サラブレッド雌馬の生産性向上のための交配から妊娠早期までの繁殖管理の改善に関する研究

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(1)Title. サラブレッド雌馬の生産性向上のための交配から妊娠早期までの繁殖管理の改善に関する研究( 本文(Fulltext) ). Author(s). 宮越, 大輔. Report No.(Doctoral Degree). 博士(獣医学) 乙第156号. Issue Date. 2018-03-13. Type. 博士論文. Version. ETD. URL. http://hdl.handle.net/20.500.12099/75235. ※この資料の著作権は、各資料の著者・学協会・出版社等に帰属します。.

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(5) ǰɤɲyɜə ............................................................................................................... 9 Ŏˀ........................................................................................................................... 10 ɸ

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(7) ʩǨuɜə .............................................................................................................. 13

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(9) ǱǜuǞȟ .............................................................................................................. 15

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(11) ˖Ǽƀ˛ ............................................................................................................ 15

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(14) ˖ǼǞȟu˖Ǽ̯ɜ .......................................................................................... 15

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(16) ʏˋˈǸ ............................................................................................................ 16

(17) ʍǺ ......................................................................................................................... 17

(18) ̤̺yɁƵuǍīɅy̔÷ ................................................................................ 17

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(20) ůŹyȤʯăyÅÉ¬Z}ǪţīʬyɝơuǍīɅy̔÷ ...................... 17

(21) 4 ˂ĘyǀÐŊǚuǍīɅy̔÷ .................................................................. 18

(22) īʬyûǚuǍīɅy̔÷ ............................................................................... 18 ii.

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(24) ʢž ......................................................................................................................... 19

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(26) ǱǜuǞȟ .............................................................................................................. 29

(27) ˖Ǽƀ˛ ............................................................................................................ 29

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(32) ǱǜuǞȟ .............................................................................................................. 51

(33) ˖Ǽƀ˛ ............................................................................................................ 51

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(35) ˖ǼǞȟZ}˖Ǽ̯ɜ ................................................................................... 51

(36) ʏˋˈǸ ............................................................................................................ 51. iii.

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(201) Ǡɜt˥b ->8E&>13:-:/E";?? yɗɉɅz

(202)

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(205) uŘŁd sZkyþzŐŒŕ̸ʨɈŚxɑwʂŐ͏ʻŐ͏

(206) ̬Ő͏

(207)

(208) SmŪŬ Ǡã̙xZb "-@1&>13:-:/E ";?? yɗɉɅz

(209) uŘŁdsW

(210) S&>13:-:/E";?? y ɗɉɅxƠ̮ÐY˄ŋuesʚȒ̤̺yƔ̿

(211) ̤̺yɁ Ƶ͂ǯʌɊɳʪȞÕ̓ďŭƣđŊɗƲtyĵʪ

(212) Ţʪxƀ gȫʪčʝ

(213) wv\ʢYsZ&>13:-:/E";?? xz̤̺āy ŢȆw˄ŋ\̔˽esWuǏžt]S ǠǰtzȔƔʆ . ̳y ÂµÅ¨¬\ɷˤ̺uesɉɊdsZ byXo \Ǡ̽ŒǞtɉɊdsWS btxǠǰŐĉw}xǠ̽ŒǞxZ` ÂµÅ¨¬y &>13:-:/E ";?? x̔gţˆȈw˖ǼzʿsZhkyɗɉɅw}ɗɉ˄ŋxq Wsǣ[xdsWwWSǰɶyɜəzǠǰyǠ̽ŒǞxZ` &>13:-:/E ";?? yɗɉɅǣ[xekyɗɉxƠ̮ÐY̤̺āy˄ŋȂˌg. 27.

(214) butVS . . . 28.

(215)

(216) ǱǜuǞȟ .

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(222). Ɣ͏ ̳

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(225)

(226) ˖ǼǞȟZ}˖Ǽ̯ɜ. ÂµÅ¨¬ʚȒȾř ȾřyʚȒ̤̺ƀ˛ues˖ǼŷǟemSô ˒̺zÛ̌ƣđŊyŪŬˏǝãÎđŊŪŬˏǝ͏Û̌ƣǠǚyÒŦþz Ǡx˦̭ȠˏǝˁʝɋWsĵʪuˏǝdmʚȒ̤̺tVSkyƣ Û̌ƣ ˾xċƗ˦̭ȠˏǝˁʝxsŪŬˏǝʿWãÎċŪŬˏ ǝ͏Û̌ƣǠǚyÒŦþz ǠÛ̌ƣ [ Ǡty &>13:-:/E";?? yǬȸˏǝedxʠƔȸØxĎɉem[ʥ]Ĵ˖ǼʿpmSʥ] Ĵ˖ǼyǬğŊɼz ótVpmSđŊŪŬˏǝtŢʪ\˓ mřĻ ćsyɕótʌɝʰtyʪʬɦɥxȫʪčʝʿpmSǰɤɲtzÛ̌ ƣđŊŪŬˏǝtzĵʪtVpm\Û̌ ͇ ˾yċŪŬˏǝǦxÏĵʪtV pmy &>13:-:/E";??-E? ċŪŬˏǝxsĵʪɨ˓emyo ɉß\Ďɉew[pmy &>13:-:/E";??-E 2;-85:3͂ɉƣɝȐZ} ďŭÒyȐÚĿ̓uesŶʟeklyɗɉɅȚ mSdxÛ ̌ƣđŊyŪŬˏǝ[ĎɊtyǓʤ %B1>-88&>13:-:/E";?? uWsŶʟ emSmʚȒ̤̺yȐÚzċŪŬˏǝ[ĎɊty̓xȐÚemyu. 29.

(227) ŶʟemS ô˒̺zkyɁƵxǯʌɊ̺ɳʪ̺kesȞÕ̺xď̷emSǯʌ Ɋ̺zĵʪďŭʌ̻esWwWʚȒ̤̺uŶʟeɳʪ̺zėƔÏĵʪ e^zėƔĵʪƣŪŬȧŧ˥bem̺uŶʟemSdxȞÕ̺z˖ǼƔx ďŭeȞÕesW̺uŶʟemS ĵʪyǬȸxqWszʌɝʰxs˦̭ȠˏǝˁʝɋWsˏǝÈˍ̐emS ˦̭Ƞɏăˏǝˁʝx#F Ã®¶ÆÉµǎʑe ¾É¬ɏăxɝʰ Ŝ[ůŹĉyʪůyǬȸxŪŬˏǝŷǟemS &>13:-:/E";?? yɗɉx̔˄ŋ˖m xãÎy̯ɜxqWs˖ mS ħʪVWzıʪyɨ˓͐

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(235) xwtʑ `mSgsy˄ŋ\

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(237) qyÄÉ¶tʿpmS qzgsy˖Ǽƀ˛̺Ŀ ÄÉ¶kesX͉qzȞÕ̺yyÄÉ¶tVSďŭƣđŊɗƲty Û̌[ľ[x̔gˈǸyȞÕ̺yyÄÉ¶ɋWsŢŠ̎ŊƑďǸ ŷǟemSáy˄ŋxqWszô˒̺ćsĿŢŠ̎ŊƑďǸx̔˽ ȂˌemS . . . 31.

(238) ʍǺ &>13:-:/E";??yɗɉɅ. &>13:-:/E";??-E? ͂đŊŪŬˏǝIċŪŬˏǝ̓yɗɉɅz% &>13:-:/E";??-E 2;-85:3͂ċŪŬˏǝIĎɊ̓yɗɉɅz % tVpmSkly &>13:-:/E";?? yþ[ɾĎdm %B1>-88&>13:-:/E";??͂đŊŪŬˏǝIĎɊ̓yɗɉɅz %tVpmS ʚȒ̤̺yȐÚ͂ċŪŬˏǝIĎɊ̓yɗɉɅz % tVpmS

(239) ŢʪxZ`ȫʪčʝu &>13:-:/E";?? y̔÷ đŊyŪŬˏǝy̡xZ`ıʪɅz tVpm͂)-.81 ̓SȫʪčʝzđŊyŪŬˏǝǦxʿm͂Û̌ƣ

(240) Ǡ͐ÒŦþ Ǡ̓S&>13:-:/E";??-E yɗɉɅzȫʪčʝʿpmıʪʞt ħʪʞtz tVpmS&>13:-:/E";??-E 2;-85:3 yɗɉɅzıʪʞtz . ħʪʞtz tVpmS &>13:-:/E";??-E? Z} &>13:-:/E";??-E 2;8-5:3 yWhy ɗɉɅxZWsıʪʞuħîʞuy̓xʏˋųəwǬƳƋz˓ w[ pmS ̤̺yɁƵZ}Ɣ̿u &>13:-:/E";?? y̔÷ &>13:-:/E";??-E? yɗɉɅ &>13:-:/E";??-E? yɗɉɅzǯʌɊ̺t ́

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(242) t ȞÕ̺t tVpm͂)-.81

(243) ̓ SŢŠ̎Ŋ ƑďǸxZWs&>13:-:/E";??-E? yɗɉɅx̤̺yɁƵzƠ̮İ dw[pmS45?=A->1@1?@ tzǯʌɊ̺uáyʞy̓xǬƳƋ\˓ m͂& ̓S &>13:-:/E";??-E? yɗɉɅz ȏt

(244) ȏt ȏt

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(247) uwƔ̿yśěxéWɗɉɅ śěem͂)-.81

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(249)

(250) Ȟ Õ ̺ t t V p m ͂ )-.81 ̓S 45?=A->1 @1?@ Z } Ţ Š ̎ Ŋ Ƒ ď Ǹ x Z W s &>13:-:/E ";?? -E? 2;-85:3 yɗɉɅx̤̺yɁƵzƠ̮İdw[pmS &>13:-:/E";??-E 2;-85:3 yɗɉɅz ȏt ȏt ȏãÍt uw45?=A->1@1?@ tzÄÉ¶̓tƋz˓ w[ pmSŢŠ̎ŊƑďǸtzô˒̺yƔ̿z &>13:-:/E";??-E 2;8-5:3 yɗɉɅuyǬƳw̔÷z˓ w[pmS klyƔ̿ÄÉ¶xZ`ǯʌɊ̺uɳʪ̺Z}ȞÕ̺y &>13:-:/E";??-E? 2;-85:3 ȂˌguǬƳƋz˓ w[pmS. 33.

(251) ( u &>13:-:/E";?? y̔÷. ( zđŊŪŬˏǝǦuċŪŬˏǝǦxȭŶekyŠĠxô˒̺ ÍǢʞʒǈʞZ}ìÎʞxĢďemSmċŪŬˏǝǦy ( x͍ ǯȮʞu ãÍʞuxĢďemS( xĢďemʞu &>13:-:/E";??-E? Z} &>13:-:/E";??-E 2;-85:3yɗɉɅuy̔˽ƮxqWsȂ ˌem)-.81 S &>13:-:/E";??-E? yɗɉɅzÍǢʞz ʒǈʞz

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(255) z( \ ãÍyy͂uȕ ˭esǬƳx̽[pm)-.81 S ďŭƣđŊɗƲu &>13:-:/E";?? y̔÷. . 34.

(256) ďŭƣ

(257) ǠãĉxÛ̌em̤̺ďŭƣđŊɗƲtyĵʪʞďŭƣ

(258) Ǡ ã̙xÛ̌em̤̺ďŭƣ͊Ŋɜã̙yɗƲtyĵʪʞuemSďŭƣđŊ ɗƲtĵʪemʞtz&>13:-:/E";??-E? yɗɉɅz &>13:-:/E";??-E? 2;-85:3 z

(259)

(260)

(261) tVďŭƣ

(262) ɗƲɜã ̙tĵʪemʞtz&>13:-:/E";??-E? yɗɉɅz

(263) &>13:-:/E";??-E? 2;-85:3 z

(264) tVpmSŢŠ̎ŊƑďǸ xZWsďŭƣđŊɗƲz &>13:-:/E";??-E? yɗɉɅuǬƳw̔÷ \˓ m& .

(265) S45?=A->1@1?@ xˈǸt&>13:-:/E";?? -E? yɗɉɅzďŭƣ

(266) ɗƲɜã̙yĵʪxȕďŭƣđŊɗƲt yĵʪxZWsǬƳx̽[pm& . 53 ̓Sm&>13:-:/E";?? -E? 2;-85:3 xqWsʏˋųəwǬƳƋz˓ wWyyļȆyĂ Ľ\mS dxďŭƣđŊɗƲtyĵʪxqWsďŭƣ ǠãĉtyÛ̌tĵʪe mʞuďŭƣ ͇

(267) ǠyÛ̌tĵʪemʞy͊ʞxď`ȕ˭ȂˌemS &>13:-:/E";??-E? yɗɉɅzďŭƣ ǠãĉtyÛ̌tĵʪem ʞ͂ %͏

(268)

(269) ̓uďŭƣ ͇

(270) ǠyÛ̌tĵʪemʞ͂ %

(271) ̓ u t ʏ ˋ ə w Ǭ Ƴ Ƌ z ˓ w [ p m S m &>13:-:/E ";?? -E? 2;-85:3 xZWsďŭƣ ǠãĉtyÛ̌tĵʪemʞ %͏ uďŭƣ ͇

(272) ǠyÛ̌tĵʪemʞ %͏ xʏˋųəw Ƌz˓ w[pmS ̾î·Ä¾ÇǀÐu &>13:-:/E";?? y̔÷. ¶Æ¢£©ÆÇ͂

(273). J. 93 ̳̓yɻʦĉħŊǀÐy &>13:-:/E";??-E? yɗɉɅxİgƠ̮xqWsȂˌemʍǺ45?=A->1@1?@ x ǀÐʞ͂ %͏

(274)

(275) ̓u̩ǀÐʞ͂

(276) %͏

(277) ̓xǬƳwƋ\˓ . 35.

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(279) ͐

(280) ̓z¡£«͇ʞ͂ ͐ ̓ &>13:-:/E";??-E? yɗɉɅ\ǬƳx̽[pm&. .

(281) Sm&>13:-:/E";??-E? yɗɉɅz ȏãÎtz¡£ «͂̓ͅʞ %͏ z͆¡£«͇͂̓ʞ%͏ xȕ˭eǬ Ƴx̽Wþɫem& . Se[e ȏãÍtz͆&>13:-:/E";??-E? yɗɉɅx¡£«yǬȸxƋɑz˓ w[pm)-.81 S &>13:-:/E";??-E? 2;-85:3 yɗɉɅzůŹĉʲ¡£«yǬȸuy̓x 45?=A->1@1?@ Z}ŢŠ̎ˈǸyWhyʏˋˈǸxZWsǬƳwɞ̔\ ˓ w[pmS. 36.

(282) ʢž ǰɤɲxZWsÛ̌ƣ ǠxŷǟdđŊŪŬˏǝ[ďŭty̓x yʚȒ̤̺xZWs &>13:-:/E";?? \˓ mSbyʍǺ[͆ġȥ̄ tyɷˤ̺ɉɊxZWs &>13:-:/E";?? xʌȪəǓŧzţ]WuʢY Sm%B1>-88&>13:-:/E";?? yɗɉɅ\ʆ tVpmbuz͆bt yɤɲŘŁ

(283) uļɯƗtV̬ŐZ}Ç¬tyŘŁ

(284) ìWɗɉɅtVpmS ŢŠ̎ŊƑďǸyʍǺđŊŪŬˏǝIċŪŬˏǝ̓xZ` ( yìÎ ċŪŬˏǝǦy ( ǯȮďŭƣđŊɗƲtyĵʪZ}ůŹĉʲ¡£«y űő\ &>13:-:/E";??'-@1-E? ÍǢdi˄ŋtVpmSmŢ Š ̎ Ŋ Ƒ ď Ǹ y ʍ Ǻ ċ Ū Ŭ ˏ ǝ Ǧ y ( ǯ Ȯ \ &>13:-:/E ";?? -E? 2;8-5:3 yɗɉɅÍǢdi˄ŋtVpmS ɊėɊƣxZ`ì ( z͆ĵʪɅw}x ->8E&>13:-:/E";?? ÍǢd iuŘŁdsW

(285) Sbt͆ʚȒ̤̺yǽ̹ɁƵz ->8E&>13:-:/E ";?? yɗɉɅxǬƳwƠ̮ÐYbu\ɫd->8E&>13:-:/E";?? yɗɉɅìÎdim xzʚȒ̤̺xĤď̎yʷˣw̸ǜÐYbu\ ǏŨdsW SǰɤɲyʍǺ[͆đŊŪŬˏǝ[ċŪŬˏǝty ̓xZ` ( yìÎZ}ċŪŬˏǝǦy ǯȮy ( \ &>13:-:/E";?? y ɗɉÃ£ÍǢdimSbybu[ŷ̡yʳƖɆřxZWsz͆đŊŪ Ŭˏǝƣx ( yìÎ˥bdwWẊĐw̸̹ɿɇʿXbu\Ț SɀxȞÕ̺xZWszß̺yƸ̒xéWȞÕ̎\śěkxéW ƪ˄w¯ÄÉśěguʢYm ̸̹ɿɇxȡƳ\ƪ˄tV uʢYS bt͆ďŭƣđŊɗƲtyĵʪtz ->8E&>13:-:/E";?? \̽Wbu\ ɫdsW

(286) SǰɤɲxZWs͆&>13:-:/E";??-E? y. 37.

(287) ɗɉɅz͆ďŭƣđŊɗƲtyĵʪ %͏ tz͆ďŭƣ͊ɗƲã̙ tyĵʪ %͏

(288) xȕ˭eǬƳx̽WʍǺuwpm . S mǰɤɲtz͆&>13:-:/E";??-E 2;-85:3 yɗɉɅxZWs͆đŊɗ Ʋtyĵʪt̽WþɫemSkyʍǺ%B1>-88&>13:-:/E";?? yɗɉɅz͆ ďŭƣđŊɗƲtyĵʪ͂

(289) ̓tz͆ďŭƣ

(290) ɗƲɜã̙tyĵʪ͂̓ xȕ̽þɫemSbyʍǺz͆ďŭƣđŊɗƲtyĵʪtz͊Ŋɜã ̙yĵʪuȕ˭e ->8E&>13:-:/E";?? \̽WugbtyŘŁuļÌy ʍǺtV

(291) Se[ẽĮyŘŁyÒxz͆ďŭƣđŊɗƲty ĵʪu

(292) ɗƲɜã̙tyĵʪut͆->8E&>13:-:/E";?? yɗɉ\ļɯƗug ŘŁ˓ SbyXwʍǺyɞ̆\wj˥by[x qWszÏǣtVSǰɤɲyʍǺ[ &>13:-:/E";??-E? Z} &>13:-:/E";??-E 2;-85:3 yɗɉɅìÎdim xz͆ʚȒ̤̺yď ŭƣyɁƵ^˅Ȅ ɁƵ\ƭe^wW̤̺tzďŭƣđŊɗƲ˅˹ ]tVuʢYS ɀxǠǰŐĉxZ`̃Įy˖Ǽ[͆ďŭƣđŊɗƲtyÛ̌\Ìʶət Vďŭ̺y \ďŭƣđŊɗƲtyÛ̌ŷǟesZďŭƣyɁ Ƶ\ʷũtwWʚȒ̤̺Û̌ʿpsWĸʭƮ\̽^ɆřyɄģƎx ďŭƣyʚȒ̤̺yɁƵy˅Ȅ \ &>13:-:/E";?? yɗɉɅìÎdim x̍˄tVuʢYS ";E u ?455 z͆ďŭƣđŊɗƲtyÛ̌xZWsǍī\ďŭ ƣ Ǡã̙x˥bpmřĻxĵʪɅ\̽WbuŘŁesWSm 8-:/4->0z͆ďŭƣ ǠxÛ̌ĵʪemʚȒ̤̺y &>13:-:/E";?? yɗɉɅ͂ ͏

(293) ̓zďŭƣ

(294) ǠxÛ̌ĵʪemʚȒ̤̺tyɗ ɉɅ͂ ͏

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(297) tVďŭƣ Ǡã̙yÛ̌ty. 38.

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(300) uƋz˓ w[pmSbybu[͆ǰɤɲtyďŭƣđŊɗƲty Û̌ty &>13:-:/E";?? z 8-:/4->0 yʍǺxȕ˭e̽^ɀxďŭƣ

(301) ǠtyÛ̌xZWs̽Wþ\ɫdmSbyXwʍǺyɞ̆\˥b˄ ŋues˖Ǽƀ˛Ńɰ\ɑwuWXȶdxďŭƣđŊɗƲtyÛ̌ xôdmʚȒ̤̺yɁƵ\Ơ̮İemĸʭƮ\ǏždSǰɤɲtz ˖Ǽƀ˛\ ÂµÅ¨¬ɰtVyxƀe8-:/4->0 yɤɲtzÉ ¥É·É£ɰ˖Ǽƀ˛uesWm ͆byŃɰyƋ\ʍǺxƠ̮ÐYs WĸʭƮ\VSm8-:/4->0 yɤɲtzďŭƣđŊɗƲtyÛ̌zď ŭƣ Ǡ

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(316) SǰɤɲxZW s͆ʚȒ̤̺yƔ̿\ÍǢgxéW &>13:-:/E";??-E? yśě\ ˓ mSě̿xéXʚȒʭĚyìÎz͆ě̿xůŹĉʲy˸ʿƮŠĠ yƠ̮uʢYsW S ,;;0? z͆ʚȒ̤̺yɁƵ\ʚȒƸʙxƠ̮ÐYuŘŁesWS ɀxǯʌɊ̺tz͆ɳʪ̺Z}ȞÕ̺xȕ˭e ->8E&>13:-:/E";?? Z } "-@1&>13:-:/E";?? yɗɉɅ\ìWuŘŁdsWSÍˍyɗ. 41.

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