6 7 8 9 1 0 Altruism Concern for one‘s own Reputation

Monitoring Condition

Concern for Direct Response

(45)

2.4.2

Tane, & Takezawa, 2011

β

Sparks, & Barclay, 2013

Matsugasaki, Tsukamoto, & Ohtsubo, 2015 1

1

(46)
(47)

3.1 3.1.1

×

1 9

1 Takahashi,

Diamond, Bieniarz, Yen, & Burd, 1980; Grewen, Davenport, & Light, 2010; Galbally, Lewis, IJzendoorn, & Permezel, 2011

Neumann & Slattery, 2015

Feldman, 2015 Insel, 2010; Feldman, 2012 Kosfeld, Heinrichs, Zak, Fischbacher, & Fehr, 2005; Baumgartner, Heinrichs, Vonlanthen, Fischbacher, & Fehr, 2008

Zak, Stanton, & Ahmadi, 2007

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Domes, Heinrichs, Gläscher, Büchel, Braus, & Herpertz, 2007

Kirsch, Esslinger, Chen, Mier, Lis, Siddhanti, Gruppe, Mattay, Gallhofer, & Meyer-Lindenberg, 2005; Baumgartner et al., 2008

Adolphs, Tranel, Damasio, & Damasio, 1995; Rauch, Shin, & Wright, 2003; Killgore, & Yurgelun-Todd, 2005

Kirsch et al., 2005

(49)

9

(50)

3.1.2

1

De Dreu & Kret, 2015 De Dreu 2010

2

ingroup love

outgroup hate 1 egoist

ingroup love outgroup hate

Social value orientation SVO SVO

ingroup love De

(51)

Ma 2014 Chinese Taiwan, S Korea, and Japan

Ma, Luo, Geng, Zhao, Zhang, & Kendrick, 2014

fMRI

OXTR rs53576 GG

Caucasian Asian

ACC

Luo, Li, Ma, Zhang, Rao, & Han, 2015

(52)

3.1.3

Bales, 2014; Dumais, & Veenema, 2015

Champagne, Diorio, Sharma, & Meaney, 2001; Pedersen, Caldwell, Walker, Ayers, & Mason, 1994

Neumann & Slattery, 2015

Nagasawa, Mitsui, En, Ohtani, Ohta, Sakuma, Onaka, Mogi, & Kikusui, 2015

(53)

3.1.4

Rilling 2012; 2014

Rilling, DeMarco, Hackett, Thompson, Ditzen, Patel, & Pagnoni, 2012; Rilling, DeMarco, Hackett, Chen, Gautam, Stair, Haroon, Thompson, Ditzen, Patel, & Pagnoni, 2014

2

a b

c

(54)

1

1 Rilling et al.,

2014

Rilling et al., 2012

Scheele, Striepens, Kendrick, Schwering, Noelle, Wille, Schläpfer, Maier, & Hurlemann, 2014

3.1.5

6

(55)

2009 Buttelmann Böhm 2014 6 8 β i.e., β 6 8 6 8 6 8 8 6 Buttelmann, & Böhm, 2014 6 8 β 5-6

(56)

Schug et al., 2013 5-6 4-6 Friend 1 Non-friend 1 Stranger Friend 1 Non-friend 1 Moore, 2009 × β

(57)

1

3.1.6 salivary oxytocin; sOT

Szeto, McCabe, Nation, Tabak, Rossetti, McCullough, Schneiderman, & Mendez, 2011 Seltzer, Ziegler, & Pollak, 2010

Weisman, Delaherche, Rondeau, Chetouani, Cohen, & Feldman, 2013; Feldman, Gordon, Influs, Gutbir, & Ebstein, 2013; Feldman, Golan, Hirschler-Guttenberg, Ostfeld-Etzion, & Zagoory-Sharon, 2014; Fujisawa, Tanaka, Saito, Kosaka, & Tomoda, 2014 Feldman

2011 4-6

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Fujisawa 2014 sOT

sOT Fujisawa et al., 2014

sOT

sOT Wesiman,

Zagoory-Sharon, & Feldman, 2012

× sOT

sOT

3.1.7

sOT

(59)

1

(60)

3.2 3.2.1 50 24 , 26 , 56.9 , = 11.6 3.2.2 10

(61)

10

2

(62)

3.2.3 sOT

sOT Saliva Collection Aid Salimetrics, Inc., State College,

PA 11

cryogenic vial 2ml 1ml × 6 1ml

× 44

9 11

-80

MACROPHI Inc., Japan, http://www.macrophi.co.jp/english/index.htm

column-based

purification 80

Assay Buffer 4

concentration ratio 5

800 g × supernatant ELISA assay

(63)

ELISA kit Enzo Life Sciences, Inc., Farmingdale, NY

SpectraMax ® Molecular Device, Sunnyvale, LLC, Sunnyvale, CA

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3.3 3.3.1 12 13 3.24 SD = 2.48 3.68 SD = 2.49 1 1 t t (49) = 1.21, p = .234, d = 0.18 × r = .407, p = .003; r = 386, p = .006 14

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12 0 1 2 3 4 5 6 classmate stranger M ea n di ct at or 's of fe r 0 2 4 6 8 10 12 14 16 18 20 0 1 2 3 4 5 6 7 8 9 10 D ist rib ut io n classmate stranger

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14 ×

0

1

2

3

4

5

6

7

8

9

10

30

40

50

60

70

80

Di

ct

at

or

's

o

ffe

r

(I

ng

ro

up

)

Age (months)

0

1

2

3

4

5

6

7

8

9

10

30

40

50

60

70

80

Di

ct

at

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Age (months)

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3.3.2 sOT sOT sOT M = 3.24 pg/ml, SD = 1.52, range = 0.94–6.31, n = 21 M = 3.20 pg/ml, SD = 1.65, range = 0.80–6.09, n = 23 t(42) = 0.08, p = .934, d = .03 15 sOT : r = .048, p = .838; : r = .075, p = .734 16 15 sOT sOT 0 1 2 3 4 5 6 boys girls S al iva ry oxyt oci n le ve l

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16 sOT ×

0

2

4

6

8

10

30 40 50 60 70 80 90

Sa

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Age (months)

0

2

4

6

8

10

30 40 50 60 70 80 90

Sa

liv

a

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n

le

ve

l

Age (months)

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3.3.3 sOT

sOT

1

sOT sOT sOT

sOT Model 5 sOT sOT Model 4 = .866, p < .01 = 1.730, p < .05 sOT = -1.284, p < .0001 sOT × sOT sOT r = 0.44, p = .039 r = -0.57, p = .009 17

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Classmate

Model1 Model2 Model3 Model4 Model5

Age months 0.348 * 0.378 ** 0.494 ** 0.866 ** 1.023 **

Gender female = 0, male = 1 0.075 1.088 ** 1.795 * 1.739 * 3.399

sOT -0.070 0.35 * 0.358 * 1.265 1.649 *

Gender × sOT -1.194 ** -1.196 ** -1.284 *** -3.052

Age months × Gender

-0.697 -0.552 -2.261

Age months × sOT

-0.969 -1.379

Age months × Gender × sOT 1.830

R .119 .348 .366 .399 .416

adjusted R2 .053 .281 .283 .301 .302

R2

F 1.798 5.203 ** 4.388 ** 4.087 ** 3.662 **

df 3, 40 4, 39 5, 38 6, 37 7, 36

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17 sOT -8 -6 -4 -2 0 2 4 6 8 R esi du al d ict at or 's off er (C la ssm at e)

Residual salivary oxytocin level

-8 -6 -4 -2 0 2 4 6 8 R esi du al d ict at or 's off er (C la ssm at e)

(72)

3.3.4 sOT

sOT

2 sOT

sOT sOT sOT

Model 5 sOT sOT Model 4 sOT β = -0.718, p < .05 sOT sOT sOT r = -0.02, p = .942 r = -0.70, p = .001 18

(73)

Stranger

Model1 Model2 Model3 Model4 Model5

Age months 0.280 0.298 * 0.253 0.249 0.209

Gender female = 0, male = 1 -0.282 0.328 0.052 0.053 -0.362

sOT -0.265 -0.012 -0.015 -0.026 -0.122

Gender × sOT -0.720 * -0.719 * -0.718 * -0.275

Age months × Gender 0.272 0.270 0.698

Age months × sOT

0.011 0.114

Age months × Gender × sOT -0.458

R .276 .359 .362 .362 .363

adjusted R2 .222 .293 .278 .259 .239

R2

(74)

18 sOT -8 -6 -4 -2 0 2 4 6 8 R esi du al d ict at or 's off er (S tra ng er)

Residual salivary oxytocin level

-8 -6 -4 -2 0 2 4 6 8 R esi du al d ict at or 's off er (S tra ng er)

(75)

3.4 3.4.1 sOT sOT sOT sOT sOT sOT

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De Dreu, 2012; De Dreu et al., 2010; De Dreu, Greer, VanKleef, & Handgraaf, 2011; De Dreu, Shalvi, Greer, Van Kleef, & Handgraaf, 2012; Van IJzendoorn & Bakermans-Kranenburg, 2012

Rilling et al., 2012; Rilling et al., 2014; Scheele et al., 2014

3.4.2 sOT

sOT

1 sOT i.e.,

(77)

Greig, Herrmann, & Zeckhauser, 2008; Bohnet, Herrmann, & Zeckhauser, 2009; Molm, Takahashi, & Peterson, 2000

Patel, Grillon, Pacletic, Rosen, Pine, & Emst, 2015 2

Fischer-Shofty,

LevKovitz, & Shamay-Tsoory, 2012

(78)

1

Moore 2009

Moore 2009

2

Buttelmann & Böhm, 2014; Schug et al.,

2013

sharing game Fehr 2008

Fehr, Bernhard, & Rockenbach, 2008

DLPFC

(79)

3.4.4 sOT Bales, 2014

(80)

Kramer, Ziegler, White-Traut, Bello, & Schwertz, 2007

Nagasawa et al., 2015; Seltzer et al., 2010

(81)

4

2

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