• 検索結果がありません。

Fig. 16 Detail of Type II Japanese native linage. The dotted line is an enlarged tree.

Number in parentheses are the number of individuals.

Ishikawa, Nozaki (8) Ishikawa, Anamizu (20)

Ishikawa, Nagasaki (6)

0.02

Type II

Table 1 Comparison of the characteristics of each dojo loach specimen collected from the Naka-ikemi Wetland, Fukui Prefecture, Japan.

Specimen no.

No. in

Fig. 3 Date of collection Sex mtDNA lineages

Total number of vertebrae

Nuclear DNA lineages

Total number of AFLP bands

1 A 23 May 2013 Male Type II 47 Type II

2 a 23 May 2013 Female Type II 47 Type II

3 a 23 May 2013 Female Type II 47 Type II

4 a 23 May 2013 Male Type II 48 Type II

5 b 24 June 2013 Female Type II 46 Type II

6 b 24 June 2013 Male Type I 48 Type I

7 b 24 June 2013 Female Type I 49 Type I

8 b 24 June 2013 Male Type I 50 Type I

9 b 24 June 2013 Female Type I 48 Type I

10 b 24 June 2013 Male Type I 49 Type I

11 b 24 June 2013 Female Type II 47 Type II 87

12 b 24 June 2013 Male Type I 49 Type I 58

13 b 24 June 2013 Male Type I 49 Type I 55

14 c 24 June 2013 Male Type I 48 Type I

15 c 20 November 2013 Female Type II 47 Type II

16 c 20 November 2013 Male Type II 46 Type II

17 c 20 November 2013 Female Type II 45 Type II

18 c 20 November 2013 Male Type II 47 Type II

19 c 20 November 2013 Male Type II 47 Type II

20 d 24 June 2013 Male Type I 48 Type I

21 d 24 June 2013 Female Type II 47 Type II

22 e 24 June 2013 Female Type II 47 Type II

23 e 24 June 2013 Male Type II 47 Type II

24 e 24 June 2013 Male Type II 48 Type II

25 e 24 June 2013 Male Type II 48 Type II

26 e 20 November 2013 Female Type II 48 Type II

27 e 20 November 2013 Female Type II 48 Type II

28 e 20 November 2013 Female Type II 46 Type II

29 e 20 November 2013 Male Type II 47 Type II

30 f 24 June 2013 Female Type II 47 Type II

31 f 24 June 2013 Female Type II 48 Type II

32 f 24 June 2013 Male Type II 48 Type II

33 f 24 June 2013 Female Type II 47 Type II

34 f 24 June 2013 Female Type II 45 Type II

35 f 24 June 2013 Male Type II 48 Type II

36 g 20 November 2013 Male Type II 47 Type II 93

37 g 20 November 2013 Male Type II 45 Type II

38 g 20 November 2013 Male Type II 46 Type II

39 g 20 November 2013 Female Type II 45 Type II 99

40 g 20 November 2013 Female Type I 51 Type I 59

41 g 20 November 2013 Male Type II 45 Type II

42 g 20 November 2013 Male Type II 46 Type II 90

43 g 20 November 2013 Male Type II 45 Type II

44 g 20 November 2013 Male Type II 44 Type II

45 g 20 November 2013 Male Type I 49 Type I 55

46 g 20 November 2013 Male Type I 48 Type I 53

47 h 20 November 2013 Male Type II 46 Type II

48 i 20 November 2013 Female Type I 49 Type I

49 24 May 2015 Male Type I Type I 55

50 24 May 2015 Female Type I Type I 55

51 24 May 2015 Female Type I Type I 55

52 24 May 2015 Female Type II Type II 99

53 24 May 2015 Male Type II Type II 99

54 24 May 2015 Male Type II Type II 99

Table 2 Comparison of the characteristics of each dojo loach specimen collected from the Naka-ikemi Wetland, Fukui Prefecture, Japan.

Specimen no.

Date of collection

No. in Fig. 4

Standard

length (mm) Sex mtDNA

lineages

1 April 17, 2016 St. 1 47 Female Type II

2 April 17, 2016 St. 1 51 Male Type II

3 April 17, 2016 St. 1 48 Female Type II

4 April 17, 2016 St. 1 61 Male Type II

5 April 17, 2016 St. 1 49 Female Type II

6 April 17, 2016 St. 1 65 Male Type II

7 April 17, 2016 St. 1 49 Male Type II

8 April 17, 2016 St. 1 79 Female Type II

9 April 17, 2016 St. 1 70 Male Type II

10 April 17, 2016 St. 1 60 Male Type II

11 April 17, 2016 St. 1 66 Male Type II

12 April 17, 2016 St. 1 71 Male Type II

13 April 17, 2016 St. 1 88 Male Type II

14 April 17, 2016 St. 1 64 Male Type II

15 April 17, 2016 St. 1 51 Female Type II

16 April 17, 2016 St. 1 48 Female Type II

17 April 17, 2016 St. 1 43 Female Type II

18 April 17, 2016 St. 1 49 Female Type II

19 April 17, 2016 St. 1 67 Female Type II

20 April 17, 2016 St. 1 32 Female Type II

21 April 17, 2016 St. 1 62 Female Type II

22 April 17, 2016 St. 1 44 Female Type II

23 April 17, 2016 St. 1 45 Female Type II

24 April 17, 2016 St. 1 57 Female Type II

25 April 17, 2016 St. 1 50 Male Type II

26 April 17, 2016 St. 1 112 Female Type II

27 April 17, 2016 St. 1 49 Male Type II

28 April 17, 2016 St. 1 40 Female Type II

29 April 17, 2016 St. 1 56 Male Type II

30 April 17, 2016 St. 1 45 Female Type II

31 April 17, 2016 St. 1 53 Female Type II

32 April 17, 2016 St. 2 55 Male Type II

33 April 17, 2016 St. 2 31 Unknown Type II

34 April 17, 2016 St. 2 34 Unknown Type I

35 April 17, 2016 St. 2 49 Unknown Type II

36 April 17, 2016 St. 2 35 Unknown Type II

37 April 17, 2016 St. 2 35 Unknown Type II

38 April 17, 2016 St. 2 35 Unknown Type I

39 April 17, 2016 St. 2 35 Unknown Type II

40 April 17, 2016 St. 2 80 Female Type II

41 April 17, 2016 St. 2 77 Female Type II

42 April 17, 2016 St. 2 78 Female Type II

43 April 17, 2016 St. 2 94 Female Type II

44 April 17, 2016 St. 3 35 Unknown Type I

45 April 17, 2016 St. 3 80 Male Type II

46 May 30, 2016 St. 1 48 Female Type II

47 May 30, 2016 St. 1 35 Unknown Type II

48 May 30, 2016 St. 1 40 Male Type II

49 May 30, 2016 St. 1 51 Male Type II

50 May 30, 2016 St. 1 76 Female Type II

51 May 30, 2016 St. 1 35 Unknown Type II

52 May 30, 2016 St. 1 49 Male Type II

53 May 30, 2016 St. 1 35 Unknown Type II

54 May 30, 2016 St. 1 81 Female Type II

55 May 30, 2016 St. 1 81 Male Type II

56 May 30, 2016 St. 1 118 Male Type II

57 May 30, 2016 St. 1 81 Female Type II

58 May 30, 2016 St. 1 81 Male Type II

59 May 30, 2016 St. 1 81 Female Type II

60 May 30, 2016 St. 1 130 Female Type II

61 May 30, 2016 St. 1 100 Female Type II

62 May 30, 2016 St. 2 35 Unknown Type II

63 May 30, 2016 St. 2 35 Unknown Type II

64 May 30, 2016 St. 2 58 Female Type II

65 May 30, 2016 St. 2 65 Male Type II

66 May 30, 2016 St. 2 71 Female Type II

67 May 30, 2016 St. 2 72 Female Type II

68 May 30, 2016 St. 2 58 Female Type II

69 May 30, 2016 St. 2 58 Unknown Type II

70 May 30, 2016 St. 2 56 Female Type II

71 May 30, 2016 St. 2 35 Unknown Type II

72 May 30, 2016 St. 2 35 Unknown Type I

73 May 30, 2016 St. 2 62 Unknown Type I

74 May 30, 2016 St. 2 58 Male Type II

75 May 30, 2016 St. 2 97 Female Type II

76 May 30, 2016 St. 2 81 Male Type II

77 May 30, 2016 St. 2 81 Male Type I

78 May 30, 2016 St. 2 78 Female Type II

79 July 3, 2016 St. 1 24 Unknown Type II

80 July 3, 2016 St. 1 47 Male Type II

81 July 3, 2016 St. 1 58 Male Type II

82 July 3, 2016 St. 1 56 Female Type II

83 July 3, 2016 St. 1 70 Male Type II

84 July 3, 2016 St. 1 77 Female Type II

85 July 3, 2016 St. 1 81 Female Type II

86 July 3, 2016 St. 1 90 Male Type II

87 July 3, 2016 St. 1 78 Male Type II

88 July 3, 2016 St. 1 92 Male Type II

89 July 3, 2016 St. 1 112 Female Type II

90 July 3, 2016 St. 1 121 Female Type II

91 July 3, 2016 St. 2 64 Female Type I

92 July 3, 2016 St. 2 28 Unknown Type II

93 July 3, 2016 St. 2 32 Unknown Type II

94 July 3, 2016 St. 2 33 Unknown Type II

95 July 3, 2016 St. 2 29 Unknown Type II

96 July 3, 2016 St. 2 34 Unknown Type II

97 July 3, 2016 St. 2 26 Unknown Type II

98 July 3, 2016 St. 2 44 Unknown Type I

99 July 3, 2016 St. 2 33 Unknown Type II

100 July 3, 2016 St. 3 83 Female Type I

101 July 3, 2016 St. 3 84 Male Type I

102 July 3, 2016 St. 3 96 Female Type I

103 August 14, 2016 St. 1 35 Unknown Type II

104 August 14, 2016 St. 1 35 Unknown Type II

105 August 14, 2016 St. 1 103 Female Type II

106 August 14, 2016 St. 1 110 Male Type II

107 August 14, 2016 St. 1 81 Male Type II

108 August 14, 2016 St. 1 53 Female Type II

109 August 14, 2016 St. 1 115 Male Type II

110 August 14, 2016 St. 1 53 Female Type II

111 August 14, 2016 St. 1 110 Male Type II

112 August 14, 2016 St. 1 118 Male Type II

113 August 14, 2016 St. 1 80 Female Type II

114 August 14, 2016 St. 1 115 Female Type II

115 August 14, 2016 St. 1 133 Male Type II

116 August 14, 2016 St. 1 98 Female Type II

117 August 14, 2016 St. 1 106 Male Type II

118 August 14, 2016 St. 1 129 Female Type II

119 August 14, 2016 St. 2 33 Unknown Type II

120 August 14, 2016 St. 2 40 Male Type II

121 August 14, 2016 St. 2 37 Unknown Type II

122 August 14, 2016 St. 3 102 Female Type II

123 August 14, 2016 St. 3 70 Male Type II

124 August 14, 2016 St. 3 52 Female Type II

125 August 14, 2016 St. 3 97 Female Type I

126 August 14, 2016 St. 3 114 Male Type I

127 August 14, 2016 St. 3 103 Male Type II

128 August 14, 2016 St. 3 94 Male Type I

129 August 14, 2016 St. 3 108 Male Type I

130 August 14, 2016 St. 3 110 Female Type I

131 October 23, 2016 St. 3 Type I

132 October 23, 2016 St. 3 Type I

133 October 23, 2016 St. 3 Type I

134 October 23, 2016 St. 3 Type I

135 October 23, 2016 St. 3 Type I

136 October 23, 2016 St. 3 Type I

137 October 23, 2016 St. 3 Type I

138 October 23, 2016 St. 3 Type I

139 October 23, 2016 St. 3 Type I

140 October 23, 2016 St. 3 Type II

141 July 17, 2017 St. 3 Type I

142 July 17, 2017 St. 3 Type I

143 July 17, 2017 St. 3 Type I

144 July 17, 2017 St. 3 Type I

145 July 17, 2017 St. 3 Type I

146 July 17, 2017 St. 3 Type I

147 July 17, 2017 St. 3 Type I

148 July 17, 2017 St. 3 Type I

149 July 17, 2017 St. 3 Type I

150 July 17, 2017 St. 3 Type II

Table 3 Sampling locations, number of individuals and Accession number of Misgurnus anguillicaudatus and its related species, used in the phylogenetic analyses.

NO. No. in Fig. 7

Locality

N Accession No.

Prefecture/Country City, Town, Lake or River

1 1 Hokkaido Kutcharo Lake 8

2 2 Nayoro 2

3 AB473261

4 AB473262

5 AB473263

6 3 Memanbetsu 4

7 4 Abashiri 15

8 5 Kotanbetsu 1

9 Furano AB473266

10 AB473267

11 6 Hokuryu 6

12 7 Sorachi District 4

13 8 Monbetsu 3

14 Atsuma AB473279

15 AB473267

16 Mukawa AB473267

17 Kyowa AB473276

18 AB473277

19 AB473267

20 Imakane AB473274

21 AB473275

22 AB473267

Misgurnus anguillicaudatus

23 9 Jyunsai Lake 31

24 10 Hokkaido Onuma Lake 18

25 Aomori Hirakawa AB473266

26 Towada AB473274

27 AB473267

28 Fujisaki AB473266

29 Goshogawara AB473277

30 AB473281

31 Tohoku AB473292

32 11 Yokohama 11

33 12 Shichinohe 3

34 Iwate Oshu AB473277

35 Hanamaki AB473267

36 Miyagi Yamamoto AB473301

37 AB473274

38 Imaizumi EF508556

39 EF508557

40 Natori AB473266

41 AB473267

42 13 8

43 14 Toyosato 7

44 Akita Hago AB473281

45 Kazuno AB473267

46 Yokote AB473266

47 Yamagata Tsuruoka AB473281

48 Fukushima Yabuki AB473266

49 Sugakawa AB473308

50 Fukushima Sugakawa AB473309

51 Ibaraki Kashima AB614357

52 Tochigi Otawara AB473281

53 Sakura AB473312

54 AB473313

55 AB473314

56 Gunma Kawaba AB473317

57 Saitama Hanno AB473279

58 Chiba Minamiboso AB473279

59 Niigata Niigata AB473266

60 AB473281

61 AB473279

62 AB473288

63 Nagaoka AB473267

64 15 Sado 4

65 16 Tokamachi 11

66 Toyama Himi AB473325

67 17 Ishikawa Wajima 42

68 18 Anamizu 20

69 19 Nagasaki 9

70 20 Nozaki 8

71 21 Neyamu 1

72 22 Kouda 29

73 23 Enome 4

74 24 Hannoura 8

75 AB599978

76 Ishikawa Nanao AB614359

77 25 Shinbo 21

78 26 Akaura 3

79 Hodatsu AB473274

80 AB473331

81 AB473333

82 Kanazawa AB473288

83 AB473326

84 AB473329

85 Komatsu AB473330

86 27 Fukui Tsuruga, Naka-ikemi Wetland 12

87 Tsuruga, Ikenokouchi 1

88 28 Fukui 3

89 Echizen AB898249

90 29 1

91 AB473266

92 AB473336

93 30 Kurokawa 5

94 31 Obama 1

95 AB899674

96 AB899684

97 Katsuyama AB473334

98 32 1

99 Aichi Owariasahi AB473266

100 Yatomi AB473279

101 33 Anjo 3

102 34 Aichi Inuyama 1

103 35 Nagoya 1

104 Gifu Gujo AB473301

105 36 Kiso River 1

106 37 Kaizucho 3

107 Shiga Azuchi AB473281

108 Higashiomi AB473301

109 AB473350

110 Kyoto Kyotango AB473301

111 38 9

112 39 Miyazu 2

113 40 Maizuru 1

114 Nara Yamazoe AB473359

115 41 Kashihara 2

116 Hyogo Kato AB473267

117 Wakayama Minabe AB473362

118 AB473363

119 AB473365

120 42 Kujinokawa 1

121 Tottori Oyama AB473368

122 Okayama Kasaoka AB473267

123 43 Maniwa 3

124 Yamaguchi Ube AB473383

125 Hagi AB473384

126 Hofu AB473386

127 Iwaguni AB614358

128 44 Yamaguchi Tawarayama 1

129 Kagawa Takamatsu AB473396

130 Ehime Seiyo AB473398

131 Kochi Haruno AB473267

132 Fukuoka Fukuoka AB978285

133 Miyako AB473400

134 Saga Yoshinogari AB473401

135 Nagasaki Tsushima AB473402

136 Miyazaki Ebino AB473406

137 45 Kagoshima Ichikikushikino 2

138 China Hejiang AY625700

139 Russia Sakhalin JN858856

140 JN858857

141 JN858858

142 JN858859

143 AB473261

144 AB473277

145 AB473308

146 EF508557

147 Amur River JN858854

148 Lotos Lake JN858860

149 Poland Warmia-Masuria EF508561

Misgurnus nikolskyi

Misgurnus fossilis

Misgurnus dabryanus

150 China Huangshan AY625701

151 Russia Amur River EF508565

152 47 Kyoro Maizuru 1

153 Korea EU670767

154 Russia Amur River JN858850

155 Korea EU508498

156 Germany Niedersachsen EU508508

157 Slovakia Presov AY887851

158 India West Bengal EU508583

Misgurnus mohoity

Cobitis lutheri

Cobitis taenia

OutGroup

Sabanejewia balcanica

Pangio pangia

Table 4. The list summarizes the results of D-loop Haplotypes, Nuclear DNA linages and phylogeny Groups.

No. mtDNA linages

D-loop Haplotypes

Nuclear DNA

linages Groups N

1 Type I Haplotype III Group 1 1

Haplotype I Type I / Type I Group 3 1 Haplotype II Type I / Type I Group 3 6

2 Type I Group 1 2

3 Type I Group 3

4 Type I Group 3

5 Type I Group 3

6 Type I Group 1 2

Group 3 2

7 Type I Haplotype I Type I / Type I Group 3 4

Haplotype II Type I / Type I Group 3 11

8 Type I Group 1 1

9 Type II Japanese native

10 Type II Japanese native

11 Type I Haplotype III Group 1 3

Type II Haplotype V Japanese native 3

12 Type II Japanese native 4

13 Type II Japanese native 3

14 Type II Japanese native

15 Type II Japanese native

16 Type II Japanese native

17 Type I Group 3

Misgurnus anguillicaudatus

18 Type II Japanese native

19 Type II Japanese native

20 Type II Japanese native

21 Type I Group 3

22 Type II Japanese native

23 Type I Haplotype III Type I / Type II Group 1 11

Type II Type II / Type II Japanese native 20

24 Type I Haplotype III Type I / Type II Group 1 7

Type II Type II / Type II Japanese native 11

25 Type II Japanese native

26 Type II Japanese native

27 Type II Japanese native

28 Type II Japanese native

29 Type I Group 1

30 Type II Japanese native

31 Type II Japanese native

32 Type II Japanese native 11

33 Type II Japanese native 3

34 Type I Group 1

35 Type II Japanese native

36 Type II Japanese native

37 Type II Japanese native

38 Type II Asian continent origin

39 Type I Group 4

40 Type II Japanese native

41 Type II Japanese native

42 Type II Japanese native 8

43 Type II Japanese native 7

44 Type II Japanese native

45 Type II Japanese native

46 Type II Japanese native

47 Type II Japanese native

48 Type II Japanese native

49 Type II Japanese native

50 Type II Japanese native

51 Type II Asian continent origin

52 Type II Japanese native

53 Type I Group 6

54 Type I Group 6

55 Type I Group 6

56 Type II Asian continent origin

57 Type II Asian continent origin

58 Type II Asian continent origin

59 Type II Japanese native

60 Type II Japanese native

61 Type II Asian continent origin

62 Type II Japanese native

63 Type II Japanese native

64 Type II Japanese native 4

65 Type II Japanese native 11

66 Type II Japanese native

67 Type I Hap A Group 1 4

Type II Hap B Type II / Type II Japanese native 38

68 Type II Japanese native 20

69 Type II Japanese native 9

70 Type II Japanese native 8

71 Type II Japanese native 1

72 Type I Hap A Type I / Type II Group 1 4

Type II Type II / Type II Japanese native 25

73 Type II Japanese native 4

74 Type II Japanese native 8

75 Type II Japanese native

76 Type I Group 1

77 Type I Haplotype XI Type I / Type I Group 6 20

Type II Hap C Type II / Type II Japanese native 1

78 Type II Japanese native 3

79 Type II Japanese native

80 Type II Japanese native

81 Type II Japanese native

82 Type II Japanese native

83 Type II Japanese native

84 Type II Japanese native

85 Type II Japanese native

86 Type I Hap D Type I / Type I Group 2 6

Type II Hap B Type II / Type II Japanese native 6

87 Type I Group 2 1

88 Type II Haplotype VII Asian continent origin 3

89 Type I Group 1

90 Type II Japanese native 1

91 Type II Japanese native

92 Type II Japanese native

93 Type II Asian continent origin 5

94 Type II Japanese native 1

95 Type II Japanese native

96 Type II Japanese native

97 Type II Japanese native

98 Type II Japanese native 1

99 Type II Japanese native

100 Type II Asian continent origin

101 Type II Asian continent origin 3

102 Type II Japanese native 1

103 Type II Japanese native 1

104 Type II Japanese native

105 Type II Japanese native 1

106 Type II Japanese native 3

107 Type II Japanese native

108 Type II Japanese native

109 Type II Japanese native

110 Type II Japanese native

111 Type II Japanese native 9

112 Type II Japanese native 2

113 Type II Asian continent origin 1

114 Type II Asian continent origin

115 Type II Haplotype VII Asian continent origin 2

116 Type II Japanese native

117 Type II Japanese native

118 Type II Japanese native

119 Type II Japanese native

120 Type II Japanese native 1

121 Type II Japanese native

122 Type II Japanese native

123 Type II Japanese native 3

124 Type II Japanese native

125 Type II Japanese native

126 Type II Asian continent origin

127 Type II Japanese native

128 Type II Japanese native 1

129 Type II Japanese native

130 Type II Japanese native

131 Type II Japanese native

132 Type II Japanese native

133 Type II Japanese native

134 Type II Japanese native

135 Type II Japanese native

136 Type II Japanese native

137 Type II Japanese native 2

138 Type II Asian continent origin

139 Type I Group 1

140 Type I Group 1

Misgurnus nikolskyi

141 Type I Group 1

142 Type I Group 3

143 Type I Group 3

144 Type I Group 1

145 Type I Group 5

146 Type I Group 4

147

148

149

150

151

152 1

153

154

155

156

157

158

Misgurnus fossilis

Misgurnus dabryanus

Misgurnus mohoity

Cobitis lutheri

Cobitis taenis

OutGroup

Sabanejewia balcanica

Pangio pangia

Table 5 Sampling locations, mtDNA linages, D-loop Haplotypes and number of individuals of Misgurnus anguillicaudatus, used in the phylogenetic analyses.

NO. No. in Fig. 7

Locality mtDNA

linages

D-loop

Haplotypes N Prefecture/Country City, Town, Lake or River

1 1 Hokkaido Kutcharo Lake Type I Haplotype I 6

Type I Haplotype II 1 Type I Haplotype III 1

7 4 Abashiri Type I Haplotype I 4

Type I Haplotype II 2

11 6 Hokuryu Type I Haplotype III 3

Type II Haplotype V 2

23 9 Jyunsai Lake Type I Haplotype III 1

24 10 Onuma Lake Type I Haplotype III 2

67 17 Ishikawa Wajima Type I Hap A 1

Type II Hap B 2

72 22 Kouda Type I Haplotype III 1

Type I Hap A 1

77 25 Shinbo Type I Haplotype XI 4

Type II Hap C 1

86 27 Fukui Tsuruga, Naka-ikemi Wetland Type I Hap D 6

Type II Hap B 6

88 28 Fukui Type II Haplotype VII 1

115 41 Nara Kashihara Type II Haplotype VII 2

Misgurnus anguillicaudatus

Table 6 Species and DDBJ accession numbers of DNA sequences used in the phylogenetic analyses of D-loop.

Species Haplotypes Accession number

Misgurnus anguillicaudatus Haplotype I AB306717

AB306721 AB306723

Haplotype II AB306724

AB306725

Haplotype III AB306729

AB306730

Haplotype IV AB306731

Haplotype V AB306733

Haplotype VII AB306762

Haplotype VIII AB306783

Haplotype X AB306786

Haplotype XI AB306789

AB306790

Haplotype XII AB306791

Misgurnus nikolskyi AB242171

Misgurnus dabryanus Hap 1 LC025649

Misgurnus mohoity Hap 5 LC025653

Cobitis striata striata AB054125

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