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(1)Late. Miocene. Flora. Tatsumitoge. Honshu,. Southwest. Tottori. of Japan. Prefecture,. (I). By. OZAKI*. Kimihiko. Ⅰ. IntrodllCtion Many. arLd Acknowledgements fossils were gain from. plant. Tottori. Okayama Prefectures and Some those fossils were of. deposite. Hoki丑ora. Onbara且orule's. made. following. for. Japan. Japan;. species in it for. by. their. It is parpose it with Miocene under. TANAI. and. the. to. paper. of. and. fossils. of. UEMURA. Japan. to permission locality. Mr. UEMURA. author. *. GeologlCal. is. to. chance. and. stored. Science. National. south-. collections Museum. Prefectural. Tottori. the. and. three. on. study. in the. describe. Tatsumitoge且ora. the. its comparisons with discuss the physical and. of this且ora. was. already. his great. Museum. compare. and some. with. late. other. environmental. conditions. in. 1976. Professor. T.. reported. 1976･. loan. much lnstitute. appreciation of Hokkaido. Mineralogy the. course. also author National Science. all. kindly obliged. their. the. on. late. to. University. study. by. T･. staylng. Museum,. have. six. Palaeomonthes. to gratitude Of the GeologlCal. his deep T.. ONOE. Prefectural. from collections identi丘cation the. Professor. for. TANAI,. who. in the. and. acknowledges. fossil. discussed to. during. Tottori. Of the of. this. of. University. The. Of the I. YAMANA and. kind. The. in the. to. along. to express wishes and of Geology. September. survey. a. of Japan. advice and facility during botanical Laboratory of Hokkaido. K.. was. is notable from reported. from. I have. many. Messrs.. report. of. vicinity. this且ora. on. author.. author Department. from. as. lived.. maple. The. the. this. of Japan, they. which on Only. in the. road. 且oras. Miocene. uranium-mineral ONOE (1961). Ningyotogeflorules･. and a. of. courtesy. of. aoras. and. Tatsumitogeflora. as. Survey. Hokiflora,. the. late. brief. the. and. This. locality. Geological. the. of Japan. a. Tatsumitoge. the. of Japan,. few. fossil且ora.. Japanese. from. gained. as. to. compare. 1968･. thing,. one. the. TANAI. border. around. those central northeastern reported is of many of the且ora number composed w叫preserved fossil species the Hokiflora many unreported and includes. are. another,. and. in. et al･. points:. western. reported. in 1963,. localities. YAMANA. by. by. Onbara of three Mitoku, a on from discuvered exposure. was. of. throughprospecting. is composed. which. Tatsumitoge鮎ra the. Mio-Pliocene. the. Emeritous. for. Museum the of TI. their. Tasumitoge some. fossils･. SHIKAMA. Of.

(2) 32. K.. Yokohama Akita. National University,. thanks logy. are. University. for. also. their. and. Professor. and. valuable. for. his. Emeritous. suggestion. Dr. Goo. to. extended. Palaeontology. OzAKI. due. also. This. Dr.. to. study. a. on. from. ⅠⅠ. Geographic l.. Fossil. is. Prefectures. as area. mountain. 750m about Tatsumi-toge. of the. of. in. have. the. 1.. been. Technology,. the. Grant. in. Aid. for. level. sea. border. the. on. This. district. of is situated boundary. the. and. is about. and. Tottori. reglOn. mudstone Prefecture.. Tottori. 1311T. JAPAN. of Yoko-. from. collected. Yazu-gun,. Saji-mura,. (pass). Chugoku. the. and. Education.･. 且ora. above. in Text一点g.. shown. Department. Settings. Saji River. the. of. of GeoThanks. species. the. Of. by. partly. of. Tatsumitoge. of the. plants. of aupperreacb The locality northeast. Ministry. the. and Geologic Environment. Present. chinese. OKUDA. supported. of Cordial. Of Nankin畠Institute. kind. 丘nancially. was. Research. Scienti丘c. S.. HuzIOEA. this study.. concerning. SHUANG-ⅩING. suggestion Dr. K. FuJIWARA and Vegetation Science, Institute of En立ironmental Science hama National University, for their suggestion. are. K.. 200m. and in the of. 32113. east_. Okayama backbone. Okayama. 3.13. and. ヰ○. SEA ■5'. L3. i5. 0＋o. ▲一′-一￣￣ヽ一_. '､'.-･汐準_･Q J>hL7yo. KURAYOSHI Mitoku. TOTTORI. ♂. P. /ダ. ▲. MT.. MITOKU. PREP. ▲. ′■ヽ__･. ---'へ?. ′. ノ /I. Ni ngyotoge. /. Tatsumitoge. Onbara. -ヽ. ー･ー■-･･＼･_.J4･_･/･-･＼. /メ. ヽ l､. A,･-･/. ＼. く. ､. ∼.. OKAYAMA. J. Text-丘g.. 1.. Map. PREP.. showing. o. the. fossil. 5 kTh. localities.. 't.

(3) Late. Tottori. Prefectures. Tbere lower. Miocene. in this. is. The. climate. and in the. A. the. Onbara. by. area. from. there. fossil. the. around. conditions. Sea:. Japan. the. falls. climatic. formed. plane,. and. level,. sea. a. at. level.. sea. by. snow. the. above. highland. Ningyotoge. the. much. 1250m. to. level.. sea. above. 33. I. watershed.. is in且uenced. area. in winter. precipitation is rather transitional. the. 800m. in this. Flora. is l150m. between. to. a. one. above. found. 700m. on. higher. peneplanes; is about 500m. volcanic rocks, height of about. is. area. are. one. Tatsumitoge. much. locality,. Sea. Japan. are. it. but to. side. the. Paci丘c. side. According. to. Saji district. of the. D.. japonica. DoN.. area. small Fagus the. The. the. Fagus. (1974),the. the. sometimes. covers. peaks. around. of Mt. Tatsumitoge. the. He. state.. (899.9m). Mitoku was. been. has. mountain. natural. one and of which Tatsumitoge pass･. and. divided. at 9km about situated by SASAKI (1958).. distinguished a. as. protected. shrine. into. the月.ora. forest. two. to. into. two. covers. subassociations 400. about at 600m. about. mountain. leaf forest 2.. 600. Sasa. and. above. is. there. no. developes 400m. about. is. association acuta. up. divided. subassociation grows. subassociation 600m. covers. of. almost. to. up. the. deciduous. warm-temperate. top. of broad. zone.. Settlng. Tottori. Prefecture. central fectures in western. border. the. various. of Tottori. volcanic.. rocks. of. and. Okayama. the. late. Pre-. Miocene-. are distributed. These terrestrial sediments with widely Misasa Group, cover Sangun the unconformably mainly late Mesozoic It is that and rocks. granitic well-known deposits. The the contains sometimes uranium-mineral. in the. metamorphic. rocks. Misasa. to. Honshu,. accompanied included. rocks,. that. former. The covers. metternichii from. facies. kept. Aucubeto-CyQlo-. Cyclobalanopsis. :. Rhododendron. senanensis. It is noteworthy. GeologlC. Pliocene. m,. facies. one. and. to. m. and. ln. the. m. northwestern The forest has. associations:. and Saseto-Fagetum crenatae. stenophyllae 400m the level the latter sea and about above Saseto-Fagetum the top of the mountain. crenatae. to. 800. than. more. ridges. balanopsidetum up. vegetation. afforestation of CryPtomeria forest are. in some recognized. upper. 且ora. of. this. by. OcⅡⅠ. level.. sea. part. SⅢIMIZ口and. Saji River, reaches of the Tatsumitoge fossil locality to the. the. near. forest. by. is widely occupied Some associations of. along. from. extends. investigation. the. Group. sedimentary. in. rocks. distribution.. These. the. Group. Misasa. have. sediments. develop. locally. been. difBcult. reported. by. to. in. lenticularly. and. correlate. each. other. by. lithology. The yAMA. but. geology and OzAWA. there. are. uranium-mineral in the group. of. this. was. region (1961),YAMADA. many. confusion FuKUOKA deposits,. whole. area. in the. FuJIWARA. (1961,1966), YAMANA on. stratigraphy. and. view. KuBO. point. of. et al.. Through. (1969). (1957),MuRA(1968) and others,. et al.. prospectlng. investigated. volcanostratlgraphy,. the and. the Misasa they.

(4) 34. K.. Table. 1.. Volcano. OzAKI. Misasa. correlation of stratigrapbic Fukuoka KロBO, and. (simpli丘ed from. CENTRALAREA. Group. 1969). NINGYOTOGEAREA. EASTERNAREA V. VV. vL. VV. lV. M. VV. VV. lvv. Ⅰ S. VsuGEGATANI.”.. A. ㌶:㌶:v TATSUMI-. TAKASHIMI.ZUF.. VV. ⅠⅠⅠ.. VV. 1vv. VV. S. L.M.V. VV. TOGEF.. NINGYOTOGEM.(N). lsABIM.. A. V. リ･M･(0) L･M･v(K. NISHIOSHIKAM. VV. G. VV. R. YOSH川ARAM.(”). ⅠⅠ. 0. TOCHI-. U.M.(T). VV. WARAF∴. M.M. L.M.. VV. ､U P. VV. lv. ∫. トⅠⅤ: Volcanic. (M). :. F∴. cycles,. Mitokuflorule,. (K). 且ora,. divided. (0). it into. sandy. and. mudstone. conglomeratic formation in toge且ora, of the. are. Onbara. TANAI. 且orules WARA. the. and. the. three. florule,. three. (T). areas. :. Tatsumitoge. member. The. member mudstone,. Formation. isolated. described. with the is made upper. conglomerate, Tatsumi-. upper. member uncon-. conglomerate. and. conglomerate,. fossil. and. of. plants. of the. unconformably. Onbara. is divided. and. mudstone. covered. this. cover. underlain. The. tuffaceous. upper by. the. Ningyotoge. by F口Jト stratlgraphy KuBO, to the stratigraphy of FuKUOKA and in the Yoshiwara Memeber, the Onbaraflorule. sediments,. followed. of. consists. tuffbreccia.. Mitoku,. of conglomerate,. of the. of basal. up. of basal. consists. basalts,. Kamejiri. Tochiwara. fossils, the. plant. member. is mainly. three. The. basalts. part mudstone of the upper The Tatsumitoge Formation. mudstone, in the preserved. (1961). made. in. area. eastern. up. Kamejiri many. preserved. tuffaceous are. is. 1.. in Table. shown. as. is composed. member. Formation lower. al. (1957). According Mitoku且orule is included et. Well. the. and. Tatsumitoge. andesites. and ONOE from. Ningyotoge. mudstone.. part.. from. Tochiwara. sandstone, florule The. member. Akawase. mudstone;. of sandstone. tuffaceous. upper. Formation.. tw■o members;. alternation. the. and. the. Volcanics. Ⅴ:. in mainly. ; middle. mudstone. sandstone its eastern. Tochiwara. formably into. sandy. obtained. by. (N). :. KuBO, the lithology and of the fossils are included is as follows. plant lower into three members: member. the Tatsumitoge which Formation is divided conglomerate. cycles. volcanic FuKUOKA. to. Member,. Onbaraflorule,. four. According. M.:. basalts. Kamejiri. :. Formation,. :. to. the.

(5) Late. is in the 且orule III.. upper. Ningyotoge. Systematic. for. Museum,. National. Tottori), Yokohama).. University,. Abies. DescrlptlOn:. Abies. both. hokiensis. OzAKI,. new. Discussion: medium to. of My. those. and. 4mm. straight. in. the. end. slightly rounded. belongs to seed. A･. of. et. distal. distal. nearly且at. ZロCC･. (HAYATA). Collection:. of. similar winged in central Honsbu. PP. growing 16219.. Abies. ugoensis. NSM. shape. of Shikoku,. to. angle. about by. the. It is slightlysimilar the Miocene middle of. the. seed. in the. Abies. genus. from. in the. separable. ITO. the. wlng･. (1961). TANAI. an. making. Seed. wide. burged. slightly. margln,. the. lOmm. and. Wing. wide･. to. distributed. kawakamii. is. long. outer. end. by. aburaensis, reported Hokkaido, but is closely. specimen. 15mm. deltoid,. singlewinged. the. and. 8). long. A. southwestern. SIEB･ A･. size. SPeCies. 6mm. marglnS,. PP. MILL.. seed. almost. the. 1.. Pinaceae. Winged. triangular-reniform, margln,. NSM. are. YNU. and. (Pl. 1,丘g.. by. in Table. shown. (National Science Japan, Tsukuba), TPM (Tottori lnstitute, Yokobama (Geological. of specimens (Geological Survey of. Genus. 55o. Ningyotoge. the. and. as. respectively. Family. inner. 35. Formation. repository. GSJ. Tokyo),. Prefectural. Member. I. Descriptions. Abbreviations Museum,. Flora. Tatsumitoge. Tatsumitoge. of the. member. is in the. Miocene. seed. and. also. vlng.. and. A.. extant. to. homolePis the. extant. in Formosa.. HuzIOKA. UEMURA. et. (Pl. 1,丘gs.ト4) Abies. HuzIOKA. ugoensis. Bull.. UEMURA,. et. Natl.. Sci.. 16(4). Mus.. 697.. :. Pl.. 1.. 1-5,. i.. 1a. (1973). Discussion base. sucker-like that to. the. Detached. :. leaves. of the. fossils. leaves. represent. A･. a. by. and. species HuzIOKA. ugoensis reported Miyataflora of Akita Prefecture･. jirma. reported. onbaraflorule mentary. by. TANAI. of Okayama The. A･メrma. SIEB･. (1961). A･ on. Prefecture,. Abies･. of. and. UEMURA. basis but. be. A･. of. a. expanded indicate seeds,. specimens. (1973). may. ugoensis the. These. by. from. are. late. A.. conspecificwith. single. protojirma. cone-scale. is based. referred Miocene. on. proto-. from. the. a. fra竿-. I refer my specimens (leaves and seeds) two fossil species are above-noted closely similar to the modern et Zucc･ in Honshu to Kyushu growing of Japan.. cone-scale･. ugoensis･. Thus,. andwinged seeds, characterized deltoid shape of the winged. tウA･.

(6) 36. K.. Collections:. NSM. PP. 16096,. OzAKI. 16115,. Genus. 16116;. Keteleeria. Keteleeria. TANAI,. ezoana. 41. TANAI. ezoana. 40,. and. This. MAST.. euelyniana Collection: NSM. My. from. PP. 16097,. Genus. Picea. kaneharai. TANAI. Discussion ONOE. relatively to. f･. 1.. Pl.. 16,. the. Kyushu,. 16117,. Japan,. P. polita to. and. ONOE. et. 19, 20). Surv.. Rep.. Japan are. P.. extant. 1.. 18了: 17. Pl.. Picea. to. referred. (SIEB. et Zucc.). the. 16178.. DIETRICH. Miocene CARR.. species. (1961)･ TANAI has. been. is closely Honshu in central. of Japan, and. growing MAST.. neoueitchii. 9. I.. kaneharae. This. shape.. small size and oblanceolate known from to late the middle. living. base. ezoana. Formosa.. and. TANAI. seeds. winged. leaf K.. of southern davidiana BEISSN.. K.. living. their. well to. A.. it represent. that. Yoshioka且ora. the. 16111,. 1,丘gs.. Geol.. ONOE,. Several. :. by. et. to. China. kaneharae (Pl.. similar. 25l.. :. sucker-like identical to. are. Miocene. similar. 16095,. small. specimens. Of western. Picea. et. ll(2). indicate. seeds,. middle. is closely. species. K.. of winged. margln.. (1961). TANAI. Hokkaido.. Picea. IV.. of the叩eCimens. shape. leaf. rolled. by. to. features. semi-elliptical. slightly. and. Several. :. :. reported. Univ.. (1961).. Discussion Keteleeria. 5-7). Sci. Hokkaido. Jour. Fac.. 497.. TPM. CARR.. (Pl. 1,丘gs. Keteleeria. 4806;. GSJ. in central. growing. China. Collections:. NSM. PP. 16002,. 16112,. Picea. ugoana. 16113;. HuzIOKA,. ugoana 20a. et. TANAI,. NISfJIDA,. Jour. Fac.. Discussion P. ugoana. seeds.. They. similar. to. Japan.. Minn.. TPM. 383.. 19, 20). Coll. AkitaUniv.. 3(4) :62.. A.. Pl. 1. f. 17121,. 17a-. (1964).. HuzIOKA. to. Jour.. 4831;. HuzIOKA. (Pl. 1,丘gs. Picea. 4802,. GSJ. :. Bull.. Sci.. Small,. HuzIOKA are. the. most extant. Sado. Mus.. Hokkaido. Univ.. 3:. 10. Pl.. IV.. oblanceolate-oblong from the Miocenelof. ll(2). to. similar P. koyamae. the. extant. SHIRASAWA,. 1. :. i. 254.. 2. (1960)I 1,f.. Pl.. 28,. 29. (1961)･. seeds of Picea are Japan by the rhomboid. referable. P.. are. winged. bicolor both. MAYR･. living. ln. and Central. shape. of. also Honshu,.

(7) Late. Collection:. PP. NSM. Tatsumitoge. Miocene. 16002,. 16112,. CARRIERE. Pseudotsuga. sp.. 1,丘g.. (Pl.. the. and lacking. though. in the. P.. extant. Pliocene. number. the. of. by. its. ellipsoidal This. cone-scales.. is closely of the cone-scale, in the orbicular My cone-scale.. part. SHIRASAWA. from. P. subrotunda district (MIKI. of Kinki. Pseudotsuga. to. shape. and. marglnal. japonica. distinguishable. 22). is referable. cone. single. size, and. shape. the. A. 37. B.. 16113A,. Pseudotsuga. Genus. Discussion:. Flora二. MIKI. and. by. 1957). P.. the. size. the. and. to. similar. is. specimen from. MIKI. gondylocarpa. cone. cone,. shape. the the. of. cone_scales. Collection:. PP. NSM. 6411.. Genus. Pseudolarix. Pseudolarix. GoRDON TANAI. japonica (Pl. 1,丘gs.. TANAI. PseudolarixjaPonica. ONOE,. GeoI. Floras. Japan. et. et. ONOE. Japan. Rep.. 9-15). Surv.. 187:. 17-18.. 1,. Pl.. 5. f.. (1961). TANAI. N.. et. descrlptlOn. Supplementary long,. 6.5cm base;. indistinct. 0.1cm. about. apex. Tert.. SuzuKI,. ; several. Leaves. :. in the. wide. to. Obtuse. base. round;. striations. 1. 102.. :. obsubulate, part. middle truncate;. I.. generally. falcate,. 2.0. gently. tapering. to. and. the. (1963).. 7-9. margin to. parallel. running. 1.. Pl.. entire;. midvein. ; texture. margln. to. the fine,. Chartace-. OuS.. Discussion are. on. the. shape to. of the. similar Collections TPM. by. texture,. chartaceous. the. base,. extant :. the. basis. the. winged-seeds. mented. of detached. number. NSM. of TANAI. parallel place. P.. SuzuKI. (1963).. several striations leaves in Pseudolarix. GoRD.. 16093AIH,. 19189,. 51, 230.. Genus Tsuga. Tsuga. hokiensis (Pl. 1,軸s.. CARRIERE. OzAKI,. new. 1961) The. above. This. in central. growing 16094-16096,. SPeCies. 21, 23, 25, 26). winged-seeds. this species. ONOE,. et. running these. and. To. (TANAI N.. and. cone-scales. et ONOE.. cone-scale. kaemPferi PP. leaves,. TANAI. PseudolarixjoPonica. to. referred. established. A. :. originally. was. above. suppledescribed. leaves. the. species. and is closely. China.. 16195,. 16207,. 16242;.

(8) K.. 38. Type. Holotype,. :. DescrlptlOn. Cone. :. obliquely. apex;. deltoid,. at. 4mm at. apex,. to. obtuse. 6. I.. specimens. but. Of seven wings of seed･ shorter 丘ora of Japan, my specimens HuzIOEA. et. (TANAI,. cenica. is also. also. 1961). of Tsuga. separable by. separable. the. shape. in T･. rotundata. the. of. from. my. specimens. PP. 16098,. 16099;. distinct;. and. in the. cone. smaller from. the. sieboldii seeds. miyataensis. (HuzIOKA. (MIKI, 1957) the. and. and. cone-scale･ T･ mL'0and. (TANAI,. T･ aburaensis. size. and. Neogene. the. T･. to. longer. and. cone-scale･. in. Tsuga. extant. similar Formation. Miyata longer. from. the. reported. closely. Miocene. late. the. distinguishable. are. are. specimens. 1961). from. 1973), but. UEMURA, My. UEMURA. to. similar. most. rounded. longand. emarginate. thick. midvein. different. species. wide,. part･. long･. are. are. cone-scales aplCal. wide･ 6mm. ovate,. 2.5mm. to. enitre;. closely. cones,. and. wide; in. 5mm. to. wing. 1.5. lmm. about are. 4. and. and. margln. l･3cm. to. truncate. wide;. long. lamina,. the. long. 2mm. and. 1･2. almost. 9mm. to. base;. at. round. leaves. the. with. wide. deltoid,. petiolefine oblique These Discussion in. (Late Miocene)･. Formation. long,. 2･4cm. to. 3mm■1ong. to. CARR.. Tochiwara. linear, 8 to 12mm. Leaves. wide.. lcm. about. quadranglarly winged-seed. 2･3. oblong,. round,. seed. Upper. Prefecture.. Tottori. Saji-mura,. 459.. TPM. OzAKI. the. of. shape. winged-seed. NSM. Collections:. Family Genus. R. BROWN TANAI. Protokonishii (Pl.. TANAI. ♪rotokonishii. Taxodiaceae. Cunninghamia. Cunninghamia. cuTminghamia. 459, 460.. TPM. 1,丘gs.. 24,. ONOE. 28). Geol･. ONOE,. et. et. Surv･. Rep･. Japan. 187:. f･. 18･ Pl･ 1,. 1. (1961). Discussion Cunninghamia domly. :. arranged. cone. attaching. konishii by. ture. konishii. TANAI TANAI HAYATA,. features. Several. :. subulate. foliar. foliaged. to. a. et. ONOE and and. dorsal. shoot. from. reported. (1961).. ONOE is also. My. similar. leaf, and. of the. This. Onbaraflorule. ran-. ill-preserved. splny. is similar. to. C･ lanceolata. HooK･. both. China. collections:. NSM. Genus. PP. 6413,. 16135;. Metasequoia. Metasequoia. MIKI. occidentalis (Pl.. 1,figs.. 4789,. GSJ. eX. 4838;. Hu. (NEWB･). 27, 29, 30). et. to. of Okayama. species. to. it represents margln･. identical. are. specimens the. leaf. the. on. spines. side. that. indicate. specimens. minute. shape,. the. on. stomata. of the. TPM. 54.. CHENG CIiANEY. ･. the. C･ protoPrefecextant. growing. C･. in.

(9) Late. Metasequoia. there. Discussion. shoots. its. Collections:. NSM. PP. 16202;. Genus. lip. a. mouth,. lower. the. and surface Discussion : Two. rather月.at dish-like. Collection. lateral. the. SIEB.. Family Genus Cyclocarya. Cyclocarya. (TANAI. ezoana. 1.4. long. and. the. at. cm. at. 2.Ocm. abaxial. abaxial. side On. margln. side.. are. to similar in southwest. closely growing. those. the. of. Japan.. ILJINSKAJA. N.. et. 2,丘gs.. N･. et. about. thickening. upper. Cyclocarya. (Pl.. 450.. Juglandaceae. (TANAI. ezoana. 449,. side,. Zucc.. et. known. widely. 5ト53,. depressed. cone-scales. uerticillata GSJ 4805.. :. been. cone-scales. Zucc.. et. orbicular,. on. the. 31, 32). deeply. rather. Sciadopitys. extant. 225. :. sp.. 1,丘gs.. and. has. TPM. 4788;. SIEB.. depressed. cone-axis. the. of. 40(3). Metasequoia. to. of. species. 4787,. Sciadopitys. Cone-scales. toward. thinning. wide, like. Soc.. identi丘ed. arrangement. This. GSJ. (Pl. :. 39. Philosoph.. are. cone. decussate. Sciadopitys. DescrlptlOn. Amer.. single. leaves.. arrangement of opposite Tertiary the of Japan.. and from. a. and. by. CHANEY. (NEWB.). occidentalis. Trans.. I. (1951).. cited.. Foliaged. :. Flora. Tatsumitoge. CliANEY,. (NEWB.). occidentalis. an￠ all丘gs.. Miocene. 3, 4;. Text-丘g.. WoLFE. SuzuKI). WoLFE. SuzuKI). et. TANAI. et. 2D) TANAI,. U.S.. Geol.. Surv.. Pap.. Prof.. (in press). Pterocarya. TANAI. ezoana. 11,Pl.. IsIIIIDA, Mem.. (in press), based 丘nely. the. enter. the. from. those. of the. shape. and. from extant. the. 1. Japan. Mineral.. and. of the. the loops. Tatsumitoge. C. paliurus. Miocene. margln. and. of. leaflets. end. locality. (BATAL.). by. Platycarya. distal. genus. :. 110.. Pl. 6, f. 2-5, 1. 37(1) :70. Pl. 5.f.. was. My. specimens. branch. of･ the gepus dichotomized. by. generally. of the. near. Cyclocarya secondary. 8,. 9,. (exclud･. is closely. in central. ezoana,. and charac-. secondary margln. and. separable veins, round. No. veins.. are. are. asymmetrical. secondary C. ezoana. growing. lea且ets. features. the. of the found.. ILJINSKAJA. Cyclocarya with Pterocarya by WoLFE. characteristic. dichotomously. The. genus. the. Alaskan. Pterocarya. genus. identical. are. from. sometimes teeth･. marginal. those. on. serrate. secondaries. from. Floras. Geol. leaflets. transferred. recently. was. :. Tert.. (1963).. Univ.. oblong. TANAI. veins. Kyoto. These. :. which. by. 10. (1970).. Discussion. terized. Suzu王くⅠ,. 21,f.. Sci.. Fac.. i. 2). ing. N･. et. 1,pl.. 19.f.. and basal. winged-nut. similar. to southern. to. the. China..

(10) K.. 40. Collections:. NSM. PP. OzAKI. 16064,E 16160;. GSJ､4791,. Genus. Juglans. Linn.. Juglans. japonica. TANAI. (Pl. 2,丘g. TANAI,. JuglansjaiIOnica. i. 9). (excluding Discussion by. TANAI. that. margln. Or. serration. widely northern pl. 6.丘g.. foliar. ll(2). in and be. 275.. :. i.. Pl. 6.. 10. to. Japan. and. also. Korea.. A. specimen from this. run. along. marginal ailanthifolia CARR.. MAXIM. I. mandushurica figured as by japonica I.. to. by. species. its. the. 丘ne. the. end, and living I.. the. secondary. then. margln. distal. similar. Juglans japonica. to. brochidodromous. the. in the. is closely. excluded. to. up. marglnal. species. identical. are. characteristic. teeth. This. 9). shape, curving. enter. may. IV.. Univ.. leaflets. slightly the. growing China. 478B.. 7). Hokkaido. fragmentary. oblong. extend. Sci.. TPM. (1961).. Two. :. the. veins. Jour. Fac.. 4801;. in growing TANAI (1961,. intersecondary. simple. VelnSJ. NSM. Collections:. PP. 16066;. YNU. Genus. 1080.. Pterocarya. KuNTH. Pterocarya. sp.. (Pl. 2,丘gs. Discussion secondary. indicate that serration leaves were figured as. Many. carya.. from. description P.. (1931).. has by. widely. Miocene. is closely. in Japan.. been. not. from. reported. leaves. My various authors. PP Collections: NSM. balsamoides K.,. GoEPP.Foss. Sci.. Rep.. in central the. identical. are. P.. Without KoNNO. by. Honshu P･. extant to. KoNNO,. rhoifolia SIEB･ but. asymmetrosa,. validly published, although”P･ Miocene of Japan and its adjacent to fragmentary. are. Genus. SuzuKI,. Omi且ora. species. veins, and of Ptero-. 16135,. 16140;. El.. the. to establish. GSJ. a. speci丘c. region status･. 4802.. Salicaceae. Populus. cf. balsamoides. Fukushima. et. asymmetrosa”. Family. Populus. asymmetrosa. tertiary a. the. 16082,. Populus. Pterocarya. to. brochidodromous. shape,. percurrent of loops, fossils the represent. similar. leaves. My. has. asymmetrosa,. been. late. the. asymmetrosa. growing. epithet,. series. 10). to elliptic foliar. ovate. asymmetric a form that. veins. fine marginal. Zucc.. The. :. 8,. Schossnits. Univ.. LINN.. GoEPPERT 23. Pl.. 10:. 15.. 26. Pl. 2.. I. 5, 6 (1855). i. 8, pl. 3. i.. 1,. 2. (1961)..

(11) Late. Discussion indicate. a. close. base,. cordate. Only. :. ill-preserved. one. slightly. zlg-Zag. close. veins p. balsamoides. course. in the. This. species. My. is somewhat. leaf. HuzIOKA. by. described. TPM. secondary. kitamiana. TANAI. Discussion. N.. et. This. :. P.. extant. HuzIOKA. sambonsgii. 14,: 137.. n.s.. Type:. Miocene).. to. 4. 60o,. veins. to. near. 5 alternate. ultimate. area. triangular. palrS,. slightly the. weakly. veinlets. margin area. the. and incurved. a. Of the. palr. 2A,. States･. E) Palaeont･. Proc･. Soc･. Japan. (Late. Formation. at. form. percurrent;. twice. apex. a. at. turning marglnal. margmal quaternary. branching. side;. loops teeth,. with while. veins ; petiole. more. distal. the. off the the. or. secon-. remaining. at 40o. the midvein some. end,. length. leaf. the. from. glVing. medium, diverglng. tOW･ard. of. leaving. apICally,. Vein veins. zlg-gag. half. dentate. margin. secondary. abaxial. spaced,. wide;. cuneate. widely. ; prlmary. basal. about. at apex,. obtuse 6･7cm. to. extending. midvein,. Irregularly. entering. largely. species. Tochiwara. to deltoid,. with. half ;. zlg-lag, to. Text･fig1. 3･7. the. in marglnal. costal. 4.5 to 8･2cm. 45o from. is similar. Prefecture･. long. its upper. ln. Populus. to. Species. new. Upper. 16075･. identical. United. (in part), Trans･ (1954)･. margin reaching to 5 7 tertiary Off veins. running. forking. f･. 10: 7･ Pl･ 2,. Pap･. This. SuzuKI. oval. and. glVlng. daries. PP. is. eastern. OzAKI,. broadly. teeth. of about. up. more,. Leaves. :. zlg-芝ag. turning. (only). Saji-mura,. ; each. angle. K･. 7. Tottori. base,. serrate. an. Sp･. Japan. teeth･. Of the. 4,figs.ト4;. pl.. et. I,l. 16･f･. NSM. totruncateat. MICHX･. 4;. Holotype,. DescriptlOn. at. Soc･. dentate. hokiensis. ag.. (pl. 3,. running. teeth･. 2B). fragmentary,. land. large. Populus. to. two. the. N･ SuzuKI. et. Palaeont･. though. grandidentata PP NSM 16073.. Collection:. pupulus. marginal. 5; Text一点g.. SuzuKI,. specimen,. in its characteristic. kitamiana to. marglnal. (1965).. 8. the. in. larger. the. and. TANAI. kitamiana. (Pl. 3,丘g.. 3,. larger. from. veins. L･. 91.. Populus. populus. balsamifera. is distinguishable. but. to. similar. P･ sanzugau)aensis, and Miyata the late Miocene. from. (1973, 1974). slightly. Of northeastern. species. P. mLyataenSis. to. Prefecture,. of. number. HENRY. two. the. P･. extant. maximowiczii. from. similar. in Akita. less. in the species Collection:. P.. UEMURA. and. Sanzugawa且oras. and. to. the. to. that. secondary. CamptOdromous leaf is closely. My. part･. similar. characterristics foliar shape,. round. Vein,. is. 41. I. several. ; deltoidly. basal. America and also slightly of North Asia ; but my leaf is distinguishable teeth.. has. Of prlmary. space. GoEPP... Flora. leaf. of Populus. relationship. have. that. Tatsumitoge. Miocene. branches. or. adjacents;. tertiary. tertiaries. in inter-. orthogonally l･7 than. reticulate; cm･.

(12) 42. K.. OzAKI. A. C. ヽ. I l. l. /. 亡m. フ1. 2mm. B 一. 2cm. E. D. 2o. 2mm. 0. Text･丘g. A. Populus. hokiensis. B. Populus. kitamiana. C. Caprinus. D. Cyclocarya. E. Populus. n.. sp.. Venation. NSM. TANAI. hokiensis. n.. sp.. et. n･. sp.. PP N.. NSM. (TANAlet. ezoana. hokiensis. 2.. N.. Holotype,. of fossils.. characters 16076.. SuzuKI PP. 2cm. NSM. PP. 16073.. 16150.. SuzuKI) NSM. WoI.FE PP. 16075.. et. TANAI. GSJ. 4801..

(13) Late. Discussion: followings. among. Among. the. the. but. marginal P. inoueL'reported. Formation. in northern. from. primary. straight. nearly. is. new. species P. davidiana. DoDE. Among. NSM. 16075,. 16076;. Slaix. hokkaidoensis. hokkaidoensis. f.. 6.. TANAI. et. 7. (1965).. I6.i.. pl.. Discussion. A. :. is similar in Japan. to. veins. NSM. Collection:. PP. Holotype,. :. at. an. :. acute. at. of 40o of. in intercostal. and. Leaf apex,. 6;. in. ultimate. the. areas. Upper. SEEMEN. species growing. SPeCies 3A,. ち). Tochiwara. veins. once. about. base,. (Late Miocene)･. Formation. area. to. to. secondaries. loops. making thrice branching;. angular. in marginal. meshes,. poly卯nal. the apICal. form. meshes or. within. entering. petiole. thick,. from. midvein the midvein. area. the the more. joining. and. portion loops;. 2･6cm. and. serrate;. diverglng. alternate,. extending. polygonal. making. irregularly. margln. to. (estimated). long. 8cm. 6 pairs,. 60o, irregularly. making. marginal. at. round. superadjacent. veinlets. 4825.. new. Texト丘gs.. lanceolate-oblong,. veins to. ; quaternary. meshes. narrow. This. teeth･. marglnal. its. Pl･ 15･. Prefecture.. ; secondary. angle. branches. by. hokkaidoensis. S. gilginata. and. OzAKI,. hokiensis. GSJ. Tottori. Description. straight. arranged. 10: 9･. Pap.. Sp.. Japan. Salix. to. KIMURA. (Pl. 3,丘g.. wide,. Soc.. 16206.. Salix. Saji-mura,. 493.. Korea.. and. Type. China･. northern. 2). Palaeont.. coarsely. and. S. koriyanagi. living. the. SuzuKI,. this of Populus to in Japan and. N. SuzuKI. et. 2,丘g.. leaf is referred. single. less secondary. shape,. N.. is different. but. LINN.. TANAI. (Pl. salix. GSJ. shape. marginal Pliocene Oya. the. growing. to central 4798; TPM. foliar larger. the. species. MIQ･. Pre-. Fukusbima. specimens,. living. in. and. Genus Salix. from. sieboldii. in Korea. PP. (1954) my. the. P.. to. similar. by. resembles. et SuzuKI. round. latter. the. TAKAIiASHI also. growing. Collections:. and. vein.. closely. from. species. new. HuzIOKA. rather. ; zlg-. lamina･. the. of. in. the. secondary. Japan, this. of. by. former. Of. base. and. the. by. Kyushu. 5 palrS. the. Neogene. teeth. marglnal. to. by. mainly. P. sambonsgii Formation Shiotsubo. Miocene. dentation,. 4. from. extends. SuzuKI. et. dentate. veins;. the. 43. characterized. ; nearly. from. reported. and dentation.. in. shape. palr. is distinguishable. finer. is. poplar. of. secondary. and basal. HuzIOKA aizuana from the late. reported. fecture,. prlmary. fossil poplar P.. species foliar. round. which. resembles both. the. of. veins. new. deltoidly. :. course. zag. This. I. Flora. Tatsumitoge. Miocene. tertiary making. veins angular. tertiary. meshes. teeth;. marginal. than. 1 cm. long･.

(14) 44. K.. OzAKI. C. B. A. 【. l F r. 0. i. 也. ∫ I. I. (. ∼. I. I ∫ l. 04. mm. ,.F 04mm /. ト5cm. D 0 i;. E ヽゝ. r. ′一′. I. ゝへよ ⊥ナ. 3cm. Lt. 5mm ￣｢■｢｢ー. Text･丘g.. 3.. Tbe. details. A,B.. of venation. their. some. Salix. C.. Salix. D.. Salix. E.. Salix. F.. Salix. sp.. related hokiensis B.. vulpina. NSM. of. characters extant. n.. sp.. PP. ANDERS.. sp.. A.. TPM. 284.. sp.. C.. NSM. PP. species. GSJ 4825.. 16027. YNUPB. 16209.. 355.. fossils. and.

(15) Late. Discussion: KIMURA dary. This. Miocene. fossil. Tatsumitoge. is similar. leaf. from. Miocene. is different. 1965), but. Collection. in the. secondary. serrate. 50｡. about up. loops;. secondary in the. way. quaternary. 1 petiole thick, about Discussion: These specimens shape,. form a. a. series. similar Honsbu. to. s. inouei. HuzIOKA. in the. KIMURA. but. those. in Japan,. 1.3 to. 1.9cm. side with. short. 16226;. glandular. the. my. villow. as. this. S･ vulpina. of. similar. veins･ TPM. 284,. Salix. sp.. to. concave. aplCal. at. which fossil leaves･. former. the. once. the. in. the. s'econdaries to establish. are. somewhat of. and such. northern marglnal. elliptical. Among. them,. and S･ alopechlea in the cuspidate to. ANDERSON. 477.. B. 1, 5, 6; Text･丘g.. 3C) 10. about at. apex, _cuneate. in aplCal point. Salix. veins have. the. mainly. genus. in less secondary of. from. reticulate. veinlets. specimens Aniaiflora. Miocene. early. in the. and. random. :. in the. superadjacent. fragmentary. too. at. teeth. venation. are. 6 an. bending. doⅥⅦ. the. the. brochidodromous. convex. or. up. marglnal. to. margin. and. percurrent. ultimate. narrow-oblanceolate,. acuminate and. loops;. apex,. midvein. curving. tertiaries. the. at. side. the. of. the. referable. especially. secondary. Leaves. wide,. teeth. marglnal. basal. :. are. serration. marglnal. PP. small long･. species. (Pl. 2,丘gs. Descripiton. and. brochidodromous,. case. to. is di庁erent. living. many. apex and the rounded NSM collections:. cm. from. resemble. growing. (TANAI. wide;. aplCal. weakly. all the specimens fossil willow, the. foliar shape,. leaves. fossil. areas. 6cm. to. branches. with. between. and. but. Among. are There serration. incurved leaves with. my. teeth. marglnal of loops,. status.. speci丘c. SuzuKI. et. rounded. gentlycurving. some. extending. veins forming. area. marglnal. or. branching;. foliar. in. and. loops. veins loops forming. area. of the blade,. in intercostal. ; tertiary. marginal. the. large. forming. portion,. secondaries. and. branches. on. or. veins from. diverglng. spaced,. in the middle. angle. some. with. lower. 60o. on. secondary. curving;. irregularly. palrS,. to. secon-. 3E). concave. apex. pointed. slightly. midvein. 9 alternate. to. with. Text-fig･. 2;. slightly. long. 12cm. teeth. side;. less. venation･. cuspidate (estimated) and 3.5. elliptlCal,. 5 to. ; marglnal. basal. on. Leaves. base,. at. rounded. Salixbakko. extant. A. sp.. 1,. (pl. 3,丘gs. :. of the. foliar. Salix. DescriptlOn. leaves. to. 45. 4825.. GSJ. :. I. in but is different shape, TANAI S･ misaotatewakii resembles Hokkaido Rubeshibe且ora of northeastern. in Japan in the growing S. hokiensis somewhat veins.. reported suzuKI,. Flora. ; midvein. side, long thick. cm. long. base, and. ; secondary. (estimated). and. margln. Serrulate;. slightly. convex. veins. diverglng. in.

(16) 46. K.. from an. the narrower. way,. polygonal. orthogonal. basal. ; ultimate. in. angle. part,. meshes with in intercostal. reticulate area. marglnal. 45o. abont. in the. angle. making. an. at. midvein. OzAKI. zlg-Zag. superadjacent area. and to. and. middle. extending. once. veinlets. the. upper. the. veins in. teeth. marglnal. branching;. at. its. on. ; tertiary. secondaries. entering. thrice. partsand. branching. and. petiole. 1. about. cm. long.. Discussion: Salix. to. able. Six by. their. though. narrow. teeth,. glandular. with. leaves,. the. and. HuzIOKA akitaensis different in cuneate base S･. willow,. et. acute. and. hokkaidoensis. TANAI. N.. et. SuzuKI. by. my. (1965),is. margin. somewhat. Another. teeth･. refer-. surrulate. are. (1974), thoughshorter;. marglnal. are. apex,. shape,fine leaves. These. venation･ UEMURA. their. ln. foliar. oblanceolate. S･. to. lacking. mostly. similar. but. they. comparable. distinguishable. The. are. fossil from. leaves. specimens serrulate Tatsumitoge are someclosely margln･ in foliar shape fine acute teeth what similar and S. babylonica to those of LINNリWhich is orlglnally in China been has and growlng widely cultivated in the world. Collections:. NSM. PP. 16207,. B;. 16208A,. Salix (Pl. 3,丘g. Description base. stout,. ; midvein. 9 palrS,. considerably. with from. branches the. midvein distinct but in. midvein･ margln or. iJICurVed, of. slightly. making. ; marglnal. teeth. by. the. above-described. in. the. Tertiary. of. S･ vulpina Collection: NSM extant. up. apICal. a. nearly. tertiary. This. Japan. ANDERS. PP. is. fossil. leaf. no. or. similar. is somewhat. cf.. Betulaceae. Alnus. GAERTNER. japonica (THUNB.) (Pl. 5,丘g.. angle. to. COnneCtlng leaving. in intercostal. veins. to. perpendicular. is referred. 16209.. Genus. about. of 40｡. margln,. slightly. in Japan.. growing. Family. Alnus. apex. leaf. There. characters･. veins. and. the. veins not preserved,･ irregularly spaced, straight. straight. complete. the. an. apex. 3 intersecondaries. almost. acute. with. at. of loops;finer. series. side, almost. at. along. ; 2 to. acute. ; secondary. midvein. slender;. side ; petiole missing. Discussion･･ A single,. the. creeplng. secondaries. area. on. the. right angle, irregularly percurrent,. thin,. concave. l･9cmwide, arcuate. gently. nearly. B.. 3F). and. from. slenderly. superadjacent at. marginal. Serrulate. prominent,. diverglng. spaced,. Text一点g.. long. oblong'5･Ocm. slightly. 1rregularly. 80o,. area. Leaf. :. 429, 480A,. C. sp.. 3;. TPM. 1). STEUD.. to. species. on. concave. the to. similar. genus my. basal. Salix. specimen to. those. ｡f.

(17) Late. Discussion: Alnus. Only. hold. the. fossil. organs. other. use. the. of of. species. TANAI. et. (Acad.. SINICA,. ONOE. the. and. of. extant. Alnus,. in. growing. A. japonica,. A.. bifurcating. the number and Collection:. TPM. secondary. description. from. is closely A. incana. similar MoENCIi.. Japan. by. My. way. and. enter. teeth.. marglnal. I with. then. collection, fossil leaf.. Among. A.. the. TANAI. eX miojaponica (1961), A.. ellipsophylla LI (1943) by the foliar shape,. small. teeth.. marglnal. sp.. 4,丘g.. My. Europe.. Of the. 12). fragmentary. leaves has. protohirsuta,. epithet, has. protohirsuta”. been. widely. been. identical. are. invalidly from. reported. the. A.. to. al-. published, Miocene. of. authors.. leaves. fragmentary. too. are. Collections:. NSM. PP. to. 16134;. Betula. Betula. LINN.. TANAI. protoglobispica. TANAI. et. ONOE,. status.. speci丘c. 288, 464.. (Pl. 5,丘g. protoglobispica. a. establish. TPM. Genus. Betula. the. as Alnus protohirsuta ENDO single leaf was丘gured Without ENDO A. Miocene Sendai area by the (1955). Protohirsuta of to TuRCZ. A. hirsuta Asia the extant and also to Of eastern. but. some. shape,. A. :. "A.. extant. 351.. (Pl.. though. this. and. veins,. Alnus. Protohirsuta,. to. WEYLAND. palaeojaponica. oblong on. the foliar. of. arranged. Tatsumitoge. is separable from K. SuzuKI (1961), A. protojaponica. 1978),and. Discussion. bifurcate. leaf. my. leaves. to. coarsely. small and Alnus in the. 47. in the. Japan. sometimes. name,. I. Flora. identical. nearly. that. veins. teeth,. marglnal is no. There. STEUD.. secondary. characteristic. Tatsumitoge. is. leaf. one. japonica (THUNB.). the. Miocene. Geol.. ONOE. et. 5). Surv.. Rep.. Japan. 187:. 24.. Pl. 3,. i.. 1,. 2. (1961). Discussion. bispica TANAI by. the. oval. et. shape,. nearly. is closely. Honshu,. Collection. identical. are leaves complate from the Hokiflora reported. nearly. ONOE. foliar. protoglobispica central. Two. :. similar. the. TANAI. to. widely round B. globispica. symmetric to. by. extant. Japan. :. TPM. 462, 463.. Betula. cf.. mioluminifera. (Pl. 3,五g.. Hu. 7, pl. 5.丘gs.. et. CHANEY. 2, 12a). to. Betula and cuneate. StIIRAI. Protoglo-. ONOE. (1961) base.a. growing. in.

(18) 48. K.. OzAIくI. Palaeont. Sinica A. 1 : 30. Pl. 4. i. 6, Hu et CIIANEY, mioluminifera 1 (excluding Pl. 5.i. 6) (1938). Pl. 9.i. HuzIOKA, Jour. Min. Coll. Akita Univ. A. 3(4) : 72. Pl. 6,i. 4 (1964).. Beiula. f･. Pl･ 7･. 1-3,. Discussion are. and. are. also. and. the. small. marglnal in Japan. NSM. :. B.. PP. CHANEY. et. 16138A,. ENDO. WINKL.. 16139,. are. are. is. Genus. invalidly. new. speciefic B.. miomaximowicziana (1955) based. ENDO. formation. on. Prefecture･. of Akita. All. published･ The status･. a. But, leaves. my. uB･. miomaximo-. REGEL. maximowicziana. in. growing. 9, pl.. ユour. Fac.. HuzIOKA,. 471, 496.. LINN.. heigunensis. (Pl. 3,丘g. heigunensis. 328, 377,. Carpinus. Carpinus. J. by. and. Collections:. Carpinus. Betula. to. description. been. extant. 10, ll). identical. without Sanzugawa. the. to southern. sp.. in southern Saghalien. PP NSM 16007, 16092; TPM. Japan. northern. to. similar. in central. growing. 9; pl. 4.丘gs.. has epithet, miomaximowicziana, too a fragmentary to establish. wicziana”. slightly. 16159A.. nearly. which wasfirst丘gured late leaf from Miocene. single the. leaves. My. to. round. fine. luminifera. (Pl. 2,丘g. :. shape,. base. Betula. Discussion. are My leaves veinlet. from the Miocene reported. ultimate. rather marginal serration･ and B. onbarensis from the Onbaraflorule reported basal but are different in the round shape and are to B･ REGEL specimens similar schumidtii. My. to. and. but. CarPinus,. of. to. (1961), teeth･. those. lanceolate-oblong. the. similar. ONOE. TANAI. Collection. Hu. inequilateral. slightly. by. growing China.. by. to. similar. branched. more. mioluminifera of China. leaves. somewhat. are. or. once. Betula. to similar Shanwang且ora. These. leaves. in the. separable. cordate. My. :. HuzIOKA. 4.丘gs.. 5-9) lmp.. S°i. Hokkaido. Univ.. ⅠⅤ. 4(ト2). :. 149･ ′･. (1938).. Carpinus. f.. KoNNO,. subyedoensis 1-4. et. SuzuKI, TANAl, TANAI. Geol.. nudum).. Central. in. Shinano,. Honma,. pl･ 7･. (1931).. HuzIOKA. TANAI. (nomen. K., et. NISIIIDA,. Sci.. ONOE,. et. HuzIO壬くA, NATSUO,. N.. Geol.. Surv.. Sci.. 1. ibid. 1. :. :. 195. 235.. HuzIOKA,. Jour. Min.. TANAI. N.. et. Mus.. Jap.. Flora. Pl. 40. Pl. 47.i.. Coll.. Akita. SuzuIくⅠ, Palaeont.. 14. Pl. 3. f. 3-5 (1960). 10: 39. Pl. 8. i. 1･･-5 (1961).. 3:. Univ.. Hokkaido. SuzuIくⅠ, Tert.. ibid.. Sado. Fukushima. Fac.. Jour.. Publ.. Rep.. Rep.. 187:. 29. Pl. 4.f.. Univ.. IV.. ll(2). 1. of Japan. i.. 5 4,. 299.. 118. Pl. 9.i.. 3,. Pl. 6,. 6,. 7,. 10. ll.. i.. 1,. 9,. 10,. i,. 6-8. (1961). 10,. Pl. 10.i.. 13 6. (1961). (1963).. (1963). (1963).. 5. Univ. Soc.. :. :. A.. Jap.. 3(4) Sp.. :. Pap.. 76. Pl. 7. 10:. 19. I,l. 13.. (1964). i. 8 (1965)..

(19) Late. IsfIIDA,. Men･. TANAI. Fac.. N.. et. SuzuKI,. OzAKI,. Sci.. Min.. Rep.. 4･5cm. double. on. secondary. vein from. generally the. midvein. apical. Sending. o庁. parts middle in intercostal. veinlets Discussion KoNNO. My. to. invalidly. published,. the. Miocene. from. the. C. subyedoensis,. late. the. but. based. Inland. veins. were丘gured.as Miocene Omi且ora. Carpinus. bracts. of Carpinus MAXIM.. NSM. TPM. PP. 16000,. 268, 306, 310,. that. 16099,. (Pl. 5,丘g. Holotype,. NSM. Miocene).. Saji-mura,. Tottori. Description with. several. apex. acute. :. Scale. fine acute with. round. PP. ; ultimate. C..tschonoskii. been. and been. are. unfortunately from. reported. widely. various. growing. leaves. The. authors. heigunensis,. bract岳 was. which. lsland in of Heigun ”subyedoensis” for the. closely. similar. 16141-16143,. 16153,. to. those. 16154;. GSJ. the. of. 4788,. OzAKI,. 9; Text一点g.. 16150.. Species. new. 2C). Upper. Tochiwara. Formation. (Late. Prefecture.. semi-oval, teeth,. has. the Miocene to heigunensis. are. percurrent. orthogonal. extant. bracts. Carpinus. and. 469.. hokiensis. Carpinus. Type:. has. by. up. basal. falcatibracteataHu both. and near. subyedoensis in central Honshu. the. subyedoensis,. to. parts. meshes. 1938) from. leaves. C. tschonoskii. 4816;. Japan.. western. and. by. to. diverging. making. curving.. region. I prefer. prlmary. 丘ne,. veins. slightly. arejdentical. (HuzIOEA,. of. long,. subyedoensis”. bracts. on. Sea,. termination. pentagonal. the epithet,. locality. slg-. curving in the. margln. tertiary. represented. although”C. of Japan and its adjacent. Collections: 4815,. and. from. speciemens. Tatsumitoge. reported. extant. bracts. middle. slightly. to. quadrangular 1 cm about. peliole leaves. at. basal. the. slightly. ; quaternary. making. or. at. to. the. near. C. subyedoensis is closely similar in Japan and Korea, and also to C.. identical. fossil. straight. (1931).. growing China. central. Seto. 4 branches. or. or. spaced. description. MAXIM. in. almost. 3. round,. veins. largest. middle. closely. Many. Without. KoNNO. by. angle. rather. simple;. :. in the. part,. to. Simple craspedodromous; 12 to 15 pairs, alternate,. veins. part,･. veins. secondary. side,. aplCal. ;丘fth-order or. plnnate,. (estimated). cm. obtuse. concave. in the. area,. none. basal. on. 40o. (1970).. Pl. 3. I. 1-3. 329.. :. 6 to 9 base. par. 1 branch. and. ⅠⅤ. 15(1-2). attenuate-acuminate,. sigmoidal. about. in the. margln,. meshes. an. at. Univ.. 4 teeth. secondary. 3,8. 3了(1) : 74. Pl. 6.i.2,. attenuate-acuminate,. Venation. straight;. 49. lanceolate-oblong,. generally. and. veins･. 30o. about. the. side. Leaves. teeth. marglnal. aplCal. each. at. ;. Mug.. apex. I. 5(1) : 48. Pl. 4.i.2 (1972). 16(4) : 707. Pl. 5. i. 3-6 (1973) 25 ⅠⅠ.21 : 10. Pl. 1.i. 24, (1974).. A.. Sci. Univ.. :. with. Hokkaido. Univ.. Natl.. wide,. serrate. of lamina. moidal. Natl.. Flora. andMineral.. Sci.. Akita. Bull.. description. 2･5 to. margln part. Coll.. Yokohama. Supplementary 1ong･ and. Jour. Fac.. UEMURA,. et. Tatsumitoge. Sci. KyotoUniv.Geol.. (1972). HuzIOKA,写Jour･ HuzIOKA. Miocene. the. terminal;. 17mm. long. and. other side･almost base round with. 8mm entire short. wide, and thick. one. side bulged. slightly bulged stalk;. longest. ,I.

(20) 50. K.. prlmary. Veins from. vein reaches. upper. tending. from. the. one. bract. length up. curving. connecting apex. the. the. small. secondaries. former are. One. the. growing. been. Collection:. the. none. and. venation. PP. Discussion. the two. Miocene bracts,. Pl. 9. Fac.. Hu. mioturczam'nowii. (Hu. These. the. of. to. close. two. extant. bract. extant. the. species. similar. to. this. Sci. Kyoto. the. leaf. Univ.. 12b). Palaeont.. Geol.. Sinica.. 1. A.. i.. Pl. 9.. 33.. :. by. may. Hu. basis. 6,. 7. excluded. 1938:. a. pl. 12,丘g.. leaf. a. 1). 7. (1970).. fragmentary this. leaf and its by species. bract,. Miocene as. 4,. (1938) from. CEANEY. and. of a from. (1970)figured. from the species One leaf,丘gured. of this CIiANEY,. be. 3了(1) : 73. Pl. 6.i.. Mineral.. first reported in China on the. IsHIDA. in Japan.. and. was. specimen. serration. leaf. 8,. CHANEY. et. (1938).. species Sbanwang且ora. single marginal described single kawa Prefecture Shanwang. f. 2). CIIANEY,. et. This. but. among. meshes.. especially. fossil. no. sometimes. meshes. those. Hu,. the. 16150.. Hu. mioiurczaninowii. Men.. to. teeth. is. percurrent,. tertiary. is similar. and. from. length. orthogonal. the. There. (Pl. 5,丘gs.. IsliIDA,. of. another part. reported.. NSM. (excluding. apICal. making. marglnal. China.. CarPinus. Carpinus. margin. till the bract. Vein. one-third. with. C. austrosinensis. and. till about veln,. among bract. prlmary. secondary. the. and. exprlmaries at its distal part. one. side. irregularly. ; tertiaries. up. four up. margin. one-forth. well-preserved. SOuthwestern. has. specimen. the. along. along. veinlets. entire. o庁one. curves. or. curving. the. sending. which. three. weakly. the prlmaries. BuRKILL. shape, ln. about. connecting. about. teeth. ; ultimate. pubescens in. at. marglnal. Discussion:. and. part,. 30o. side. in. slgmOidally. between. veins. or. up. base. secondaries. of. toothed. teeth;. curving the. slightly. ; Secondary. Carpinus. one. from. with. entering. straight. larger. its upper. at. angle. almost. base:. the. an. in the. marglnal. reaching forking near the. base. bending. apex, at. tooth;. marglnal. and. of. the. reaching. basal■position. the. OzAKI. and supplementary Norosbi且ora of lshiC. from of miocordata. to transferred recently Betula mioluminifera (Plant fossils of China, vol. 3, pl. 48,fig. 6),but it seems to be included in C. mioturczaninowii. A bract (Plant of C. mioturczaninou)ii fossils of China 3, 47,丘g. be to those vol. pl. 7) may similar rather of C. and. NATIiORST. stenophylla C. mioturczaninowii ing. in central Collections:. to. is similar. Japan,. western. NSM. PP. to. the. Korea. extant. was. C. turczaninowii. 16037,､16138B,. and 16149,. 16151,. B.. Carpinus. subcordata. (Pl. 3,丘g.. HANCE. grow-. China.. NATHORST. 8, pl. 5,丘g.. 4). 16156;. GSJ. 4813,. 4814A,.

(21) 1Jate Miocene. Car♪inus. NATHORST,. subcordata. 20. Vt.. Kgl.. Sv.. are. referable. oval. shape. My. leaves. Korea. and. China.. leaves. Two. :. teeth･. deeply. of the leaves, are. Collections:. Acad.. to. akin. NSM. Description:. PP. the. angular. apex. 5 teeth with. prlmary. TPM. thin,. few. turning tertiaries. lanceolate;. to. lateral. base,. upper. strongest; and. midrib. reteculate. Discussion: extant. Carpinus. fossil. leaves. are. distinguishable. (K.. SuzuKI,. is separable A. shape･ of. ONOE. (1961). my. also. in finer. 1961). from. from. my. bract. those. of. C･. specimens,. are. seems. specimens Collections:. reaching. veins. too. similar. marginal. Miocene. leaves. in. more. its leaf of. but. PP. similar to. Corylus. veins to. to. middle making. pentagonal to. similar Korea. and. C･ tschonoskii lamina.. Formation. These. MAXIM., C.. of Fukushima. those. but. are. protolaxLj7ora Prefecture. longer and the foliar veins ONOE, and 1961) is closely similar to C･ miocenica indicated by TANAI･ and to. establish. 16145-16148;. Genus. the. from. secondary. (TANAI. miocem'ca. closely. broader. and. Ryozen. from. ;. falcately. diverglng. ; tertiary. in Japan. the. of. lobe. single. meshes.. are. growing. those. serration. the. to be rather ill-preserved. NSM. specimens. or. none. quadrangular. quaternary. veins Bne,. quaternaries. Veins. teeth. glVing. tertiary. ; main. diverglng. making. the. laxijiora BLUME to. wide. to marglnal. none veinlets within These leaf and bract. ; ultimate. cm. portion; straight,. teeth,. veinlets. 5 prlmary. to. secondaries. ; quaternary. the. of. deltoid;. l･O. and 4. ;. spacing. ; ultimate. meshes. upper. teeth;. ;. veins. almost. marglnal. enter丘ner. fine, tri-. secondary. extending. closely. serrate,. teeth. the. at. entering. Which rather. two. of each. end undulate. and. long. lamina,･. of. 2.5 to. and. doubly. ; margin. 10 palrS,. margln. l･7cm. in Japan,. growing. (estimated). long. part. the. at. margln. small,. of the. orthogonally meshes. lobes. widest. about. areas,. about. BLUME. round. sometimes. pentagonal. midvein. part. the. in intercostal. quadrangular. at. alternate,. the. near. and. 5cm. to. base. vein. near. up. base. NATHORST subcordata fine cuspidate marginal. 6, 7). 4. oval. straight, to. petiole missing. Bracts three-lobed,. the. 18,. sp.. largest. apex,. thin,. opposite. percurrent. making. secondary. rather. slightly. off. 14-16,. i.. 502.. attenuate-acuminate; per. thin,. to. ovate. cuspidate. Vein. secondaries. a. 20: 39. Pl. 3.. C. cordata. extant. 16155;. Leaves. wide;. 4 to. then. 51. Carpinus. cordate. (Pl. 5,丘gs.. with. I. Handl.. to. CarPinus. 2.8cm. Flora. (1883).. Discussion in the. Tatsumitoge. those a. of. speci丘c. TPM. Corylus. subsieboldiana. C. tschonoskii. 468.. LINN.. K.. SuzuKI. status.. MAXIM.. My.

(22) 52. OzAKI. K.. (Pl. 5,丘gs. Corylus. K.. subsieboldiana. Rep.. 10, ll). Suzu王くL, Sci.. Fukushima. Univ.. 10:. 45.. f.. 10,. Pl.. 3,. 4. (1961). Discussion. foliar. ovate. pentagonally. leaves. These. :. serratioⅠ1 that. marglnal. somewhat. smaller the living. to similar is slightly similar. the. C.. subsieboldiana. its. with. obtuse. widely. BLUME. sieboldiana. in. growing. MAXIM.. C. mandshLm'ca. to. Corylus. to. t)asal shape, and ratber丘ne each at termination of secondary veins, although This fossil is closely specimens. orlglnal species. shape,. exserts. than. identical. are. Japan. ln. growing. Korea. and. northern. and. and. western. China. Collections:. NSM. PP. 16137;. TPM. 465.. Family Genus Castanea. Fagaceae. Castanea. MILL. TANAI. miocrenata. (Pl. 6,丘g. Casianea. miocrenaia. Castanea. vulgaris. TANAI. 10a. 15,f.. Pl.. Discussion CARR.. by. separable tertiary in. My. leaves. shape,. in. secondary. arranged are. of secondary is separable shape,. Kydshu,. (1961) differs. veins, from the. C･マrenaia SIEB･. lo. SuzuKI but. df the. 30. Pl. 5.i.. Akad.. (1961). 20(2) : 84.. 1-4. Handl.. other. of. secondary. Zucc･. et. somewhat. the. miocrenata base,. secondary. C.. which. is widely. PP､6445;. GSJ. tertiary. veins. Fukushima. in. in number C. veins. miocrenata by the oblong foliar. tertiary Castanea. and prominent is closely similar distributed in southwestern. awn. to. l. Genus Fagus. 4801,. Fagus. PalieojaPonica (Pl. 6,丘g.. 4802,. 4808;. TPM. LINN. PTANAI 7). et. ONOE. spaced ONOE of the. number. Japa?I. collection占:NSM. et. are. miocrenata. veins C. miocrenata. veins.. TANAI. spaced Formation. resembles. but. the coarsely. cordate. coursely and Hirasawa. in closely spaced fossil species of. and. acutissima. teeth,. marginal. veins to. round. Quercus. extant. and. Castanea. to. referable. asymmetrically. less number. the. terminal. K.. of the. shape. marglnal secondary veins, aristate Miocene C. lanceolata described from the by. 187:. Vet.. Sv.. those foliar. teeth. Prefecture. Rep.. Japan Kgl.. resemble. BLUME. irregularly. the. veins. foliar. the. Surv.. NATHORST,. leaves. Q. variabilis. and. Geol.. 1). (1883). These. :. ONOE,. et. fossilis. LAMARK. ONOE. et. 96, 205.. at. the. the extant Hokkaido.

(23) Late. Fagus. TANAI. palaeoja♪onica. Miocene. Tatsumitoge. ONOE,. et. Geol.. Flora. Surv.. 53. I. 187:. Rep.. Japan. 3-5. f.. Pl. 7.. 35.. (1961). Fagus. protojaPonica. 11(2):. 308.. TANAlet. N.. HuzIOIくA. 5-9. f. Fagus. HEER 6. FagusjaPonica (1958). MuRAI,. ibid.. Sci.. Mus.. 16(4) Geol.. KoNNO,. :. by. Hokkaido. ⅠⅤ.. Univ.. i.2,. Pl. 6.. Pl. 8.. 710.. Central. MuRAI,. Palaeolongipetiolata MuRAI,. MAXIM:. non. Rep.. 4,. 15(2): 26. Pl. 7.i. fossilis NATHORST,. 5,. 4-8. i.. 5. Pl.20.i.. (1973).. in. Shinano,. Rep.. Tech.. Homma,. their. reported. from by. Kgl.. are. Sv.. Univ.. Pl. 7.. Univ.. 15(2). 29.. :. is. species. marginal marginalminute. MAXIM.. F.. Collections:. PP. NSM. 16085;. Fagus. F.. 4839i. 84. Pl.. :. TANAI spaced. SuzuKI of. (1959) is to. charac-. the. extant. Korea. westhern. 15. et. K.. is similar. Of. F.. and. I. TANAI. (NATHORST). stuxbergii. _ト3. Fukushima'Prefecture. protojaPonica palaeojaPonica. 4819,. GSJ. i.. closely. protojaPonica. of. multinerui云NAKAI Chin畠.. western. central. 一and. SuzuKI. F.. teeth.. Of Japan, Of. :. 20(2). ′Handl.. margln. Formation. teeth. 20. Pl. 2.. :. palaeojaponica. F.. from. separable. Tennoji. crenate. Fagus. cremate. veins,. ll. (1962).. Vet.∫ Acad.. to. referred. secondary. Miocene. the. SEEM.. engleriana. of. This. the. japonica. leaves. number. its. by. These. :. veins.. lwate. Tech. Iwate. i. Pl. 8こj.. ENDL.. tertiary. F.. 10: 21.. (1883).. 6. terized. Pap.. Sp.. Sci.. (1962).. Discussion. Japan. Natl.. SAPORTA;. ∀ar.. auct.. sieboldii. ONOE. Japan. Fac.. (1931).. Pl. 7.f.. f.. Soc.. Jour.. (1965).. Bull. non. auct.. antipojii. Fagus. SuzuKI:. (1961).. i. 8). UEMURA,. et. TANAI,. K. 6. Palaeont.. 16.. Pl.. pliocenica. 5,. 15.i.. SuzuKI,. (excluding Fagus. non. auct.. Pl.. ′l′＼. _/). Fagus. 4,. f.. 5,. 8,. Discussion:. ”,. A. textj.'2g-j. of leaved from collections. rec′entfi■'discdssed is related. TANAI. by to. the. extant. PALIB. Chin包;I '･F:Jl^h･d5,atae. crenata. BLUME. grOY(ing. 16234-16238;. GSJ. NSML,PP. in. four. the. Tatsumitoge This. (1976). central. starlk ∫o/i･cupules. Colfe.ttions :. and. TANAI･. Fag?(亨}tuxbergii (NATEORST). stuxbergii. Sci･ Hokkaido J叩･ Fac･ (ゝi976).(see synonymy).. number. of the. nearly､由half to. (p/i.6,figsy. 2-6). (NAT甲声苧T) TANAI,. stuxbergi 1.. }. Japan, 16012,. F. lucida eX. basis. the. on. REIiD.. HAyATA. especially. 16016,. 4772;;4778, 4789･; 4809. 16038, ; TPM. Quercus. miocrisPula. the丘rst. -16053,. 16058,. 93, 197, 211,. ;. QuダrCuS. of the. growing. 1. Genus. for these. et WILS.. to. LINN･ HuzIOKA. 17(2). :. 276･ Plc. occupleS. all of which are locality,. cupules,. epithet. IV･. Univ･. referred was. specimens. F.. Mogispecimens. ln. grOWlng. in. Western. Formosa. one. 16062, 313.. in. and the. F.. short. 16085-16088,.

(24) 54. K.. OzAIくⅠ. (Pl. 6,丘g. Ouercus. HuzIOKA. miocrispula. Proc.. Trams.. 9). Palaeont.. Soc.. 187:. 30. Pl. 6, i.. 15:. Japan. Pl. 25,. 196.. 3. i.. (1954). TANAI. ONOE,. et. Geol.. Surv.. Jour. Fac. Sci. Hokkaido Pl. 13.i. 2) (1961).. HuzIOKA. Quercus. UEMURA,. et. 3. ll.f.. Pl.. Discussion. ing. these. Q. mongolica and Q. dentata. are. and. have. and. by. the. FISIiER. Collections. NSM. :. PP. aliena 3,. 71l.. veins. known. the. But,. extant. leaves. the. foliar. obovate teeth,. marglnal. above-. four. those. to. some. THUNB.,. serrata. species. leaves.. oblanceolate-oblong. Regard-. are. 0.. basal. the. ln. of. shape, 10. generally. Q. miocrisPula. long.. (1954).. The. 0.. closely. ∀ar.. mongolica in the cool-temperate. to. was. similar. grosseserrata. forests. of. 16157,. 16185-16190,. OzAKI,. K.. SuzuKI,. 16283;. new. 1,. 4772,. GSJ. 4823.. species. 2). Sci.. Fukushima. Rep.. 10: 55. Pl. 12.. Uuiv.. Tottori. concave. to on. apex;. the. 16159.. Bull.. aplCal. wide,. apex. and. strong,. to subopposite,. midvein. at. 16. an. base. cuspidate,. dentate;. side Vein. Tochiwara. Upper. oblanceolate-oblong. undulately. prlmary alternate. from. UEMURA,. et. Natl.. Mus.. 16(4). (Late. Formation. Prefecture.. large,. (estimated). serrate. HuzIOKA. PP. NSM. Leaves. :. 9cm. HuzIOKA:. (only) (1973).. Saji-mura,. strong,. diverging. there MIQ.,. separate. Protoaliena. non. Holotype,. Description. acute. 6. 6.I.. Pl.. Miocene).. up. miocrisPula Q. Proto-. and. in taxonomy.. by. acute. sp.. lacking. resembrance. only. Quercus. which. 5mm petiole less than Korea by HuzIOKA of. Bl.UME:. non. auct.. Type:. Slightly. 60.. 4. f. (1961). Quercus auct. miocrispula. coarsely. 19:. followlng. of the. n.. Acutually,. (Pl. 7.丘gs.. 6 to. of. all. :. are. (BLUME) tO. base, large. 16083,. Quercus. and. Bull.. Asia.. northeastern. Quercus. Res.. leaves. Japan,. of. and its variety, luxuriant growth. WILSリShow. et. to. case. short Miocene. oaks. THUNB.,. auriculate. Q. mongolica. species, (BL.) REED.. Mus.. confused. grosseserrata. characterized. veins from. sometimes. close. some. slightly. 15 secondary first reported. Kai. laeves. their. deciduous var.. Q. miocrisPula. Ho-on. (1961). 7 (excluding. 2,. (1974).. 1. Pl. 7.f.. distinguish. to. when. been. of the. is difficult in. narrow. has. species. fossil species. species. 351.. :. (only) i.. 15.. Pl.. of leaf impressions et ONOE, Q. Protoaliena. especially. among. mentioned. Saito. 311.. 1. basis. TANAI. species. aliena. 17(4). Mus.. di氏cult. the. on. only. et ONOE,. BLUME. ll(2). IV.. OKUTSU,. living. the. hybrids 0.. non. oaks. Thus,. part.. Sci.. It is sometimes. :. TANAI. Nat.. BLUME:. 0. protoserrata. dentata. Univ.. :. (1940).. fossil desiduous HuzIOKA,. Buu.. auct.. crisPula. Rep.. Japan. TANAl,. convex. straight neary. angle. basal to. parallel of 40o to. 19. (estimated). cm. to obtuse,. cuneate teeth. marglnal on. to. large,. side with. curving each. long. margin. straight. slightly. to. curving. secondary gently; 16 to 20 palrS,. other,. 50o, spacing. Irregularly. in. :.

(25) Late. some. two. almost. running. part,. secondaries. ; quaternary. the. loops. form. not preserved; veinlets Discussion: This. to. veins. the. making. margln;. except. basal. tertiary. veins. pentagonal. mainly. i.5. thick,. and. 55. teeth. marglnal. at. stout. petiole. I. Flora. Tatsumitoge. straight. which. 丘ne, percurrent. Miocene. or. rather. ; ultimate. meshes long.. 2.8cm. to. one. by a large is chracterized number of the species large are foliar size and long thick petiole. My specimens secondary veins, in southwestern Japan, closely Q. aliena BLUME akin to the extant growing in Korea f. et TIiOMS. 0. gribithii HooK southwestern and China, and growing. China;. but they. secondary. in. longer. and. leaves. of Q. miocrispula is distinguishable by the. by. and. the. oblanceolate-oblong. from. from. (1869) different. HuzIOKA. by. the. long A. 6). secondary. obtuse. specimen has. part. the. and. the. and. Collections:. to. similarity large. this. foliar. new. petiole. PP 16157-16159;. NSM. Quercus. TPM. K.. Quercus. protosalicina Pl. 3. i. 5-6. Discussion to. Quercus. mation. :. SuzuKI,. Two. leaves,. Fukusbima. K.. SuzuKI. Assoc.. (1961,. in the. many. number. size, but. the. specimen. pl. the. of. lacks. SuzuKI. Collab.. Geol.. 9: 36. Pl･ 2･. Japan. are. and. TANAI. Jour. Minn.. et. Coil.. Geol.. Akita. Univ.. of. f･. 13,. Surv. A.. Japan 3(4). :. 79.. Japan. and. are. referred Tennoji For-. the. extant. ()･. Korea･. ONOE. et 4,. to. similar. region. TANAI. half,. Miocene. late. closely. 8, pl. 7,丘gs.. ONOE,. lower. in their from. reported. Protoserrata. (Pl. 6,丘g.. HuzIOKA,. K.. thoughlacking. Prefecture,. Ouercus. Ouercus Protoserrata ′. 7 (1961).. species secondary. 3). in warm-temperate growing NSM PP 16091, 16096.. salicina Collection:. new. (1959).. Protosalicina. of BLUME. is. 343.. Protosalicina. Monog.. this. of the number TANAI ONOE and. by. and. (Pl. 7,丘g.. petiole,. Q. miocrisPula. as. 17. to. species. shape. thick. and. Q. furuhjelmi HEER. petiole. is similar to Q. Protoaliena, but. Q. miocrisPula. as. but. secondary ONOE et. TANAI. teeth. long. 16. and. the. of. number. specimen reported is included in 6). 6,fig.. 1.2cm. than. reported. close. veins. its aplCal. (1973, pl.. more. petiole. A. teeth.. large. serration,. marginal. marglnal. Q. gr塀thii. the. is similar㌧to. species and. of the. number. from. Q. Protoserrata. by. Alaska. of. marginal. UEMURA. and. from. large. and. large. large. ONOE. et. flora. Miocene. the. in the. by. veins. 6,丘g.. TANAI. 0. Protodentata. shape. and. petiole. is also separable foliar shpe,. Q. Protoaliena. veins.. petiole. HuzIOKA in foliar longer. in the. Q. aliena. 17 in number, new This. than. more. generally. thick. from. distinguishable. are. veins,. mainly. new. 5) Rep.. 187:. Pl. 9.. f.. 32. Pl. 3. 1. (1964)･. f･. 6･. Pl･ 6･.

(26) 56. K.. HuzIOKA. Quercus. UEMURA,. et. miocrispula. 30. Pl. 6. HuzIOIくA. i.. Discussion oblong. the. with HuzIOKA. and. Mus･. 16(4). TANAI. Pl･ 9･. 711･. :. ONOE,. et. and. shape,. slender ONOE,. Sci.. Mus.. 16(4). Geol･. f･.ミβJA,97芦)･ Suiv･ Jap早n. oaks. 1961,. are. small. rather. long. petiole. Pl. 9. f.. 711.. :. of Q. Protoserrata. foliar. rather. other. Natl.. leaves. deciduous. (TANAI. 9, f,. Bull.. The. :. Sci･. 4817,. NSM. PP. belong. 16072,. teeth. marglnal 30mm. 16120,. 16178-16183,. incurved compared. specimens reported UEMURA, pl. 6, f. 6; HuzIOKA and to by Q. Protoserrata the long. 16088,. by. Q. miocrisPula. as. 1973,. pl.. petiole and ONOE, 1961, aやd its long petiole.. 16229,. 16233;. GSJ. 4839.. (cupula). sp.. (Pl. 6,且g. Discussion splrally. Three. :. arranged. depressed. scale-like. specimens Collections. :. probably NSM PP. protrusions, belong 16193;. to. Quercus. by. Collection:. :. These. its fine NSM. amentaceous. cupules. resemble. Q. Protoserrata. TPM. are. which those TANAI. covered. of 0. serrata. by. THUNB.. ONOE.. et. 476.. sp.. (Pl. 6,fig. Discussion. 10). pelv‥iformis. Quercus?. of. with. length,. Two. Quercus. These. (oily) (1973).. characterized. generally. 6 to. of. 1. 1) considered its foliar shape･ A specimen as Q･ Protodentata reported (TANAI to Q. Protoserrata or pl. 7,丘g. 2) is similar Q. Protoaliena by to. are. Collections:. 18了:. Rep･. (only) (1961).. oblanceolate. tips. pointed. 6. Natl･ HuzIOKA:. non. UEMURA,. et. to. Bull･. auct･. OzAKI. (ament) ll). specimens. rachis and simple 6406. 16190B.. may. be. referred. to. male且owers. corolla.. PP. (To. be. continued).

(27) Explanation. 1-7. of Plate.

(28) Explanation (all Figs.. 1-4.. Figs･. 5-7･. Fig･ Figs.. 8･. hokiensis Pseudolarix. 16232,. 16207,. 16189.. Figs.. 16-18.. Picea. ugoana. Figs.. 19, 20.. Figs.. 2l, 23, 25, 26. 22.. Picea. Pseudotsuga. Figs.. 24, 28.. Figs.. 27, 29, 30.. TANAI. NSM. PP. et. ONOE.. sp.. cf. sieboldiiLCARR. NSM PP 16198. NSM. Cunninghamia. PP. stated) 497,. NSM. 16097,. ′16219･ PP NSM. 16112,. 16178. ×1･5･. 16195,. 16092,. 16113A,. TPM. 383. TPM. 459. 31, 32.. Sciadopitys. sp.. TANAI. occidentalis. GSJ. 4806A･. GSJ. 16195B,. GSJ (remains),. ×1.5,. GSJ. ONOE,. et. (NEWB.). CHANCY,. 4805. (outer. and. inner. B･. 16195A,. 16113B.. TPM. 500. Figs.. 16115,. PP. 4802. ×1.5･. TPM. 459. (cast. 6411.. protokonishii. Metasequoia. 16096, PP. Tsuga 4802,. 21), GSJ. HuzIOKA,. kaneharae. TPM. ONOE,. et. 1. otherwise. NSM. Hol･otyp?, japonica TANAI. n･,叩･. 9-15.. Fig.. unless. size. Abies HuzIOKA et UEMURA, ugoensis ezoana PP Keteleeria TAN今Ⅰ, NSM. Abies. 丘g.. natural. Plate. of. side).. 4832. ×2/3, GSJ 499, GSJ. 4789･. 4788,. TPM. of.

(29) K.. OzAKI. Sec.. 8 10. 23. 2丁. 28. II, Plate. 1.

(30) ExplanatiorL (all丘gures Figs. Fig. Figs.. l, 5, 6･ 2.. Salix. 3, 4.. Salix. BI. sp･. TPM TANAI. hokkaidoensis Cyclocarya. 4791 et. (TANAI. ezoana. Of. in natural. are. 480･ N･ et. Plate. NSM. PP. SuzuK[1 N･. Figs. Fig.. 7.. Juglans japonica. 8, 10. 9.. Betula. Pterocarya sp.. TPM. TANAI, sp･. NSM. 328.. YNU PP. IO80･ 16082,. 16135･. size) 16201･. NSM. SuzuK,). 478. Fig.. 2. PP WoLFEet. 16206･ TANAl･. GSJ4801･. TPM.

(31) K.. OzAKI. Sec.. ⅠⅠ,Plate. 2.

(32) ExplanatiorL (au natural Figs. 2.. Salix. sp.. A,. C,. NSM. Fig.. 3. Salix. Fig.. 4. Populus. hokiensis. Fig.. 5.. Po?ulus. kitamiana. Fig.. 6. Salix. Fig.. 7. Betula. Fig.. 8. Carpinus. Fig.. 9. Car?inus. sp.. hokiensis. TPM. 477. PP. TANAt sp.. x3/4,. otherwise. stated). 284.. TPM. Holotype, et. N･. Holotype,. Hu et cf. mioluminifera NATFIORST, subcordata heigunensis. unless. 16209. sp.. n.. n.. size. 3. Plate. Of. HuzIOKA,. NSM. PP. GSJ. NSM. PP. 16073･. 4825.. CHAN巴Y, NSM. 16075･. NSM. SuzuKt,. NSM PP. PP. 16155･ 16142･. PP. 16159A･.

(33) K.. OzAKI. See.. 1I, Plate. 3.

(34) Expl乱n&tion. (au Figs.. I-4･ 501,. Figs･ Figs. Fig.. Populus. GSJ. 5-9･. Car?inus. GSJ. 10, 1l.. Betula. Alnus. n･. size. sp.. unless. Paratypes,. Plate. 4. therwise TPM. stated) 493. ×3/4, NSM. PP. 16076,. TPM. NSM. PP. 16153B,. 4798.. 16153A,. 12.. natural. hokicnsis. of. sp.. heigunensis. HuzIOIくA,. NSM. PP. 16143,. 4778. sp.. NSM. TPM. 47l. PP. 16134.. x3/5,. NSM. PP. 16092.. TPM. 306,.

(35) K.. OzAKI. See.. II, Plate. 4.

(36) Expl乱A&tioTL. (all natural Fig. Figs.. 1.. Alnus. 2, 12a. 3.. Car♪inus. Fig.. 4.. Carpinus. Fig.. 5.. Betula. Figs Fig. Figs. 6, 7. 8, 12b. 9.. Hu. Carpinus. sp.. TPM. TANAI 468,. et. NSM. Carpinus. Hu mioturczaninowii hokicnsis n. sp. Holotype,. Corylus. subsieboldiana. K.. wise. TPM. 351.. NSM. PP. stated) NSM. PP. 16221. 16139,. 16138A.. ×1.5.. 502.. ONOE, PP. 5. other. CEANEY,. et. NATHORST, cf. steno♪hylla NATIiORST, TPM subcordata. Protoglobis?ica. Carpinus. 10, ll.. unless STEUD.,. (TrfUNB.) cf. japonica Bctula cf. mioluminifera. Fig.. Figs. size. Plate. Of. TPM. 462.. 16146. et. NSM. SuzuKI,. CHANEY, PP TPM. NSM. PP. 16156,. 16150. 465,. NSM. PP. 16137.. 16138B..

(37) K.. OzAKI. Sec.. ⅠⅠ,Plate. 5.

(38) Explanation. (all natural Fig. Figs.. I.. Castanea. 216･. miocrenata. Fagz＋s stuxbergii. size. TANA[. et. oE. Plate. unless. otherwise. ONOE･. NSM. PP. TANAI,. (NATf10RST). 6. NSM. stated) 16197･ PP. 16238･. 16235. Fig. Fig. Fig.. 7. 8.. Fagus. Palaeoja?onica. Quercus?rotoserrata. 9.. Fig.. Quercus 10. Oucrcus. Fig.. ll.. Quercus?. miocrispula sp. sp.. ONOE,. TANAlet TANAlet. ONOE,. HuzlOKA,. NSM. (cupula)･ NSM (ament)･ NSM. PP PP. GSJ NSM PP. PP. 16238･. 16193･ 16190. 4839･. x2･. 16178. ×2/3･. 16234,. 16038･. 16236･.

(39) K.. OzAKI. See.. II, Plate. 6.

(40) Exphnatio-n. (all natural ド g. F g. F g. F gs.. size. Plate. of unless. otherwise. 7 stated). NSM PP 16159. A. Qucrcus?rotoaliena sp. Holotype, 2. n. Paratype, TPM 343 ×3/4. Quercus?rotoaliena sp. 3. K. SuzuKI, NSM PP 16096. Quercus?rotosalicina TANAlet ONOE, NSM PP 16179B, 4, 5. Quercus ♪rotoserrata 1.. 16229..

(41) K1. 0zAKI. Sec･. IT, Plate. 7.

(42)