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(2) K.. 20. OzAKI. A. B. 初. =;こ;;司 Clematis. 1.. Fig.. PP. NSM. sp.. 16079,. Clematis. :. ultimate. veinlets. those. Smilax. These. fragmentary. Two. Discussion:. acrodromous. 2;. of leaflets. to. once. are. by. the. entire. to. in. areolation. leaves. are. of. which from. separable. and. Ceratopbyllaceae. Family Genus. Hu. miodemersum (Pl. 3丘gs.. Ceratophyllum. Discussion are. to. similar. to. of this species : NSM. record. Collections. CrIANEY,. et. Ceratophyllum. leaves. linear. Hu. 19,figs・. which. 4-6・. CHANEY・. et. This LINN・. C・ demersum extant cosmopolitan species, rare fossil floras ofAsia・ is rather from 455. PP 16178; TPM. Steu)artia. (P′1.1,丘gs. TANAl(in. part), TANAI. the. species The. Theaceae. Stewartia. hokiana. from. radiate. the. Genus. okuisui. CHANEY. pl. 42, pl.. miodemer亨um. Family. Steu)artia. et. 4, 6) 1938,. branching. Dicotomously. :. referred. is closely. Hu. miodemersum. LINN.. Ceratophyllum. Ceratophyllum. stem. small. veinlets・ complicated ultimate C. patens MoRR. et DECNE. Of. extant. similar C. chinensis OsBECK Of China. NSM PP 16079, 16233. Collection:. Japan. feature. characteristic. and The. and. of the. a. show. margin,. areolation. those. (×10).. 1). branched.. twice. smaller. are. Fig.. lea且ets. venation,. B. (×2),. sp.. 1Clematis (Pl. 8,fig.. A. OzAKI, 2,. 5-7;. et. N.. LINN. new. Fig.. SPeCies 2). SuzuKI,. 1965,. p.. 142,. pl.. 14,fig・. 81. (only).. Holotype:. TPM. 307.. Upper. Tochiwara. Formation. (Late. Miocene)・. SajL.
(3) Late. Miocene. Tatsumitoge. Flora. 21. A. B. 2o Fig・ B.. 2・. Venation. NSM. PP. Description apex. Leaves. :. base. acute,. oblong, to. cuneate. secondary. curvlng. uP. apex. area. hSlngle. once. Discussion relationship -that induced dromous. to. the. 284. lower. the. and. corner. area. 3.5. (×4.5),. to5.Ocmwide, by. serrate concave. the the. of. Steu)artia. vary. a. short. :. off. length. veins. that. series. of loops. are. of. 8. 10. at. an. midvein. to. lamina,. gently. making series of loops irregularly in percurrent, loop. to. near. the. the.tooth veinlets. a. close. foliar. shape,. aplCal. side of the teeth, brocbidoin the lower half of the. the. arranged in. short. a. of the. oblong. aplCal. very. sending. from. part. short. and. straight,. diverglng. veins. veins intercostal. in. from. genus. by. form. and. TPM. secondary's veins formlng Irregular polygonal ; ultimate meshes branching; more 5mm in length. than petiole : leaves The have indicate several that characteristics. secondary. lamina. A.. and. side,. stout,. the. on. spaced. veins. extending. ; quaternary. ; midvein. ; secondary. long. basal. on. convex. apex. closely. tertiary. or. sp.. crenately. joinlng Superadjacent. and. -the margln; marglnal. n.. llcm. to. margln. teeth. veins. 45o,. to. 7. about. obtuse,. -side of the teeth; marglnal aplCal ton side with attenuate ・subopposite angle of 40o. hokiana. (×10).. Prefecture.. Tottori. mura,. of Stewartia. characters. 16039. 2o. closely its distal part,. cremate-like. margin. percurrent tertiary weakly once branched These veins, and simple ultimate closely veinlets・ specimens to living Steu)artia the MAXIM. in Honshu to related pseudo-camellia growing to. Kyushu ct. ONOE. of Japan・. new. the. extent. secondary veins TANAI (1961) as a. in the. and. ・of the by. from. This. holotype ・specimens the genus capsule.. is. from separable S. monadelpha SIEB. et. species lower. half to. of the the. S. submonadelpha by Zucc. many. lamina・. S・ okutui. number. was. reported S. pseudo-camellia, but the The speci丘c status. other. related species extant S. okutsui is too fragmentary to take a TANAI by N. SuzuKI and of S. okutsui reported Stewartia in the venation character and the entire. of. TANAI. (1965) margln,. differ. from. except. a.
(4) 22. K.. Collections:. NSM. PP. 16039;. Steu)artia. 722,. p.. Discussion: and This. but. venation,. is. species. are. in central. growing. delpha. are. 1961,. to. Japan. southern. lower. in the. PP. NSM. half. 16009,. TPM. Tottori. acuminate,. base. 2. glands. zlgZag. Leaf. OzAKI,. along. the. loops. with. an. about. the veins. a. base. margln,. quaternary. and sents. venation Sapium.. (Late Miocene).. from. the. then. the. lamina weak. then. up. Saji-. Weak. ortbogonal The foliar. of the. marglnal. The. leaf of. veins. area. this. new. the superadjacent forming and. to. palr. Of. the. with. the. in intercostal. basal. extending. margln; area. series. weakly. of the. along. superadjacent. secondaries basal. part. secondaries a. tertiaries. extending. and. distal. the. up. or. angle. lamina. the. to. middle. slightly an. at. midvein of. long,. 3.5cm. and. portion veins from. 1nterSeCOndary percurrent. ;. veins not welトpreserved. base at the of of the lamina pair glands indicate that the fossil repreof the lamina. reticulate shape,. the. connecting. and. along. ; tertiary. the. apex. wide,. extending. base. the. secondaries. one. runnlng. Vein. from. ln. secondaries. curving. margln;. prlmary. from. superadjacent. basal. the. straight. curvlng. veins. (estimated). petiole. from. branches. with. 4cm. diverglng. ZlgZag. Secondary. from. rather. veins Discussion:. 8 palrS,. about. upper. near. of the. lamina;. the. of. of loops. of 60o. angle. entire,. secondaries but runnlng. branches. simple,. 3). long,. diverglng. Series. loops. form. species. Formation. 5cm. oval,. straight. branches. margln;. of the. number. 4773.. new. Fig.. ll;. margln. base. veins. basal. nearly. forming. small. P. BR.. Tochiwara. cuneate,. the. at. 50o,. about. Upper. widely. widely. ; Secondary. extending. the. S. serrata. Prefecture.. DescriPtion:. small. 242.. 4771,. GSJ. (Pl. 2,丘g.. Holotype:. in. Zucc.. et. S. submona-. lamina.. Sapium. hokian,um. Sapium. SIEB.. monadelpha. Euphorbiaceae. Genus. to. S.. character et ONOE.. TANAI. Cheju Island of Korea. S. pseudo-camellia MAXIM.,. and. of the. 16008,. Family. mura,. UEMURA,. et. marglnal. submonadelpha. living. the. shape,. modern somewhat resembles S. hokiana n. and sp., but is distinguishable. secondary veins Collections:. then. in foliar. the. MAXIM,. of. 53, pl. 18, fig. 6. HuzlO王くA. p.. Steu)artia. to. similar. to. ONOE. et. 3, 4). variable. referred. closely. 1,. 478.. 9, 10.. leaves. These. 307, 399,. TANAI. 1,丘gs.. ONOE,. et. 12,丘gs.. pl.. TPM. submonadelpha. TANAI. submonadelpha. 1973,. 4772;. GSJ. (Pl. Stewartia. OzAKI. ;丘ner. a. species. is similar. to. the. extant. S.. sebiferum.
(5) Late. Miocene. 23. Flora. Tatsumitoge. A4S ∼. 7. B. Fig.. 3.. 242A,. Venation B.. The. basal. in China,. RoxB.. (L.). of the. part. and. growing intersecondary. the. secondary. veins. counter. but. Discussion. points. (SEIB. china,. :. SuzuKI,. K.. ProtojaPonicum This. et. but. PAX.. Zucc.) is. Collection/. entire. This. veins・ HoFFM・. et. NSM. PP. Discussion by. secondary Colleciion. :. Very. its large. 75, pl・. 17,figs・. in Japan. the. small. TPM. venation to. except. number. of. the extant. and grandular S・ jaPonicum. Hokkaido, the. K・ SuzuKI. Korea. secondary. and veins・. 16069.. Simaroubaceae. Ailanthus. sp.. (Pl. 3,丘g.. 7). fragmentary gland. on. DESF.. Ailanthus. only the. basal. one. leaflet is identiBed. margin. veins. :. 5-7・. ProtojaPonicum. is simialr. species. in. slightly different :. p.. growing. Genus. basal. and. SuzuKI. is referred to SaPium brochidodromous margin,. Family. Ailanthus. midvein. 3). 2,丘g.. 1961,. specimen. slightly undulate on the marginal. K・. Protojaponicum (Pl.. the. the. secondary. 242.. TPM. Sapium. in. of the. number between. angles. TPM. A・. in small. narrower. AI. (×2)・. sp・. veins.. Collection:. sapium. n・. of the丘g・. part. is different. in the. and. hokianum. Sapium. of. characters. 458A.. Family. Sapindaceae. and. the. with. characteristic. the. genus. incurved.
(6) 24. K.. Genus. OzAKI. Koelreuteria. Koelreuteria. Hu. miointegrifolia. Discussion. Well. :. miointegrifolia Hu tegrifoliolaMERR・. K.. and. Collections:. et. 1, 2,. CII-IANEY,. 1938,. capsules. and. preserved CHANEY.. et. NSM. This. ;. Family. bifurcate. veins at. the. margln. intercostal The on. leaaet. area・. the. or. fragmentary shoot. With the These. is similar My. margin・. jaL,anica which Collection:. to. adjacents,. is closely. enter. the. percurrent. the. to. similar. warm-temperate secondary. This. leaf, the. that. marglnal. tertiary. diverge and. veins. wbicb. developed丘ner. weakly Collection:. NSM. PP. China.. 470.. 8). is devoid. Meliosma China. and. rigida in the. midvein. thick. texture,. but. nearly. transverse in ramiBed lea月・et represents Rhus.. compare. with. than. lower. SIEB・. fourth. Aquifoliaceae. half. serration R.. et. narrow. palaeo-. to. the. lamina,. in. growlng. foliar shape,. angles. is di庁erent. order.. of the. Zucc・. oblanceolate. perpendicular. 16039.. Family. loops. make. regular. of the. at. veins. ; secondary. not. in having to. the. more. do. BLUME. (Pl. 4,丘g.. are. venation. in 1970.. sp.. the. the. LINN.. Meliosma. from. teeth. 287,. in-. Sabiaceae. thoughit. of Japan. region veins. that. Meliosma. extant. TPM. and are. veins. ill-preserved. MATSUO. Genus. margln. tertiary. Family. Discussion:. the. javanica. is too. reported PP 16245.. NSM. 4782;. characteristic. indicate. of R・ by. was. K.. extant. 12). near. and. characters. leaflet. the. sp.. veins. those. to Koelreuteria. referred to. fig. 3.. LINN.. Rhus. leaflet has. tertiary. 4770,. 1, 3, pl. 39,. in southeastern. growing. (Pl. 2,丘g. A. are. is similar. GSJ. Rhus. :. leaflets. 38,丘gs.. Anacardiaceae. Genus. Discussion. 64, pl.. p.. FRANCH.. 16047116051. CHANEY. et. 7). species. bipinnata. PP. Hu. miointegrifolia (Pl. 4,丘gs.. Koelreuteria. LAXM.. and. in. more. midvein. the. generally closely and. in.
(7) Miocene. Late. Genus Ilex. Ilex. LINN.. HuzIOKA. subcornuta. (Pl. 3,丘gs. Huz-l・OKA. )lex・subcor顎uta. Discussion:. Only. )lox. subcornuta. HuzIOKA. ilar. to. the. et. UEMURA,. two. but. 1973, well. et. extant. 10, 12) 716,. p.. (1973).. LINDL.. I. cornuta. UEMURA. et. 12, figs. 2-5.. pl.. leaves. preserved. UEMURA. 25. Flora. Tatsumitoge. identi丘ed. are. This. fossil. species. in. central. China. grOWlng. with is closely and. the. sim-. southern. Korea. Collections:. NSM. PP. 16010;. TPM. 391.. Ilex. sp.. (Pl. 7,丘g. Discussion. small. and. is similar Korea and. Japan. acute. to. one. the. its. adjacent NSM. Coll9Ction:. PP. several. THUNB.. Euonymus. number. Hu. et. HuzIOKA. has. leaf. one. elliptic foliar shape,. leaf is closely planipes KoEHNE. veins.. is leaf. in Japan, growing from Miocene the. Celastraceae. to. similar in the. the. a. acute. 8) close. apex,. E・. extant. LINN, sp.. (Pl. 2.丘g.. Euonymus. reported. The. a. area・. Euonymus. in. varieties. that. serration. secondary. its. indicate. 16092.. Genus. Only. marginal. been. not. that. characteristics. and. fossil leaf has. Family. Discussion:. 6D). brochidodromous. apex,. Similar. Fig.. foliar shape,. oblanceolate. I. crenata. extant. China. and. :. splny. with. leaf has. small. Ilex. to. relationship. close. of. Only. :. 2;. to. relationship finely. and. alatus. SIEB・. the. serrulate and. leaves. margln・. is also. similar. of This to. E・. base, serration, the and marginalfine leaf is from E. veins. separable protobungeana of the secondary TANAI ONOE CHANEY E. et (1938), (1961) and E. okamotoi palaeoseieboldiana acute. apex. This. (1974). by. marglnal. serration. TPM 508.. and負ner Collection:. the. acute. basal. shape,. Family Genus Buxus. small. of the. number. Buxaceae Buxus. protojaponica. LINN. TANAI. et. ONOE. secondary. veins.
(8) 26. K.. OzAKI. 7,丘g.. (Pl. Buxus. TANAI. ♪rotojaponica Discussion. characteristic cies. venation. TPM. Berchemia. elliptic foliar shape, that. veins. China・ growing. Paucijlora the. Japan. in. MAXIM.. 5, ll). referred. to. the. closely. to. to. to. the. NSM. PP. 16045,. 16046,. Genus Houenia. Hu. miodulcis. are. 3.. characterized veins. and. and are referred to且 Shanwang flora of longeracemosa OKUYAMA. B.. are. and BRONGN.. also. Of China. 且. to. similar. irl the number. 16157.. Houenia. THUNB. Hu. miodulcis (Pl. 4,丘g.. Discussion. 2,. secondary. apex,. B. jloribunda. and. Berchemia. genus. extant. round. pl. 40,丘gs.. the midvein, the Miocene. from. similar. obtuse. 5,. 391丘g1. euCamptOdromous. reported. the. 65, pl・. veins.. Collection:. Houenia. spe-. ja?onica (MuELL,. ∀ar.. CIIANEY. et. p・. 1938,. perpendicular. Of Japan. secondary. (Pl. 4,丘gs.. margln,. CHANEY. are. Hu. are. nearly. et. leaves. in. Zucc.. et. NECK.. BerchemL'a. miojloribunda. entire. are. Hu. My. oblong. sized SIEB.. the. with This. Rhamnaceae. CIIANEY,. et. leaves. Four. :. miojloribunda. of. Hu. mioj7oribunda. tertiary. protojaponica foliar shape・. 250.. Genus. by. 5.. Bu・xus. with. microphylla. Family. Discussion. 14,丘g.. pl.. WILLS.. et. Collection:. Berchemia. 46,. p.. small. and. B.. the.modern. REHD.. 1961,. leaf is identiBed. one. ramified. resembles. ARC.). Only. :. ONOE,. et. 4). et. The. :. CtlANEY,. 1938,. deltoid-oval to. nearly. p.. et. 10). 65, pl.. foliar. the. CHANEY. 1, 4,. 40,丘gs.. shape,. trinerved. and. 5. primary. marglnal. that. Veins, teeth. perpendicular midvein acuminate with seta A that Houenia・ two the fossils represent and single nearly complat・e fragmentary leaves are to H. Hu et CHANEY. This referred miodulcis species・ is closely similar to the extant H. dulcis TBUNB. in Korea Japan, growing and indicate. China. and MuRAI. flora. also. similar. (1963) reported. of Japan,. cordate. to. Collection:. H.. tomentella. prlmary. TPM. 483,. (MAKINO) from. masuzawaensis. this leaf is considered. but. base,負ve. H.. Veins, 504.. secondary. to. belong. veins. NAKAI. Of Japan. Miocene middle Tetracentron genus. the to. and. acute. marginal. Gosho by. teeth.. the.
(9) Genus. LINN.. Rhamnus Rhamnus. sp. 3;. (Pl. 4,丘g. Discussion. :. SIEB.. et. crenata. the former. at. that. veins. extend. form. and to. oblique. 45o,. to. the. specimen. Collection:. Sageretia. Sageretia. PP. NSM. HolotyPe,. Tottori. Saji-mura, 1.7cm. wide,. rather. low. acute,. apex. of the. base,. lateral. extending. basal. veins. apex,. 1amina. JOlnlng. the. at. other. diverglng. at. cuneate. a. angule. to. polygonal. petiole. 4. to. 6mm. strong. veins. 60o, forking. with. the. marglnal. part; area. ; ultimate. meshes in length,. weakly. tertiary. entering veinlets curvlng・. 3. from-. runnlng. middle. the. along part. extending. of the二 rather. to. lateral loops. veins. prlmaries at. among. one-third the. once. to. twice. in. the. inner. secondariesb. teeth ; quaternary. marglnal mainly. the. of. joining each and angles 4 in number, alternate,. wide. midvein. to. gently'. lamina,. diverglng. prlmaries. 60o, joinlng With the formlng and marglnal. 1n. and. midvein. of the length. veins. basal. of the. above. prlmaries. one-third. prlmaries in the. with血e on basal-. midvein;. middle. about. of the lateral. side. from. the. to. one-fourth. slightly. the. secondary. of the. to. at. Ill. and. convex. and. ; lateral. in. at. long. one-third. part. upper. veins. secodary. SeCOndaries angle of 30o to. reticulate,. making. Miocene)-. Serrulate. side. side. midvein. point. up curvlng of lamina, in aplCal from the margln. random. the. Of rather. margin,. margln. aplCal. aplCal. to. tertiary. in abaxial. middle side. (Late. 2・8cm. to. round, on. its middle. the. 2・O. Veins separating of 20o to 40o with. prlmary. palr. of about. to. of the. separating. With. angle. an. 4A・D) Formation. ovate,. concave. length. or. veins. an. at. base. parallel. the. ; secondary. straight. to. ln. zlgZag. below. then. oblong. an. making. secondary With from the apex;. midvein. small,. three. running. and. up. Tochiwara. teeth. side;. slightly. margln. the. SPeCies. Figs・. Upper. marglnal. acuminate. with. new. 4, 6, 9, 12;. ags.. 16038・. Leaves. teeth;. those. tbe. SuzuKI. Prefecture・. Description:. lamina. in. BRONGN・. OzAKI,. hokiana. (pl. 4.. JOlnlng. Thisr. China・. and K・. Protocrenata. Rhamnus-. way・. Korea. Hokkaido,. tertiary. and. the. the. to. parallel. margln. on. ana一. prlmary. 347.. TPM. Genus. curving. fork. and. secondaries. run. the. along. the. to・r. related. serration・. marglnal. side. loops. some. that. veins. secondary. is. but. between. angle. Rhamnus. extant. of the. WEBERBER,. teeth,也e. in Japan except is widely distributed is different from those of Rhamnus. c,enata. 血e. 40o. those. fragloides. Zucc.. veins that are its distal part. secondary margln. and Rhamnella in the incurved marglnal. one. 5F). Fig.. leaf resembles. fragmentary. This. 27'. Flora. Tatsumitoge. Miocene. Late. veins. branching;.
(10) 二28. K.. Discussion. Five. ・・. foliar. ; small. ・foliar shape These veins・. by. are. the. are. to. suprabasal. NSM. to. to. in. PP. 16038,. Genus. the. 16171;. もy. its foliar shape,. 4770,・. tertiary. leticulate. I(SIEB・. et. Zucc・). the. are. species of MAXIM.. secondary. veins. This. veins・. but. 235.. PLANCH.. sp.. 4;. Fig.. 5G). leaf is referable to the Parthenocissus genus incurved of marglnal teeth and rather random leaf is similar to the P・ extant iricuspidata. number. PLANCH・,. TPM. Vitaceae. Parthenocissus. fragmentary. small. but. S. paucicostata of. elliptlC. prlmary. living. the. of. number. GSJ. (Pl. 5.丘g. This. trinerved of Paliurus,. Among. those. Parthenocissus. :. leaves. the. :. Veins.. Family. Discussion. of Sageretia. ; Supurabasal. veins・. similar. China. northern. prlmary. Collections:. margln. primary. closely. feature. characteristic. slightly similar. supurabasal. leaves. in central. a. show. size ; serrulate. specimens. ・distinguishable Sageretia, these rgrowing and the. leaves. OzAKI. is. slightly different. in the. cuadate. apex. the. of. lobes.. Collection:. NSM. PP. 1622l.. Family. Tiliaceae. Genus Tilia. Tilia. distans. LINN.. NATHORST. (Pl. 5.丘g. 'Tilia disians. NATI10RST,. flgS・ト4,. 1883,. TANAr,. 1976,. 65, pl・. p・. p・ 336,. pl・. 7). 4,figs・. 5113, 2,. 4,軸・. pl・. =uzIOKA,. 7・丘g・. 1943,. 3,. pl・. 121,. p・ fig.. 9.. 2,. 21,. pl.. text・丘gs.. 5g,也.. Discussion:. Only. leaf is referred. one. discussed. by TANAI ・species This species is closely similar in western growing Japan. was. Collection:. NSM. PP. (1976) to. the. et. et. CfJANEY,. ONOE,. basis. the. T. kiushiana. Hu. miohenryana. (Pl. 5,丘g. Hu. livlng. NATHORST.. distance the. of. MAEINO. Migi et. This. specimens. SIJIRASAWA. 16077.. Tilia. JTilia miohenryana 丘g・ 3, TANAI. Tilia. to on. 1961,. 1938・ p・. 3, pl. p・. et 6,丘g.. 69, pl・. 52, pl・. CHANEY. 14,丘g.. 7). 22,fig・ 9.. 3a, pl・. 44・. flgS・. i,. 2,. pl.. 45,.
(11) Late. A. Miocene. 臥. Tatsumitoge. Flora. B. 29一. C. (. 唱. 1cm. 2cm. 2cm. D 2o. F E. ●. 2mm. 7L. 2o. 2cm. 2cm. Fig.. 4.. GSJ. 4770,. NSM. PP. Venation B,. D.. 16038.. A-D. of fossils. characters NSM PP 16038 (Holotype), F.. Acanthopanax. sp.. GSJ. Sageretia C.. 4796.. TPM. hokiana 235.. E.. n.. Hedera. sp.. A. sp..
(12) K.. 二30. Fragmentary. Discussion: Hu. et. CHANEY. OzAIくI. leaves. from. reported. the. Tilia. to. similar closely Miocene Shanwangflora are. miohenryana China in foliar. of. base and the number cordate of the secondary ・shape, lnequilateral HuzIOKA in the wider fossil species is separable from T. subnobilis is also from T・ base, cordate and inequilaterally and separable -ENDO. in the. generally. My ・shape. SHIRASAWA. leaves. large. have. of the. number. close. secondary to. similarities. PP. 6415,. 6445,. Tilia. Discussion from. 1966,. These. leaves. Miocene. Late. -to the. :. ENDO,. T.. extant. Collections:. 189,. GSJ. region. TPM. 121, 452.. Tilia. of. HuzIOKA,. Discussion. Well. Tilia the. Early leaves. growing. ・.. to. subnobilis 丘g.1.. 1943,. subnobilis. secondary veins. Miocene Yongdong且ora This. species. in northern in China.. growing Collections:. 125,. pl.. 5,丘g.. and. in the. oval. This. Honshu. similar. PP. Family Genus Rotala. Holotype: Tottori. TPM. 492.. Prefecture.. Rotala. hokiana. Upper. on. two. 29,. referable. and HuzIOKA. number. from. fragmentary. TPM. 167.. Lytbraceae. OzAKI,. (Pl. 5,丘g.. Tnura,. by. are. pl・. SHIRMONKAI T. maximou)icziana T・ REtID・ Japan and et WILS・ nobilis. 4837;. GSJ. 16078;. based. base. 360,. p.. to the living. Hokkaido,. and. leaves. foliar shape, cordate was first reported Korea. 1961,. TANAI,. fragmentary. some. of northern. is closely. NSM. species. 5). 22(IT),丘gs. 2, 3,. pl.. one. preserved. HuzIOKA. HuzIOKA. subnobilis (pl.. Tilia. 3・. reported referable to Tilia protojaPonica ENDO are Akita Prefecture and of closely similar in cool-temperate of Japan・ growing. are. SIMONKAI. 4833;. 2, 6). 23,丘g.. pl.. Sanzugawaflora. japonica. ENDO. Protojaponica. p.. 99, 168, 201, 453.. TPM. 4834-4836,. GSJ. (Pl. 6,丘gs. protojaPonica. protojaponica. larger foliar and veins living T・ maximowicziana. in Japan.. growing Collections: NSM. Tilia. the. This veins・ foliar shape. 1;. Tochiwara. LINN. new. Fig.. SPeCies 5D). Formation. (Late. Miocene)・. Saji-.
(13) Late. DescriPtion. Leaves. :. the. at. slightly narrowed Discussion:. 5. A. ∼Iiocene. 6 verticullate,. or. base,. single. troplC. Of the TPM. reglOnS. Collection:. leaves・ by. ponds,. Genus Cornus. Hu. margln.. Cornus. LINN. Hu. miowarteri. CfIANEY,. et. 1938,. et. Chaney. 1, 8). 72, pl. 47, figs.. p.. i, 2, 4.. to Hu My leaves are Cornus referable miowarteri foliar size and 3 to 4 palrS Of the secondary veins. TANAI ONOE is to (1961) this similar species, but and. Discussion: in the. small. by. reported in the undulate. NSM. PP. 16071,. My. leaves. shape,. large. Discussion:. must. veins, 2) has be. separable. omitted Collections:. type. brocbidodromous from NSM. the. the. of. extant. C.. quaternary. ovate. foliar. specimen. shape C. the of. and. and. veins. and. coriaceous. the. venation Cornus. genus. PP. 16037,. GSJ. 4786.. 16053;. Family Genus. Araliaceae MIQ.. AcanthoPanax AcanthoPanax. (Pl. 7,丘g.. 1;. sp. Fig.. the. minoensis. 4F). HEMSLEY. controversa. number of the secondary veins HuzIOKA C. the reported minoensis. similar Korea in the of the. 3). the to. but. to. similar. meshes. closely. Changgiflora ondary 17,丘g.. are. though. veinlets, are. subkousa. is. sp.. (Pl. 6,丘g.. ultimate leaves. CEANEY. 16303.. Cornus. foliar. et. C.. margln.. Collections:. in the. to. of temperate. wetlands. Cornaceae. (Pl. 6,丘gs. miowarteri. hip-. is separable The extant. world・ 492.. Family. Cornus. obtuse,. Rotala. specimen. entire and. living. the. This. the. linear, apex. entire・ to. similar. 1enear. in the verticullate small those of Hydrilla uerticL'llate CASP. in lakes, R. hipPuris is distributed species. broad. to. margln. is closely. MAKINO. puris from. linear. obscure,. veinlet. specimen. 31. Flora. Tatsumitoge. complicated is. (1972) from. number. of. (HuzIOKA, texture,. small・. then. the 1964, the. My the sec-. pl・ type.
(14) 32. K.. OzAKI. 0. m. 一.......-.....__-. _1c. 0. て≡. A. D. 2o. ⊥/. E. 8cm. 滋. 刀. 2o Ll,. 1cm. B. C. 2o. 1 Cm. -\\. 乏)2隻二プ. G. F. -ぐ. I--、_/1. \.  ̄ _/ {三. %. 77. 2cm. 2cm. クエ Fig・. 5・. Venation. Paulownia. A・. STEUD. n・. sp・. 347.. HUPB. form. E.. Aralia. Parthenocissus. sp.. that veins, margln. :. angular. angular. indicate. TPM. shanwangensis 647. C. Chionanthus 492.. Discussion. fossils. of TAO. TPM G.. characters. The. loops. tertiary that. the margln,. at. the. sp.. n.. at. fossil. the. PP TPM. some. 16223. 156.. extant. tomentosa Rotala. D. F.. species.. (TとIUNB.). Rhamnus. hokiana TPM. sp.. 1621l.. foliar shape,. obovate. loops. PP. related B. Paulownia. NSM. sp.. hokiana NSM. and. 309.. irregularly. Orthogonally margin,. represents. and a. spaced. reticulate angular. secondary. intercostal teeth. at to. the. veins tertiary upper. close the genus relationship fossil is similar to the extant A. FR. et sciadophylloides SAV・ in Korea China, but Japan, is distinguishable in the small growing and number the teeth. of marglnal. AcanthoPanax・. Collection:. This. GSJ. 4796..
(15) Late. Miocene.Tatsumitoge. Genus Aralia. hokiana. TPM. :. Tottori. mura,. OzALKI,. serrulate,. just. veins. making. also. diverglng. midvein. and. portion,. the. lower. sending. o庁some. making. an. angle. base. the. two. branches. at. of about. 40o. Aralia. to. marglnal. teeth,. marginal A・. loops. :. 1n. its with. and. making. loops. differ. marglnal. of the. in. grdbra. midveinF;. area. enterlng notL. several. of. tertiary. superad-. an. the. ■veins. the marglnal. one. the. with the. angle. to. the. strong. on. way at. part;. from. 30o. about. middle. preserved. with. basal. the. midvein the. the. abaxial 30o and of. upper. palrS. in'intercostal. teeth ; quaternary. well. indicate. that. characteristics. A・. aplCal. eそOanq TANAI. (1970). are. similar. to. basal. and. marginal shape, serration and My lea月.et is sirrlilar叶those of the in the Japa?, -but is slightl女′.'d:iffernt. secondary. veins. in growing. MATStJM.. the. part,. (1959) and. the. veins. apICal. diverging. pair. midvein,. on. the. close relairregularly apex, shape, aristate spaced form that arranged secondary veins sometimes Among也e fossil species from Japan, reported. margin. et ONOE. this lea且et, but. the. irregulary. foliar. irregularly. the. at. states. sinuous. long. ll. 3cm. margln. in. angle. from. veins. cordate,. an. make. distal. marglnal. inequilateral. celtisfoliaTANAI. ext?ntA・. Miocene).▲ Sa〕i-. inequilateral,. weak. vitb. tertiary. diverglng. palrS. percurrent,. that. strong. o庁rather. one. 60o. about. of. orthogonalreticulate ,・血er Discussion : This leaflet has. tionship. Formationノ(Late. lowest. the. margin,. from. angle. area古elakly. the. at an. send. veins. Tochiwa;a. ; midvein semicraspedodromous ll to、12 palrS, 1rregularly spaced,. veins. and. pair. 5E). :. venation. secondary. base. Fig.. SPeCies. Leaflet slightly ,1anceolate-oblong, base (estimated),Jwide, apex aristate,. 4・ 3cm. jacent. new. Prefecture.. Description and. Upper. 156.. 33. LINN.. Aralia. (pl. 2,fig:2;. Holotype. Flora. lて. 't. marglnal. small. serratioh.. Collection:. TPM. 156.. 'Genus -. Hedera. spべ∴. (pl. 3,fig. Discussion. just those Korea. the. of the. a. close. brochidodromous. margin,. above. indicate. that. acteristics entire. This. specimen_,. :. base. extant. bifurcate. and H.. rhombea. China.. and Collection:. NSM. PP. 16070.. I=INN.. yHedera. ll;. Fig.也). thoughmisslng. the. relatilOnShip. on. the. (MIQ.). of. three. venation,. way.. BEAM. aplCal Hedera:. ・palrS. This. growing. has. part, ovate. Oflateral. some. foliar. veins. charshape,. that radiate. leaf is_ closelyy:-similar. in,Japan. except. to. Hokkaido,.
(16) 34. K.. OzAKI. イて'. D. C. A. L \. 嶺. しし∼、. \ J. + -r. ㌔(.  ̄、㍉ (I(I. 感当. \ー〉丁・. \. ( Fig・ A.. 6.. Venation. characters. fossils. of. Kalopanax. dambar HUPI∋. 178. JJβ∬sp.. D.. (×8).. NSM. Genus. Kalopanax. Ealopanax. n-suzukii. acerij'oium ONOE,. et. WoI.FE. 1959,. HuzIOKA,. et. 285,. pl. 7,丘g. 71, pl. 9,丘g.. p. p.. 16092. extant. (×8).. Kalopanax. species. B. Liqui・ NAKAI. Pictus. (×3).. WoLFE. TANAI. et. Fig.. 6A). 42, pl. 22, fig. 4. 1938, p. 20, pl. 47,figs.. p.. 3, 5, TANAI. CIIANEY,. 4. TANAI 9.. C.. some. 16011. MIQ.. 3, 5;. 1980,. Hu. (NATI-IORST). 1972,. (×8).. ”-suzukii. TANAI,. et. 34 PP. KaloPanax. (Pl. 7,丘gs.. Kalopanax. related and PP NSM. TANAI. WoLFE et n・suzukii HANCE HUPB formosana. et. N.. HLTZIOKA. et. SuzuKI,. 1965,. UEMロRA,. 43, pl. 20,flg.2.. p.. 1973,. p.. 923,. 18,. pl.. 丘g.1. Liquidambar. Discussion the. Hu. miocenica. Well. :. semicraspedodromous. marginal those. serration ultimate radiate. and. TANAI. small. and. similarity. the. of the. those. of. one. leaf and. some. leaves. veins. shape,. that. new. the. pl. 3,丘g. are. 6. acute. from. base. the. (only).. identical. marginalfine. radiate. to. teeth, of. the. of fossil Kalopanax ”-suzukii closely resemble fossil Acer Liquidambar in the and marglna) distinguishable from by less branched the former latter. marginal. by. existence. teeth.. 1, 2) to the. epithet, extant. genus. K. n-suzukii, K.. of. TANAI. Hu. acerifolium (NATHORST). 5, pr: 9,figs.. (1980) reported to. are. from. of KaloPanax. pl. 8,fig.. and. of. but. base. p. 316,. leaves. leaves. venation,. veinlets from the. specimens 1883,. Some. lobed. 1976b,. foliar. palmate. venation,. teeth.. of丘ve. large. in the. TANAl,. (in part),. preserved. Kalopanax. genus. CfIANEY. et. some. 丘ne. (1976). removed. et. CHANEY. Liquidambar, for. pictus (THUNB.). the. fossil. NAEAI.. and leaves. The. veins the. which type. (NATHORSTi WoLFE. and. have which living equiva-.
(17) Miocene. Late. K.. 一ent species, ern. china. and. NAKAI. pictus (THUNB・). Clethra. 、Saji-mura,Tottori DescriPtion 2 to 3. 5cm .acute. to. Leaves. diverglng. part. about. 60o. tertiary. one. in the. up. making. secondary's. enterlng. new. north-. 4776.. SPeCies. 8,丘g.. pl.. Upper. 3;. 7). Fig・. Formation. Tochiwara. Miocene)・. (Late. the. base. or. acute. teeth. of the. lamina;. and. to. two. cuneate, to. arche teeth. and. upper. 15. by. arches. to. the. another. one. JOlnlng. reaching. two. thin,. straight. or. 20. opposite. to. to. 70o. in. basal. parallel each other, in its distal partwith. forking. then. margln,. with. each. of about. secondaries from the extending. adjacent area. marginal. percurre叫in. and. Serrulate. between. rather. almost. part,. long. lOcm. margln. three. midvein. slightly zigzag,. polygonal area. the. 6 to. linear-oblong,. to. marginal. middle. in intercostal. veins of the. corner. basal. and. 4774,. semicraspedodromous, veins from the midvein at an angle. curving. gently. angles. 6-10,. acuminate,. Secondary. palrS,. ・extending. Korea,. Japan,. LINN.. OzAKI,. lanceolate-oblong. apex. wide,. alternate and. GSJ. in. Prefecture・. :. curvlng;. slightly. wide. 16090.. teeth; sickle-shaped in the middle veins. secondary. 16161;. Clethra. hokiana. (Pl. 7,丘gs. PP. grows. Clethraceae. Genus. NSM. widely. 16028,. 16011,. Family. Holotype:. 35. Flora. U.S.S.R.. southeastern NSM PP. collections:. Tatsumitoge. forking the. the. at. sinus. bottoms. A. margln: ; qua-. B. ≧r. 0 Fig.. 7.. Venation. characters. of. Clethra. ZmT. 0. 1cm hokiana. n・. sp・. NSM. PP. 16087・. ,I.
(18) 36. K.. ternary. veins. veinlets. ortbogonal or. simple same. almost 0.6 to. to. nearly. the. from. extending. proximal. then. end,. ; ultimate. meshes the. in. midvein. trapezoidally,.. expanding. long.. 2cm. Discussion: symmetric. These. leaves. foliar. oblong. are. shape,. venation, distinguishable from. is. elliptical foliar shape,丘ner are. new. specimens et WILS. and. C.. and. NSM. FRANCE.. PP. teeth. of. and. veins. in the. new. narrowly・ This petiole.. Slender. to. the・. margln,.. This. C.. extant. in central growing 16086-16091, 16117;. 16084,. in. serrulate. NATHORST. the. Clethra. of. tertiary. spaced. maximoviczii. leaves. those. petiole, uniformly. sickle-shaped the. to. similar a. regulary. and C.. to. similar. fargesii. Collections:. closely. baying. semicraspedodromous species. 2 to 4丘ft血order. reticulate, enclosing branched; petiole丘ne,. once. vidth. OzAIくI. REHD.. monostachys China. western. GSJ 4803;ーTPM. 120, 365A,. 458.. Clethra. NATEORST. maximoviczii. (Pl. 2,丘g.. 5, pl. 3,丘g.. 5). Clethera. NATrIORST, 1883, p. 51, pl. ll,figs: 18120. TANAI, maximoviczi 3, text一点gs. 6 i, j. 丘g. 2, pl. 8,丘g. 1, pl. 9,丘g. 3, pl. 10,丘g.. Discussion NATEORST. in the. oblong. marginal. teeth,. and. is closely. similar. in southern. Hokkaido. tertiary tbrougb. runnlng. to. the. middle GSJ 4794,. and. closely. Korea,. fossil leaves. Enkianthus. Tripetalel'a Enkianthus. Enkianthus. Discussion TANAI, This. which species. :. was. the. marginal. ones. modern of the. growing'. ; many. marginal. teeth. LouR.. TANAI,. TANAI. (NATHORST). almquistii. (NATⅢORST). 1,. 1976,. p.. 2) 338,. pl. 4,軸.. 4, pl.. 5,. figs.. 1, 2, pl.. 5k.. 2, text-丘g. almquisti koreanus. apices. but. Ericaceae. (pl. 3,丘gs. almquisii. the. Zucc.. et. China,. uniformly This. veins.. of the teeth. 4818; TPM 157.. Genus. 8,丘g.. the. reach. venation, SIEB.. and from. maximouiczii. tertiary. spaced. C. barbinervis. extant. Family. Enkianthus. Clethra. semicraspedodromous. of Japan, different somewhat. of the. the. Collections:. shape,. to. referable. Kyushu. to. veins. are. regularly. fossil leaves. of the. venation of the. leaves. foliar. serrulate. species. Fragmentary. :. 1976, p. 338, pl. 7,. NATIiORST,. 1883,. HuzIOKA,. Three. leaves. discussed. is closely. 1972,. akin. p.. p.. 53, pl. 9,figs. 6, 12, 13. 10,fig. 8.. 72, pl.. are referred to Enkianthus almquistii (NATIIORST) by TANAI on basis (1976) the of the Mogi speciemns.. to. the. extant. E. subsessilis. (MIQ.) MAKINO. growing.
(19) Late. in central. Honsbu. Collection:. NSM. 16220,. Genus. Rhododendron. district. PP. Rhododendron. hokiense. LINN.. OzAKI,. (Pl. 5,丘g. TPM. Holotype: Tottori. 皿ura,. tor. tat. Leaves. (estimated). an. loops. loliar. shape, and HuzIOKA minasense. Protodilatatum. new. Formation. varieties growing Collection:. ; quaternary. but. leaves. imperfect. are. ONOE. et. in. (1961) to. similar. the. et. ONOE. ln. is. with. making upper. ; ultimate. part. veinlets by. the. distinguishable. from. R.. is also. and. R.. midvein. Rhododendron. from. separable foliar shape・. elliptical. kaempferi. PLANCtI.. This and. its. 200, 263.. TANAI. Protodilatatum. TANAlet. These. leaves. in the. rbombate. ONOE,. TPM. 1961,. p.. to. are. referred foliar shape. 55, pl.. Family Genus. and. camptodromous to. Diospyros. Ebenaceae LINN,. Diospyros nordquistii (Pl. 8,丘g.. NATHORST 8). 18,figs.. Rhododendron. 495.. 242, 451,. ONOE. et. 2). is similar R. protodilatatum 心f secondary veins. to in Honsbu Kyushu MIQ. and its varieties growlng leaves in 3's. Rhododendron have which rhombate Collection:. straight. in Japan. TPM. protodilatatum :. the. extant. (Pl. 5,丘g.. Discussion. developlng. to. referable. This species venation character. (1974) in acute apex et UEMURA TANAI. area,. oriented. veins randomly branched. once. than. more. in. 2・5. to. loops. marglnal intercostal. intersecondaries. some. ;. Rhododendron. TANAI. Saji一. 2・3. and. formlng. up,. percurrent. weakly. area. long. (estimated). 4cm. to. curvlng. gently. veins. is closely. species. Rhododendron. Miocene)・. (Late. missing, margin entire ; midvein 5 diverglng from the palrS, about. veins. 50o,. simple. well, Two. Discussion:. base. acute,. in marglnal. rather. preserved. not. SPeCies. 6). Tochiwara. elliptical, 3.7. ; secondary. of 45o to tertiary veins;. angle. marglnal of lamina,. Upper. apex. wide,. Slightly sinuous. tertiary. R.. 200.. new. Prefecture.. Description: cm. 37. Flora. Tatsumitoge. of Japan. 16217, 16301.. Kinki. to. Miocene. the and. extant some. 2, 4.. Protodilatatum small R. other. number dilatatum species. of.
(20) K.. 38. Dios如′ros. NA,l1-IORST, nordqvisii fig. 14, pl. 3). TANAl,. ing. 1883. OzAKI. 51, pl.. p.. 1976,. p.. 1,. 8,丘g.. 339,. 9,fig.. pl.. 1, 2,. 14,五gs.. pl.. 10,fig.. 7, pl.. 1.. 4, 5, (exclud(excluding pl.I. 6,丘g.6) Diospyros. Hu. miokaki. Discussion. by. secondary et CHANEY TANAI. the. shape,. My. also. D.. by. show. a. teeth. on. minute. THUNB.. D.. and. slightly small. NATHORST. 3) have丘ne. observation some. kaki. round. (excluding負g.. 3)・. nordquistii NATIiORST differnt from the type. somewhat base and. leaves丘gured. (1938, pl. 46,負g.. 6) has. extant. Two. veins.. (1976).. pl. 6,fig. to. wider. are. 2. Diospyros. to. resemble. leaves. foliar. 1,. 72. pl. 46,figs.. p.. are. speciemns My character.. venation in the speciemens Hu. 1938,. These. :. in=the. the. CIIANEY,. et. teeth. the. on. margin. reported margin. D・. the. SEIB.. japonica. of. 3) and. asmentioned TANAI. by. specimen. number. pl. 14,fig.. (1883,. (1976,I. is akin nordqvistii inZucc. both growing. et. Japan. Collections:. NSM. PP. TPM. 16194;. 291.. Family. Styracaceae. Genus Styrax. LINN.. Styrax. (Pl. 8,丘gs. Styrax. TANAI,. 1976,. Several. leaves. PrOtOjaponica Discussion. :. remotely. and. brocbidodromous. japonica. irregularly. and. the. SIEB.. et. Collectioyts:. 340,. p. are. PP. 2,fig・. pl.. acute. TPM. 16195;. Genus. leaves to. of the. southwestern. irregulary. spaced. Collection:. Chionanthus. extant. Japan, marglnal. NSM. PP. Korea. foliar. shape・. livlng. the. 5-... 331.. LINN.. ll, Fig.. 5C). LINDL.. China teeth. is closely. and. in. et. the. Fraxinus. PAXTON foliar. brochidodromous. 16223.. Genus. to. by-. TANAI. sp.. retusus. minute. n.. elliptic. similar. 302, 324,. fragmentary,. and. m,. Oleaceae. Chionanthus. leaf, though. 6. protojaponica teeth,. marglnal. 210, 252,. (Pl. 8,丘g. This. Styrax. to. Chionanthus. :. text-丘gs・. species is closely in Japan and China・. Family. Discussion. 4.. This. growing. NSM. 5, 6). referred. sapced. venation. Zucc・. TANAI. protojaponica. LINN.. similar. to. the. young'. ln. Central. growing. shape,. punctate. venation.. and-.
(21) Late. Fraxinus. Tatsllnitoge. M.ioceニe. TANAI. k-yamadae. (Pl. Fraxinus. k-yamadai. Discussion. TANAI. This. :. TANAI. yamadae. complately. NI. and. is similar. samara. to. China,. F.. Of. NuTT.. TPM. that. veins. Oblong. randomly teeth. marglnal. F・ sanzugau)aensis foliar shape, and are. similar. to. of the Collections:. number. 4,. N・. Fraxinus,. the. in a. in. the丘ne. veins. 4840;. GSJ. size.. 357,. p・. shape,. 10 flgS・. pl・. 4,. 5, text-Ag.. 6.. tertiary. apex,. acuminate. base. area. the and reach of the The fossils are to of Fraxinus. similar in the irregulary spaced marginal teeth. irregulary. and. TPM. TANAI. leanets. marglnal. NSM. N・. et. are. and. are. TANAI. 2,丘gs.. serration PP. SuzuKI,. teeth.. marglnal. and. N.. 147,. pl・. Fraxinus. to. SuzuKI. to. pl.. the. of F.. those. 25, flgS.. 2,. 3. TANAI. u)akamatsuensis. Among. veins・. similar. et. extant. species of HARA apertisquamifera. venation. 16196.. Bignoniaceae. Paulownia. Paulownia. et. 6). 1963,. secondary. 1614l,. SIEB.. et. in Academia. 8, Fig. sinica,. Zucc. TAO. shanwangensis. (Pl. 9,丘g. TAO,. 1,. similar. closely. Genus. shanwangensis. spaced. 507.. fossils. in foliar shape,. text一点g. 76.. F.. northeastern. and. shape. Japan,. 10). 1974,. intercostal. Family. Paulownia. Of. My. et UEMURA. 8,丘g.. pl.. cf. wakamatsuensis. These. :. Collection:. in the. foliar. species UEMURA. et. secondary. these. base.. the wider LINGELSH. Of RupR.. k-・. different in the tertiary veins. My specimens somewhat F. sieboldiana BLUME in the small foliar size, small extant. waka7natSuenSis. SuzuKI. in. mandshurica. UEMURA,. (Pl.. Discussion. Fraxinus. to. similar. are. Fraxinus. Fraxinus. spaethiana. HuzIOKA. inequilateral. HuzIOKA but. F.. America. 2,丘g.. et. oriented indicate. that. extant. sanzugau)aensis. HuzIOKA. sanzugawaensis :. F.. Of North. (Pl.. Discussion. is. samara. 6.. 450.. Fraxinus. Fraxinus. 13, flg.. 47, pl.. p.. slightly different. China,. western. oregona. and Collection:. of the. is. SuzuKI. 4). 1965,. but. 39. N.. et. preserved. SuzuKI,. those. LINGELSH.. inopinata. 8,丘g.. SL)-ZUKI,. N・. et. Flora. 5A) 1978,. p・. 155,. pl.. 138,. pl.. 139,. pl. 141,.
(22) 40. Oz.AKI. K.. Discussion: TAO. shanwangensis foliar shape,. and. P・. (THUNB・). tome71tOSa. fragmentary.1如ge. one. only. from. reported venation. Collection:. TPM. Widely. Tottori. in. the. the. extant. Asia・. Caprifoliaceae. hokianum. OzAKI,. 1, 2, 5-7;. (Pl. 9,丘gs. TPM. similar. ih. cultivating. to. Heptacodium'\ RE出).. Heptacodium. mura,. specimell. China. of. ,eastern. Genus. :. is. 309. Family. Holotype. flora. Paulou)nia. to. tis 'referable. Sahnwang. This. character.. STEUD・. leaf. Miocene.. new芦peCies. Figs.. 8A-E). 454・ノupper Tocムiwara▲Fわrmation. (Late. Miocene).. saji_. Prefecture.. Des?ription:..Leaves. 7. ovat?,. 9cm. to. (e,stimated)long. 4.5cm. to. andて3.,5. base. apex. missing, round, mar$1n entire ; primary separatVeins trinerved \base be′low laterals, the l畠mina, of thicker then・ the ・ngl了uit midvein extending an hlidvein 如raight; 1ateiaトprlmaries making angle the 20o i血the with about wide・. bas°.thenJ turhnlng up and course betwe占n themidvein. dive、rging basal. from. m'argin,. between. the. the. along. branched. ;・ tertiAry. prlmaries to. area. the. shallow grooves end, some five calyx-lobes spatulate,. small. loops 1. 6 to. to. somewhat similar from those leaves petiole. the. by margln. not. long,. tubes extant. of the are. calyx-tube. twice. are. most ln. and血er and. weak. Outside. of the. species.. in. reticulate,. once. to. twice. and. These. fossil calyxes. have. veins.. but. fossil. this. of the distal. resemble. end. those. living. entire is venation. trinerved leaves. wbicb. are. separable. by. species. by. the but. calyx-lobes, compare. the. from. the. the. cha-. run. and. of. coarse. Heptacodium. Veins. prlmary. base. rounded,. in foliar shape,. my. prlmaries. distal. Verticillate. vei血s making. the. Clematis,. main. apex. China These. lateral. wider. to. the. at. surface.. marginal. Central. five number. longer. the wide,. similar. eaSt9rn. baying. the. orthogonal. in diameter. on. apd 3mm nexuous, sl・ightly.. of Smilax. and. nearly loops. making. generally. 4mm. to. long. racteristic secondary and tertiary veins. The to seem to three calyxes belong shape. 3. longitudinally. 8mm:. Veins,. those. meshes. the. reticulate. extending. veins,. veinlets. to. long.. cm. growlng. prlmary. secondary. ; ultimate. 6mm. other,, midvein. trinerved. reticulate,. the tertiary. runnlng. AIRY-AtIAW. orthogonal. among. reticulation ; margln enitre. leaves Discussion: These. along. oneL_third brochidodromous,. veins. Secondary. veins. making. each. vLeins. midvein. petiole short, about Calyx-tube turbinate, about. margln,. ;. the. ;. }asminoides. margln. inner. the; 'midvein. among. ; quaternary. margln. connecting. the. and. to. parallel. the prim':aii占s占t.an 50o to 60o, parallel angle of about duplex -loops in lateral 'making ar甲, rather random. perpendicular marglnal. running. to genus. turbinate. the. those. caryx-. of Abelia,. the but.
(23) Miocene. Late. A. 41. Flora. Tatsumitoge. c塵. D. ′ ̄ー\. Jシ. L-+. 転‥‥... O_、5mm. ㌔. -J. 5mm. /. 叔鼠. 望享.  ̄. /. B I._-. /. 0.=.・. J. 5cm. if-i /. fl. E. \. 454. Venation. (Holotype),. of Heptacodium characters 494, D. NSM PP C. TPM. Genus Lonicera. Lonicera. Protojaponica (Pl. 6,丘g.. Lonicera. ProtOjaPonica. Lonicera the. プon. from extant. by. TANAI. Well. Discussion: the. margln,. hokianum E・. 16227,. 1961,. ONOE,. et. p・. TPM. to. L・. TANAI. et. 57,. ・ゝ\. 494・. Sp.. venation・ TANAI et. T血is species. aヰdthe. LIN由.. Viburnum. 5・. ∫. to. relationship. protojaPonica. Vibityn、um・. 454,. ONOE. 18,丘g・. pl・. of Okayama L/ gracilipes MIQ.盲ar. in the foliar shape PP 16003; GSJ 4767・ collections: NSM. Genus. TPM. 16115・. 5). Prefecture・. the. Onbara且orule. B・. A,. sp・. PP. LINN.. leaves has close preserved foliar shape, brochidodromous. oblong and is referable. n・. NSM. distal end of calyx-tube・ 16155, 15227; GSJ 4800;. in the wider ar弓differenit NSM PP collections・.. し. 5mm. 0 8.. m. `..〕 沙 感. ). Fig.. im. and. ONOE. the. genus. fine. reported. is similar. venation. hairs. to. character・. the.
(24) 42. K.. OzAKI. (Pl. 9,丘g. Discussion. Only. :. Viburnum. furcatum. secondary. veins, wide. percurrent. tertiary. the. middle. of the. has. weakly. BLUME. veins lamina・. that. the. nearly. Patrinia. hokiana. Tottori. mura,. Upper. closely. spaced. midvein. along. (1961) is fragmenta,y. and. to. the. the. Just. marglnal. veins. situating. in. oval,. 4. 3;. new. Fig.. species. 9). Formation. enCloslng. making. the. (Late. Miocene).. Saji-. wide, semiellipsoidal flat or slightly depressed. and. semiclub-shaped, on. 5mm. long. 3・5. to. and 4mm. 3.5. round. on. to. wide;. 4mm. wide_. main. seed,. of the or. 4 to. long. vein sending to the off many veins margin,I irregular reticulation ; margin Seed small, entire. 1・5 to 2・5mm long wlng, 1.0 to 1.5mm and. the coarse. center. small, 5mm. to. it. then. midvein,. Juss.. Tochiwara. Winged-achenia to. elliptical. developlng. and. to. Prefecture.. Description: Wing. 489・. down. oblique. OzAKI,. (Pl・ 9,丘g.. TPM. turnning. Valerianaceae. Genus. IIolotype:. and. teeth,. 374.. Family. PatrL'nia. living. the. similar. TANAI. are. to. closely. branching. perpendicular. veins that Viburnum.. TPM. are. marglnal. protofurcatum. genus. 4797;. GSJ. trl叩gular. are. V・. leaves. dichotomously. shape,. tertiary. from. omitted Collections:. in the. basal. percurrent. be. may. fragmentary. two. 4). the. margln. front・. round. entire, at. apex,. obtuse. at. back. the base,. and. surface. smooth.. Discussion PatrinL'a. by. These. :. the. shape. closed-up and. venation. 29. winged-achenia characters. of. are. the. referred wings,. and. to. the. shape. 汁. 〔 ”. /. a. a. て u. Fig1. 9・. Venation. 5mm characters. of Patrinia. hokiana. n・. sp.. TPM. 489.. genus. of. the.
(25) Late. The. closely. P.. and. BuNGE. heteroPhylla. and the collection:. in. related. Potamogeton hokiensis. potamogeton. (pl.2,丘gs.. Discussion leaves. these fossil. This. and. pondweed. OzAKI,. 9,. speciもs. is. shape. of. elongated. wlngeS・. SPeCies. new. 9). 3,丘g・. 10, pl・. Formation. (Late. Saji-. Miocene)I. :. slightly. are. specimens. to. turnnlng. apex. and. faintly. lenticular・. FIEBER. P. berchtoldii from. species is distinguishable Eastern Asia by from reported. and. down・. three-nerved. similar to some species of Potamogeton. new. alternate, long. 5cm. to. the. of southeastern P・. and. pusillus LINN・. fossil speciemens of in the linear foliar. several. authors. several. linear. submarsed. closely. living. similar. tO. One. section. the. 4. generally. clasplng,. are. linear,. stem,. sessil,. part・. middle. Among. the. of. portion. leaves. zigzag;. extending. wide・. entire, transverse These specimens. of pondweed・. Asia,. l・5mm. to. in the. taparing margln. nervulate,. 1. bending. from the extending i.5mm wide about base. new. LINN・. Tochiwara. Upper. 4831・. Stem. DescriPtion.I. acute,. Korea. Japan,. Prefecture・. Tottori. mura,. most. Potamogetonaceae. Genus. GSJ. are. 489. Family. HololyPe:. This in. species the. of. veins. in. growing. China・. two. extant. fossil seeds. these. Juss. northern. marglnal. spaced. closely TPM. wings. the. that. (TfIUNB・). uillosa. growing. from. different. somevhat. P・. 43. Flora. suggest. characters. above-described to the living related. seeds.. Tatsumitoge. Miocene. shape.. NSM. collection,s:. PP. 16221,. Gramineae. Family Genus. UNGER. Bambusium Bambusium. sp.. (Pl. 3,丘g.. The. leaves.. Bambusium. groups・. long. (estimated). 皿entary.. midvein of the. 3 to have. leaves. The and. and. have. 5. 3). to. 4・5cm 7. to. 9 parallel. wide, 10. by. though. strong. large. between. NSM. PP. 4607,. 16225A;. GSJ. the. veins. parallel. veins. are. sized. all of. the. lamina.. collections:. di氏cult. identification. of bamboos sp・ A is characterized. leaves. A. is very of bamboos Tatsumitoge from the. The. Discussion:. 4831・. GSJ. 16222;. 4832・. strong. based. only. into. separable leaves・・. 15. both veins. to. twoJ. 20cm frag一. are. specimens in. on. sides. in the. of. the・. middle.
(26) 44. K.. OzAI(I. Bambusium (Pl. 6,点g. Discussion: long. Leaves. l・5. and. are. 2cm. to. the. PP. by. leaves. 16224,. 5. belong. ℃o11ection linear. foliar. to. shape. Collection:. leaves. and. parallel. veins. inbotb. strong. veins. in. the. the. sp.. 9) found. sometimes. Gramineae,. the. lOcm. to. LINN.. Carex. (Pl. 8,丘g. Fragmentary. 7 strong. to. 7.5. size;. 16225B.. Carex. :. small. between. veins. Genus. Discussion. rather. have. 5 to 7fine. and. 4). characterized. wide・. the midvein -sides of of the lamina. .middle Collection : NSM. B. sp.. are. and. probably. from. Tatsumitoge. the. to Carex. referable. in their. parallel venation. 16231.. NSM. PP. References AcAD・. SINICA,. PEKIN INST・ BoT・ and NANKINGrINST. GEOL. PALAEONT. and Fossil Plants III, Cenozoic of All Gruops・ Plants of China Sinica, Science Press, Peking. pls・ Academia (in Chinese).. 1978.. Fossils 149 S.. ・ENDO,. 1966.. On. description. of. 190, ∫uIくUDA,. H・H・. K・. K・. KuBO, Rept・. 1943・. A,. The. 1972・. The. Tertiary. HuzIOIくA,. K・. 1974・. The. Miocene. 5(2) K・. :. K・. fecture, H・. 即ATSUO,. Flora・ MuRAI,. NAT壬10RST, Handl.. the. of. 923-948.. Miocene. :. Japan,. Hokkaido,. with 61 :188-. N.S,. Ningyo・toge. from. Flora. Shantung. 1970・. Coll.. Akita. Floras. 1973・. pls. 1-5. Prefecture,. Univ.. 20. Palaeogene Sci・. I.. Jour.. Fac.. Sci.. Aniai・typefloras. the. and. in. 3(4) :ト105, pls. 1-18. ibid. 5(1) :ト83, pis. 1-14, the. western. end. of. Honshu,. Natn.. Late. Miocene. S°i. Mus.. Miyata. 16(4). :. Flora. and. 1883. 3-92,. Japan.. 1-16.. Prefecture,. pls.ト18. Flora of Akita. Ⅰvate. Shizukuishi of the Univ. 16(2) : 45-65,. a laflore. fossile du Japon.. Pre-. 1-ll.. KyGshG. Pt. ⅠⅠ.The of Northwestern Kanazawa Arts, Univ・ 7 : 13-39, pls. 1-ll.. Paleobotany Tech.. Contribution pls.. of Akita. 66ト738,. Floras. Coll・ Liberal. Geology. :. China.. A,. ser.. of Korea. Flora from. The. Bull.. Japan.. (Part Iト3)・ Rep・ A・G・. Okayama. Province,. Tyosen.. from. plants. K・ 1974・ The Late Miocene UEMURA, Sanzugawa and Northeast Honshu, Japan. ibid. 17(4) : 325-366, pls. Ann・. Mine,. (in Japanese).. 118-141,. of Akita. Daibo. pls.ト6. UEMURA, K・. Honsbu,. S・ 1963・. Japan. of. Soc.. 1-24,. and. Northeast --HuzIOKA,. zone. Palaent.. pls.. 7(1). Aniaiflora. Jour・ Min・. Japan・. :. Tertiary. 4,. K・. BuzIOKA・. Proc,. Deposits. A. 57. some. ser.. 月uzIOKA, ibid.. Woodwardia. the. 232. Japan. 1938・. on. Univ.. 1964・. Uranium. Surv.. I :ト147,. Notes. lmp.. K・. 1969・. R・W・. Sinica. Hokkaido -HuzIOKA, Honshu,. from Trams.. species.. Geol・. CIIANEY・. and Palaeont.. Huz10KA,. Tilia. genus new. 232 pp.. 23.. pl.. Ⅰ・ and. Prefecture, Bu,. the two. Chinese. of Cbina・. Basin, pls.. Iwata. Sakito. Prefeclure,. 18-20.. Kgl.. Svensk.. Vet-Akad..
(27) Late. 0zAKI,. K・. 1979・. Honshu, OzAKI・. K・. 1980・. Flora・. On. K・. 1961・. The. plants. from. Sci.. Fukusbima. T・ 4,. 1961.. ll(2) T・. 119-398,. :. 1970・. T・. TANAI,. 1976・ Florin. T・. 10. :. The :. pls.. T・. and. border. ONOE,. Southwest. 31-56,. pls. 1-7. Tatsumitoge. Miocene Pliocene Sci.. species of Fac. Art. Rep.. pls.ト19. in. Japan.. Jour.. Fac.. Sci.. Hokkaido. Univ.. ser.. from. Kushiro. the. Ⅲokkaido,. coal丘eld,. Japan... pis. 3-20. Pliocene. of the. revision. T・. :. 1-32.. (1920). ibid. 17(2) :. between. Prefecture,. 2. 26. sec.. Miocene characteristic and Tohoku district, Japan.. change. Oligocene且oras. 3831514, The. of. 1-95,. Tottori. of Univ.. the. 277-346,. ONOE, T. 1959. A Miocene and Field, Japan. Bull. Geol. Surv.. Coal TANAI,. Univ.. part. Neogenefloral. ibid. 14(4). and. ser.. and. southern. Natl.. 45. Late Rosales and of the C, 6(2) : 33-58, pls.ト7.. Ranales. S°i. Mus.. Flora. Flora. Yokobama. important. the. Tatsumitoge. Tatsumitoge Rep・. Urticales,. Natn.. TANAI,. TANAI,. Miocene. (I)I Sci・. Bull・. SuzuKI,. TANAI,. Late. Japan. Miocene. A. 1961・. Tottori. Flora,. described. Nathorst. by. (1883)・. 1-10.. Flora. Japan. from. the. 10(4). MioIPlioceneflora Okayama. and. Mogi pls.. :. part. northern 1-24.. from. Prefectures,. the. Ningyo-toge. Japan.. Geol.. area. Surv.. Joban. the. of. on. Japan. the Rep.. 187 :ト62,. TANAI,. pls. 1-18. and OzAIくⅠ, K. Prefecture, Western T.. pls.. TANAI,. 1977.. The. Acer. genus. Jour. Fac.. Japan.. from. S°i. Hokkaido. the. Miocene. upper. Univ.. 4,. ser.. in. 1了(4):. Tottori 575-606,.. 1-5.. T.. and. SuzuKI,. N.. 1965.. Late. Palaeont. Japan. Soc. Japan Spec. WoLFE, ∫.A. and TANAI, T. 1980. The Group,. Alaska.. U.S.. Geol.. Survey. Tertiary Paper. lO. Miocene Prof.. Paper. :. Floras. from. 1-117,. pls. Point. Seldovia llO5:. 52. northeastern 1-21.. ppリ25. Flora pls.. from. Ⅲokkaido,.. the. Kena主.
(28) Explanation in natural size, unless Museum, National Science. (All丘gures NSM. PP. GSJ: TPM. :. Geological. Survey. Tottori. Prefectural. :. of. 1,. Figs.. 2, 5, 6・. 3, 4・. 1-9. otherwise stated) Plant fossils・. of Japan・ Museum.. Explanation Figs.. Plate. Steu,artia. submonadelpha. Stewartia. hokiana. of TANAI. Plate et. ONOE・. 1 NSM. PP. 16008,. 16009・. 4771.. Fig.. 7.. Stewartia. hokiana. n・. sp・. n・. sp・. Paratypes・. Holotype・. TPM. TPM 307・. 399,. GSJ. 4772・. TPM. 478・. GSJ.
(29) K・. OzAKI. See.. II, Plate. 1.
(30) Explanation F gs.. 1. 6.. Fraxinus hokiana. Holotype.. 2.. Aralia. F. g.. 3.. SaPium. protoja♪onicum. g.. 4.. Fraxinus. sanzugawaensis. ド g.. 5.. Clethra. F g.. 7.. F. 8.. Acer nordenskioeldii Euonymus TPM sp.. F g.. 9.. Potamogeton. F. 10. Potanwgeton. F g.. ll. Sapium. F g.. 12. Rhus. g.. g.. n.. sp.. n.. PP. et. N.. 2 SuzuKI.. NSM. 156. NSM et. TPM. NATrIORST.. Plate. PP. 16096・. UEMURA.. TPM. 507・. 157. 508B.. TPM. 508.. hokiensis. NSM. TPM. SuzuKI.. NATHORST.. hokiensis. hokianum. K.. HuzIOKA. maximoviczii. sp.. TANAI. cf. u)akamatsuensis. F g. F. of. n.. sp. n.. sp.. 16245.. sp.. Holotype. Paratype.. Holotype.. TPM. GSJ. 4831・. NSM 242.. PP. 16221・. PP. 1614l,. 16196・.
(31) K. OzAKI. See.. II, Plate. 2.
(32) Explanation F gs. F. g.. F. gs.. 1, 2. 3.. Enkianthus. Bambusium. 4, 6.. g.. 5.. Clethra. F g.. 7.. Ailanthus. F. 8.. Gleditsia. 9.. Potamogeton. F. g.. F g. F gs. F. g.. 10 ll.. 12.. maximoviczii sp. TPM. Ilex. Hedera. NATI-王ORST.. sp.. (xl.5). et. GSJ. NSM. Cf王ANEY.. PP. (×1.5). 16301 TPM. (×1.5), 16217. 455A,. 4794.. 458.. Hu miosinensis n. hokiensis subcornuta. Hu. miodemersum. 3. TANAl.. (NATHORST) almquistiisp. A. GSJ 4823.. Ceratophyllum. Plate. of. sp.. HuzIOI{A. NSM. GSJ. CHANEY.. et. PP. Paratype. et. NSM. UEMURA.. 16070.. 4794. PP NSM. 16222. PP. 16010,. TPM. 39l.. B・.
(33) K.. OzAKI. See.. II, Plate. 3.
(34) Explanation Figs.. Koelreuieria. 1, 2, 7.. TPM. Fig.. 3.. Rhamnus. sp.. Fig.. 4.. Sageretia. hokiana. Figs.. 5, ll.. Figs.. 6, 9, 12. 8.. Fig.. 10.. et. CHANCY.. TPM. 287,. NSM. PP. 16047,. 4770.. GSJ. Fig.. Hu. miointegrifolia. 4. Plate. of. Berchemia. Meliosma Houenia. Holotype.. sp.. Hu. mioPoribunda. Sageretia sp.. 347. n.. hokiana. NSM. miodulcis. n.. PP. 16039.. Hu. et. sp.. CflANEY.. NSM et. PP. 16038. CIiANEY.. NSM. Paratypes.. TPM. NSM. 504.. (x2). PP PP. 16046,. 16171,. TPM. 16045. 235,. GSJ. 4770..
(35) K.. OzAKI. Sec.. ⅠⅠ, Plate. 4.
(36) Explanation Fig.. 1. Rotala. Fig.. 2. Rhododendron. hokiana. Fig.. 3. Tilia. Fig.. 4. Fig.. 5. Fig.. 6. Tilia subnobilis HuzIOKA. Rhododendron hokiense. Fig.. 7. Tilia. n.. Hu. NATI寸ORST.. n.. et. CflANEY. PP. (xl.5).. 492. TANAI. et. NSM. 5. TPM. Holotype.. protodilataium. miohenryana Parthcnocissus sp.. distans. sp.. Plate. of. TPM. ONOE. 201. TPM. (×4/5).. 16211. TPM. 167.. sp.. Holotype.. NSM. PP. 16077.. TPM. 200.. 459..
(37) K・. OzAI(I. Sec.. ⅠⅠ, Plate. 5.
(38) Explanation F gS.. 1,8. Cornus. F gS・. 2,6. Tilia. F g. 3. Cornus. F g. 4. Bambusium. F ど. 5. Lonicera. ど g. 7. Tilia. of Hu. miowarteri. et. ♪rotoja♪onicaENDO. sp.. NSM sp.. PP B.,. 6. CHANEY. TPM. NSM 121,. PP. 16071,. 16303.. 452.. 16053. Tsuga. ♪rotojaPonica miohenryana. Plate. Hu. hokiensis. TANAI et. OzAKI. ONOE.. et. C壬†ANEY.. TPM. NSM. NSM 453.. PP. PP. 16224.. 16003..
(39) K.. OzAKI. Sec.. ⅠⅠ,Plate. 6.
(40) Explanation F g.. 1.. Acantho♪anax. F g. 2.. Ilex. F gS. 3. 5.. F g.. 4.. F gS. 9.. PP Clethra. WoLFE. ”-suzukii TANAI. Clethra. hokiana. et. et n.. TANAl.. ONOE.. sp.. TPM. Paratypes.. GSJ. n.. sp.. Holotype.. NSM. PP. 4774,. NSM. PP. 16011. (×2/5).. 250. NSM. 16087. hokiana. 7. 16092.. Buxus?rotojaponica. 6, 7, 8, 10. NSM. Fig.. Kalopanax. PP. Plate. 4796.. GSJ. sp.. NSM. sp.. of. 16090.. PP. 16088,. 16084,. TPM. 365A,.
(41) K.. OzAKI. See.. II, Plate. 7.
(42) Explanation F gs. F g.. 1. 3.. Clethra. 2.. Clematis. F i. 4. F gs. 5. Fraxinus 6.. hokiana NSM. sp.. Styrax. n.. PP. k-yamadae. 7.. Acer. g.. 8.. Dios♪yros. nordqvistii. F. ど. 9.. Cαrβ∬ sp. Fraxinus. NSM. 10. F g.. ll. integerrimum. Chionanthus. et. NSM. SuzuKI.. TPM. 331, TPM. MASSAL.. NATtIORST.. PP. 16089A,. TPM. 340.. TPM. NSM. 210. 199.. PP. 16194.. 16231. HuzIOKA. sanzugawaensis sp.. N.. TANAI.. (VIV.) PP. 8. 16233. TANAI. F. Plate. Paratypes.. sp.. ♪rotojaponica. F g.. F g.. of. NSM. PP. 16223.. et. UEMURA.. GSJ. 4840.. 120..
(43) K.. OzAKI. See.. II, Plate. 8.
(44) Explanation Fig. Figs.. 1.. Heptacodium. 2, 5-7.. hokianum. Heptacodium. (×2), NSM. PP. 16115,. GSJ. 3.. Patri77・ia. Fig.. 4.. Viburnum. cf. furcatum. Fig.. 8.. Paulownia. shanu・an,gensis. Fig.. hokiana. n.. sp.. hokianum. n.. sp.. Holotype. n.. 4800. TAO.. sp.. TPM. 9 454.. Paratypes.. NSM. (×1.5).. Holotype. BLUME.. Plate. of. TPM. GSJ TPM. 489. (×4).. 4797. 309. (i/2).. PP. 16227. (×2),. TPM. 494.
(45) K.. OzAKI. See.. II, Plate. 9.
(46)
図
+3
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