• 検索結果がありません。

ᢚไࡋࠊ㯤యࡢ㏥⾜࡜ඹ࡟పୗࡍࡿࠋ

59

ㅝ༸⬊Ⓨ⫱Ἴࡢฟ⌧ࢆಁ㐍ࡍࡿࠋࡑࡢᚋࠊ༸⬊࠿ࡽศἪࡉࢀࡿ࢖ࣥࣄࣅࣥ࡟ࡼ

ࡾ )6+ ศἪࡀᢚไࡉࢀ࡚⾑୰ )6+ ⃰ᗘࡀపୗࡍࡿࡀࠊ࢖ࣥࣄࣅࣥ⏕⏘ࡢపୗ࡟

కࡗ࡚෌ࡧୖ᪼ࡋࠊୖ᪼࡜పୗࢆ⧞ࡾ㏉ࡍࠋ⾑୰ࣉࣟࢪ࢙ࢫࢸࣟࣥ⃰ᗘࡣࠊ᤼

༸ᚋ࡟㯤యࡢᙧᡂ࡜ඹ࡟ୖ᪼ࢆ♧ࡋࠊ㈇ࡢࣇ࢕࣮ࢻࣂࢵࢡ࡜ࡋ࡚ࠊ/+ ࡢศἪࢆ

60

㸳㸫㸲㸫㸰㸬Ἢஙᮇ࡟࠾ࡅࡿྛ✀࣍ࣝࣔࣥศἪ 㸳㸫㸲㸫㸰㸬Ἢஙᮇ࡟࠾ࡅࡿྛ✀࣍ࣝࣔࣥศἪ 㸳㸫㸲㸫㸰㸬Ἢஙᮇ࡟࠾ࡅࡿྛ✀࣍ࣝࣔࣥศἪ 㸳㸫㸲㸫㸰㸬Ἢஙᮇ࡟࠾ࡅࡿྛ✀࣍ࣝࣔࣥศἪ

့ங㢮ࡢ㞤࡛ࡣࠊஙᏊࡢ྾ங่⃭࡟ࡼࡾࣉࣟࣛࢡࢳࣥศἪࡀஹ㐍ࡋ࡚ࠊ/+࡜

)6+ ศἪࡀᢚไࡉࢀࠊ

ẕື≀࡛ࡣἪஙᮇ㛫୰࡟ࡣࠊ᤼༸ࡀ೵Ṇࡍࡿࡇ࡜ࡀ▱ࡽࢀ

࡚࠸ࡿࠋᮏ◊✲࡟࠾࠸࡚ࠊ⾑୰ࣉࣟࣛࢡࢳࣥ⃰ᗘࡣࠊἪஙᮇ㛫୰ࡣࠊ㏻ᖖࡼࡾ

ࡶ㧗࠸್ࢆ⥔ᣢࡋࡓࠋ/+ࠊ)6+ࠊࣉࣟࢪ࢙ࢫࢸࣟࣥࠊࢸࢫࢺࢫࢸࣟࣥཬࡧ࢚ࢫ ࢺࣛࢪ࣮࢜ࣝ࡟㛵ࡋ࡚ࡣࠊἪஙᮇ୰࡛ࡶୖ᪼࡜పୗࡀ⧞ࡾ㏉ࡉࢀ࡚࠸ࡿࡇ࡜࠿

ࡽ㞤࣐࡛࢘ࡣࠊἪஙᮇ୰࡛ࡶ᤼༸ࡀ೵Ṇࡍࡿࡇ࡜࡞ࡃⓎ᝟࿘ᮇࡀ⥔ᣢࡉࢀ࡚࠸

ࡿࡇ࡜ࡀ☜ㄆࡉࢀࡓࠋ࢘ࢩ࡛ࡣࠊศፔᚋ⣙ ᪥࡟Ⓨ᝟࿘ᮇࡀᅇᖐࡋࠊࣈࢱ࡛ࡣࠊ Ἢஙᮇ୰࡟ࡣⓎ᝟࿘ᮇࡣᅇᖐࡋ࡞࠸㸦᳃ࡽࠊ㸧ࠋ࣐࡛࢘ࡣࠊศፔᚋ᪩ᮇ࡟Ⓨ

᝟ࡀᅇᖐࡋࠊศፔᚋⓎ᝟ࡀ᪩ᮇ࡟㉳ࡇࡿෆศἪᏛⓗゎᯒ࡟㛵ࡋ࡚ࡣࠊ㸲❶࡟グ

㏙ࡋࡓࠋ

㸳㸫㸲㸫㸱㸬ྛ✀࣍ࣝࣔࣥࡢศἪࣃࢱ࣮ࣥࡢẚ㍑

㸳㸫㸲㸫㸱㸬ྛ✀࣍ࣝࣔࣥࡢศἪࣃࢱ࣮ࣥࡢẚ㍑

㸳㸫㸲㸫㸱㸬ྛ✀࣍ࣝࣔࣥࡢศἪࣃࢱ࣮ࣥࡢẚ㍑

㸳㸫㸲㸫㸱㸬ྛ✀࣍ࣝࣔࣥࡢศἪࣃࢱ࣮ࣥࡢẚ㍑

㠀ዷፎᮇ࡛ࡣࠊࣉࣟࢪ࢙ࢫࢸࣟࣥ࡜ࢸࢫࢺࢫࢸࣟࣥࡢศἪࣃࢱ࣮ࣥࡣ୍⮴ࡋ

ࡓࠋ㯤యᮇ࡟㧗ࡃࠊ༸⬊ᮇ࡟ప࠿ࡗࡓࠋࡇࢀ࡟ࡼࡾࠊࢸࢫࢺࢫࢸࣟࣥࡢ୺࡞ศ Ἢ※ࡣ㯤య࡛࠶ࡾࠊ༸⬊࡛ࡣ࡞࠸࡜᥎ᐹࡉࢀࡿࠋ㯤య⣽⬊࡟ࡣࠊࣉࣟࢪ࢙ࢫࢸ

ࣟࣥ࠿ࡽࢸࢫࢺࢫࢸࣟࣥࡢኚ᥮㓝⣲࡛࠶ࡿ

F\WRFKURPH3ȘK\GUR[\ODVH O\DVH ࡀᏑᅾࡍࡿ㸦$OEUHFKWDQG'DHOV㸧ࡓࡵࠊQJPO ࡜࠸࠺㧗

⃰ᗘ࡛⾑୰࡟Ꮡᅾࡍࡿࣉࣟࢪ࢙ࢫࢸࣟࣥࡀࢸࢫࢺࢫࢸࣟࣥ࡟ኚ᥮ࡉࢀࡓࡓࡵ࡜

⪃࠼ࡽࢀࡿࠋࡋ࠿ࡋࠊ㐣ཤࡢ◊✲࡛ࡣࠊࣉࣟࢪ࢙ࢫࢸࣟࣥࡀపୗࡋࠊࢸࢫࢺࢫ

ࢸࣟࣥࡀୖ᪼ࡍࡿ࡜࠸࠺ሗ࿌㸦6LOEHU]DKQHWDO㸧ࡸⓎ᝟࿘ᮇ୰ࡣኚື

ࢆ♧ࡉ࡞࠸࡜࠸࠺ሗ࿌㸦0LQHFNHHWDO㸧ࡶᏑᅾࡍࡿࡀࠊࡇࢀࡣ ᐃ᪉ἲ

࡞࡝ࡢ㐪࠸࡛࠶ࡿ࡜᥎ᐹࡉࢀࡿࠋ

ࡲࡓࠊዷፎᮇ࡟ࡶࢸࢫࢺࢫࢸࣟࣥ࡜ࣉࣟࢪ࢙ࢫࢸࣟࣥࡣྠᵝࡢኚ໬ࢆ♧ࡋࡓࠋ 㯤య⣽⬊࡜ྠᵝ࡟ࠊ⫾┙࡟ࡶ

F\WRFKURPH3ȘK\GUR[\ODVHO\DVH

61

ࡀᏑᅾࡍࡿ㸦$OEUHFKWDQG'DHOV㸧ࡓࡵࠊ㯤యᮇ࡜ྠᵝࡢ࣓࢝ࢽࢬ࣒࡛

ศἪࣃࢱ࣮ࣥࡀ୍⮴ࡋࡓ࡜᥎ᐹࡉࢀࡿࠋ

࢚ࢫࢺࣛࢪ࣮࢜ࣝ࡜࢖ࣥࣄࣅࣥࡣ୍⮴ࡋࡓኚ໬ࢆ♧ࡋࡓࠋ࢚ࢫࢺࣛࢪ࣮࢜ࣝ

ࡣ༸⬊ࡢ㢛⢏⭷⣽⬊࠿ࡽ୺࡟ศἪࡉࢀࡿࡀࠊ࢖ࣥࣄࣅࣥࡶྠᵝ࡟༸⬊ࡢ㢛⢏⭷

⣽⬊࠿ࡽ୺࡟ศἪࡉࢀࡿ㸦1DJDPLQHHWDO㸧ࡓࡵࠊᮏ◊✲࡛ࡶࠊศἪࣃ

ࢱ࣮ࣥࡀ୍⮴ࡋࡓ࡜ゎ㔘ࡉࢀࡓࠋ

ᮏ◊✲ࡢ⤖ᯝ࠿ࡽࠊ໭ᾏ㐨࡟࠾࠸࡚㣫⫱ࡉࢀ࡚࠸ࡿ㞤ࢧࣛࣈࣞࢵࢻ࡟࠾࠸࡚ࡶ

どᗋୗ㒊࣭ୗᆶయ࣭༸ᕢ㍈ࡣࠊ㞝ࢧࣛࣈࣞࢵࢻࡢどᗋୗ㒊࣭ୗᆶయ࣭⢭ᕢ㍈࡜

㢮ఝࡋࡓ᫂ࡽ࠿࡞㛗᪥Ꮨ⠇⦾Ṫື≀ࡢ≉ᚩࢆ♧ࡍ஦ᐇࡀ☜ㄆࡉࢀࡓࠋࡇࡢ㞤ࢧ

ࣛࣈࣞࢵࢻ࡟࠾ࡅࡿどᗋୗ㒊࣭ୗᆶయ࣭༸ᕢ㍈ࡢᏘ⠇ኚ໬ࡣࠊዷፎࡸἪங࡟࠾

࠸࡚ࡶ㜼ᐖࡉࢀࡿࡇ࡜ࡀ࡞ࡃ⥔ᣢࡉࢀࡿࡇ࡜ࡀ᫂ࡽ࠿࡟࡞ࡗࡓࠋ

62

Fig. 5-1 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during complete reproductive cycle of Mare 4. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation

Gestation Gestation

0 5 10 15 100200 300400 500 600 800 1000 1200

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6 Lactation

Lactation

Date

/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

Gestation

Lactation

Gestation Gestation

0 5 10 15

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Lactation Lactation

Date

FS H (ng /m l)

Gestation

Lactation

Gestation Gestation

0 5 10 15 20 30 40 50 60

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/9

Lactation Lactation

Date

PRL(ng/ml)

Mare 4 (a)

(b)

(c)

63

Fig. 5-1 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș(f), and ir-inhibin (g) during complete reproductive cycle of Mare 4. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation

Gestation Gestation

0 50 100 150

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6 Lactation

Lactation

Date

Progesterone(ng/ml)

Gestation

Lactation

Gestation Gestation

0 100 200 300

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6 Lactation Lactation

Date

Testosterone(pg/ml)

0 20 40 60 80 100 200 400 600 800 1000

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6 Gestation

Lactation Lactation

Gestation Gestation

Lactation

Date

Estradiol-17ββββ(pg/ml)

Gestation

Lactation

Gestation Gestation

0.0 0.5 1.0 1.5 2.0 2.5 2.5 3.0 3.5 4.0 4.5 5.0

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Lactation Lactation

Date

ir-inhibin(ng/ml)

Mare 4

(d)

(e)

(f)

(g)

64

Fig. 5-2 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during complete reproductive cycle of Mare 13. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation Lactation

Gestation Gestation

0 2 4 6 8 10 100 200 300 400 500 1000 2000 3000 4000

2004/3 2004/6

2004/9 200

4/12 2005/3

2005/6 2005/9

200 5/12

2006/3 2006/6

Date

/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

Gestation

Lactation Lactation

Gestation Gestation

0 20 40 60

2004/

3 2004/

6 2004

/9 2004

/12 2005/

3 2005/

6 2005/

9 2005

/12 2006/

3 2006/

6

Date

FSH(ng/ml)

Gestation

Lactation Lactation

Gestation Gestation

0 5 10 15

2004/3 200

4/6 200

4/9 2004

/12 2005

/3 200

5/6 200

5/9 2005

/12 2006

/3 2006/9

Date

PRL(ng/ml)

Mare 13 (a)

(b)

(c)

65

Fig. 5-2 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f), and ir-inhibin (g) during complete reproductive cycle of Mare 13. Yellow box show gestation period. Arrow show lactation period. of

Gestation

Lactation Lactation

Gestation Gestation

0 10 20 30 40

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Progesterone(ng/ml)

Gestation

Lactation Lactation

Gestation Gestation

0 100 200 300 400

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Testosterone(pg/ml)

0 10 20 30 40 50 100 200 300 400 500 600 800 1000 1200 1400

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6 Gestation

Lactation Lactation

Gestation Gestation

Date

Estradiol-17ββββ(pg/ml)

Gestation

Lactation Lactation

Gestation Gestation

0.0 0.5 1.0 1.5 2.0 2.5 3 4 5 6

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

ir-inhibin(ng/ml)

Mare 13

(e)

(d) (f)

(g)

66

Fig. 5-3 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during non-pregnant period of Mare 5.

0 5 10 15

2004/3

2004/6

2004/9

2004/12

Date

/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

0 5 10 15 20 25

2004/3

2004/6

2004/9

2004/12

Date

FS H( ng/ m l)

0 1 2 3 4 5

2004/3

2004/6

2004/9

2004/12

Date

PRL(ng/ml)

Mare 5 (a)

(b)

(c)

67

Fig. 5-3 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during non-pregnant period of Mare 5.

0 10 20 30 40

2004/3

2004/6

2004/9

2004/12

Date

Progesterone(ng/ml)

0 50 100 150

2004/3

2004/6

2004/9

2004/12

Date

Testosterone(pg/ml)

0 10 20 30

2004/3

2004/6

2004/9

2004/12

Date Estradiol-17ββββ(pg/ml)

0 1 2 3 4

2004/3

2004/6

2004/9

2004/12

Date

ir-inhibin(ng/ml)

Mare 5

(d)

(e)

(f)

(g)

68

Fig. 5-4. Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during non-pregnant period of Mare 8. Arrow show lactation period.

Lactation

0 2 4 6 8 10

2004/6

2004/9

2004/12

2005/3

2005/6

2005/9

Date /+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

Lactation

0 5 10 15 20

2004/6

2004/9

2004/12

2005/3

2005/6

2005/9

Date

FS H (n g /m l)

Lactation

0 5 10 15 20 25

2004/6

2004/9

2004/12

2005/3

2005/6 2005/9

Date

PRL(ng/ml)

Mare 8 (a)

(b)

(c)

69

Fig. 5-4 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during non-pregnant period of Mare 8.

Arrow show lactation period.

Lactation

0 5 10 15 20 25

2004/

6 2004/

9 200

4/12 2005/

3 2005/

6 2005/

9

Date

Progesterone(ng/ml)

Lactation

0 50 100 150 200

2004/6 2004/9

2004/12 2005/3

2005/6 2005/9

Date

Testosterone(pg/ml)

0 10 20 30 40

2004/6 2004/9

2004/12 2005/3

2005/6 2005/9 Lactation

Date Estradiol-17ββββ(pg/ml)

Lactation

0 1 2 3 4 5

2004/

6 2004/

9 2004/

12 2005/

3 2005/

6 200

5/9

Date

ir-inhibin(ng/ml)

Mare 8

(d)

(e)

(f)

(g)

70

Fig. 5-5 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during non-pregnant period of Mare 12.

0 5 10 15 20

2004/6

2004/9

2004/12

Date

/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

0 5 10 15 20

2004/6

2004/9

2004/12

Date

FS H (n g /m l)

0 2 4 6 8 10 10 12 14 16 18 20

2004/6

2004/9

2004/12

Date

PRL(ng/ml)

Mare 12 (a)

(b)

(c)

71

Fig. 5-5 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during non-pregnant period of Mare 12.

0 10 20 30 40

2004/6

2004/9

2004/12

Date

Progesterone(ng/ml)

0 50 100 150

2004/6

2004/9

2004/12

Date

Testosterone(pg/ml)

0 10 20 30

2004/6

2004/9

2004/12

Date

Estradiol-17ββββ(pg/ml)

0 1 2 3 4 5 5 10 15 20

2004/6

2004/9

2004/12

Date

ir-inhibin(ng/ml)

Mare 12

(e)

(f)

(g)

(d)

72

Fig. 5-6 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during non-pregnant period of Mare 16.

0 5 10 15

2004/3

2004/6

2004/9

2004/12

Date

/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

0 5 10 15

2004/3

2004/6

2004/9

2004/12

Date

FS H (ng /m l)

0 5 10 15

2004/3

2004/6

2004/9

2004/12

Date

PRL(ng/ml)

Mare 16 (a)

(b)

(c)

73

Fig. 5-6 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during non-pregnant period of Mare 16.

0 10 20 30

2004/3

2004/6

2004/9

2004/12

Date

Progesterone(ng/ml)

0 50 100 150

2004/3

2004/6

2004/9

2004/12

Date

Testosterone(pg/ml)

0 20 40 60 80

2004/3

2004/6

2004/9

2004/12

Date Estradiol-17ββββ(pg/ml)

0 1 2 3 4 5

2004/3

2004/6

2004/9

2004/12

Date

ir-inhibin(ng/ml)

Mare 16 (d)

(e)

(f)

(g)

74

Fig. 5-7 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during lactation of Mare 3. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation

0 2 4 6 8 10 100 200 300400 500 500 1000 1500 2000

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date /+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

Gestation

Lactation

0 5 10 15 20 25

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

FS H (ng /m l)

Gestation

Lactation

0 5 10 15 15 20 25 30 35 40

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/9

Date

PRL(ng/ml)

Mare 3 (a)

(b)

(c)

75

Fig. 5-7 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during lactation of Mare 3. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation

0 5 10 15 20 25

2004/

3 2004/6

2004/9 200

4/12 2005/3

2005/

6 2005/9

2005/12 2006/

3 2006/

6

Date

Progesterone(ng/ml)

Gestation

Lactation

0 50 100 150 200 200 300 400

2004 /3

2004/

6 2004/

9 2004/

12 2005/

3 2005/

6 2005

/9 2005/

12 2006/

3 2006/

6

Date

Testosterone(pg/ml)

0 20 40 60 80 100 200 400 600 800 1000

200 4/3

200 4/6

200 4/9

2004/12 200

5/3 200

5/6 200

5/9 2005

/12 200

6/3 200

6/6 Gestation

Lactation

Date

Estradiol-17ββββ(pg/ml)

Gestation

Lactation

0.0 0.5 1.0 1.5 2.02 3 4 5

2004/

3 2004/

6 2004/

9 200

4/12 2005/

3 2005/6

2005/9 2005

/12 2006/3

2006/6

Date

ir-inhibin(ng/ml)

Mare 3

(d)

(e)

(f)

(g)

76

Fig. 5-8 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during lactation of Mare 9. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation

Gestation

0 5 10 15 100 200 300 400 500 1000 2000 3000 4000

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date /+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

Gestation

Lactation

Gestation

0 5 10 15 20 25 30 40 50

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

FSH(ng/ml)

Gestation

Lactation

Gestation

0 2 4 6 8 10

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/9

Date

PRL(ng/ml)

Mare 9 (a)

(b)

(c)

77

Fig. 5-8 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during lactation of Mare 9. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation

Gestation

0 10 20 30

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Progesterone(ng/ml)

Gestation

Lactation

Gestation

0 100 200 300

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Testosterone(pg/ml)

Gestation

Lactation

0 10 20 30 40 5050 100 150 200 200 400 600 800

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6 Gestation

Date Estradiol-17ββββ(pg/ml)

Gestation

Lactation

Gestation

0 1 2 3 4

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

ir-inhibin(ng/ml)

Mare 9

(d)

(e)

(f)

(g)

78

Fig. 5-9 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during lactation of Mare 11. Yellow box show gestation period. Arrow show lactation period.

Gestation Gestation

Lactation Lactation Lactation

0 2 4 6 8 10 100 200 300 400 500 1000 2000 3000 4000

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

Gestation Gestation

Lactation Lactation Lactation

0 2 4 6 8 10 10 15 20

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

FS H( ng/ m l)

Gestation Gestation

Lactation Lactation Lactation

0 2 4 6 8 10 10 15 20 25

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/9

Date

PRL(ng/ml)

Mare 11 (a)

(b)

(c)

79

Fig. 5-9 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during lactation of Mare 11. Yellow box show gestation period. Arrow show lactation period.

Gestation Gestation

Lactation Lactation Lactation

0 5 10 15 20 25

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Progesterone(ng/ml)

Gestation Gestation

Lactation Lactation Lactation

0 50 100 150 200

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Testosterone(pg/ml)

0 100 200 300

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Gestation Gestation

Lactation Lactation Lactation

Date Estradiol-17ββββ(pg/ml)

Gestation Gestation

Lactation Lactation Lactation

0.0 0.5 1.0 1.5 2.0 2.5 3 4 5 6

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

ir-inhibin(ng/ml)

Mare 11

(d)

(e)

(f)

(g)

80

Fig. 5-10 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during lactation of Mare 12. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation

0 5 10 15 100 200 300 400 500 500 1000 1500 2000 2500

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date /+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

Gestation

Lactation

0 5 10 15 20

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

FS H (ng /m l)

Gestation

Lactation

0 2 4 6 8 10 10 12 14 16 18 20

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/9

Date

PRL(ng/ml)

Mare 12 (a)

(b)

(c)

81

Fig. 5-10 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during lactation of Mare 12. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation

0 10 20 30 40

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Progesterone(ng/ml)

Gestation

Lactation

0 100 200 300 400

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Testosterone(pg/ml)

0 10 20 30 40 50 100 200 300 400

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6 Gestation

Lactation

Date

Estradiol-17ββββ(pg/ml)

Gestation

Lactation

0 1 2 3 4 55 10 15 20

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

ir-inhibin(ng/ml)

Mare 12

(d)

(e)

(f)

(g)

82

Fig. 5-11 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during lactation of Mare 17. Yellow box show gestation period. Arrow show lactation period.

Gestation Gestation

Lactation Lactation Lactation

0 2 4 6 8 10 100 200 300 400 500 600 800 1000 1200 1400

200 4/3

2004/6 200

4/9 2004/

12 200

5/3 200

5/6 2005/9

200 5/12

200 6/3

2006/6

Date

/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

Gestation Gestation

Lactation Lactation

Lactation

0 5 10 15 20

2004/3 2004/6

2004/9 2004/

12 2005/3

2005/6 2005/9

2005/

12 2006/3

2006/6

Date

F S H (ng/m l)

Gestation Gestation

Lactation Lactation

Lactation

0 5 10 15 20 20 40 60 80 100

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 200

5/12 2006/3

2006/9

Date

PRL(ng/ml)

Mare 17 (a)

(b)

(c)

83

Fig. 5-11 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during two years of Mare 17. Yellow box show gestation period. Arrow show lactation period.

Gestation Gestation

Lactation Lactation Lactation

0 5 10 15 20 25

200 4/3

2004 /6

2004 /9

2004/

12 200

5/3 200

5/6 200

5/9 2005/

12 2006

/3 2006

/6

Date

Progesterone(ng/ml)

Gestation Gestation

Lactation Lactation

Lactation

0 100 200 300 400

2004 /3

2004 /6

200 4/9

2004/12 2005

/3 2005

/6 2005

/9 2005/

12 2006

/3 2006

/6

Date

Testosterone(pg/ml)

0 500 1000 1500 2000 2500

2004/3 2004/6

2004/9 200

4/12 2005/3

2005/6 2005/9

200 5/12

2006/3 2006/6

Gestation Gestation

Lactation Lactation

Lactation

Date Estradiol-17ββββ(pg/ml)

Gestation Gestation

Lactation Lactation

Lactation

0 1 2 3 4 5

2004/3 2004

/6 2004

/9 2004/12

2005 /3

200 5/6

2005 /9

2005 /12

2006 /3

200 6/6

Date

ir-inhibin(ng/ml)

Mare 17

(d)

(e)

(f)

(g)

84

Fig. 5-12 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during two years of Mare 10. Yellow box show gestation period. Arrow show lactation period.

Gestation

Gestation Abortion Gestation

Lactation Lactation

0 5 10 15 200 400 600

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date /+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

Gestation

Gestation Abortion Gestation

Lactation

Lactation

0 5 10 15 20 20 30 40

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

FS H (n g /m l)

Gestation

Gestation Abortion Gestation

Lactation Lactation

0 5 10 15 20 30 40 50 50 100 150

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/9

Date

PRL(ng/ml)

Mare 10 (a)

(b)

(c)

85

Fig. 5-12 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during two years of Mare 10. Yellow box show gestation period. Arrow show lactation period.

Gestation

Gestation Abortion Gestation

Lactation Lactation

0 5 10 15 20 25

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Progesterone(ng/ml)

Gestation

Gestation Abortion Gestation

Lactation Lactation

0 50 100 150 200

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Testosterone(pg/ml)

0 200 400 600

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6 Gestation

Gestation Abortion Gestation

Lactation Lactation

Date

Estradiol-17ββββ(pg/ml)

Gestation

Gestation Abortion Gestation

Lactation

Lactation

0 1 2 3 4 6 8 10

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

ir-inhibin(ng/ml)

Mare 10

(d)

(e)

(f)

(g)

86

Fig. 5-13 Changes in plasma concentration of PRL (a), LH (b) and FSH (c) during two years of Mare 14. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation

Abortion Gestation

0 5 10 15

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date /+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

Gestation

Lactation

Abortion Gestation

0 5 10 15 20

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

FSH(ng/ml)

Gestation

Lactation

Abortion Gestation

0 5 10 15 20

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/9

Date

PRL(ng/ml)

Mare 14 (a)

(b)

(c)

87

Fig. 5-13 Changes in plasma concentration of progesterone (d), testosterone (e), estradiol-17ș (f) and ir-inhibin (g) during two years of Mare 14. Yellow box show gestation period. Arrow show lactation period.

Gestation

Lactation

Abortion Gestation

0 5 10 15 20 25

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

Progesterone(ng/ml)

Gestation

Lactation Gestation

0 50 100 150 200 1000500 15002000 2500

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6 Abortion

Date

Testosterone(pg/ml)

Gestation

Lactation

Abortion

0 20 40 60 80 100 200 400 600 800

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6 Gestation

Date Estradiol-17ββββ(pg/ml)

Gestation

Lactation

Gestation Abortion

0 1 2 3 4

2004/3 2004/6

2004/9 2004/12

2005/3 2005/6

2005/9 2005/12

2006/3 2006/6

Date

ir-inhibin(ng/ml)

Mare 14

(d)

(e)

(f)

(g)

88

0 5 10 15 20

0 50 100 150 200

2005/6 2005/9 2005/12 2006/3

7HVWRVWHURQHᤡSJPOᤢ7HVWRVWHURQHᤡSJPOᤢ7HVWRVWHURQHᤡSJPOᤢ7HVWRVWHURQHᤡSJPOᤢ

Progesterone Testosterone

A

Progesterone(ng/ml)

0 10 20 30

0 100 200 300

2004/3 2004/6 2004/9 2004/12 2005/3 2005/6 2005/9 2005/12 2006/3 2006/6

Date

Testosterone(pg/ml)

Progesterone Testosterone

Gestation Gestation

B

Progesterone(ng/ml)

Fig.5-14 Comparing changes in plasma concentration of testosterone (ڦ) with progesterone (ە) during estrus cycle (A) and pregnancy (B).

89 0

5 10 15

0 10 20 30 40 50

2005/6 2005/9 2005/12 2006/3 2006/6

E s tr ad io l-17 ββββ (pg/m l)

Progesterone Estradiol-17β

P ro g ester o n e( n g /m l)

Fig.5-15 Comparing changes in plasma concentration of estradiol-17ș (ڸ) with progesterone (ە) during estrus cycle.

90 0

5 10 15

0.0 0.5 1.0 1.5 2.0 2.5 5

2005/6 2005/9 2005/12 2006/3 2006/6

ir -inhi bi n( ng /m l)

P roge s te rone (ng/ m l)

Date

Fig.5-16 Comparing changes in plasma concentration of ir-inhibin (ی) with progesterone (ە) during estrus cycle.

91 0

1 2 3

0 10 20 30 40 50

2005/6 2005/9 2005/12 2006/3

Date

Estradiol-17ββββ(pg/ml) ir-inhibin Estradiol-17β

ir -i n h ib in (n g /m l)

Fig.5-17 Comparing changes in plasma concentration of ir-inhibin (ی) with estradiol-17ș (ڸ) during estrus cycle.

92

㸴❶㸴❶

㸴❶㸴❶ ග⥺ฎ⌮ග⥺ฎ⌮ග⥺ฎ⌮ග⥺ฎ⌮ἲࢆᐇ᪋ࡋࡓἲࢆᐇ᪋ࡋࡓἲࢆᐇ᪋ࡋࡓᙜἲࢆᐇ᪋ࡋࡓᙜᙜᙜṓṓṓṓ㞤㞤㞤㞤㤿㤿㤿㤿ࡢ⏕Ṫࡢ⏕Ṫࡢ⏕Ṫࡢ⏕ṪෆศἪᏛⓗኚ໬ෆศἪᏛⓗኚ໬ෆศἪᏛⓗኚ໬ෆศἪᏛⓗኚ໬

㸴㸫㸯㸬ᐇ㦂ࡢ⫼ᬒ 㸴㸫㸯㸬ᐇ㦂ࡢ⫼ᬒ 㸴㸫㸯㸬ᐇ㦂ࡢ⫼ᬒ 㸴㸫㸯㸬ᐇ㦂ࡢ⫼ᬒ

ᛶᡂ⇍࡜ࡣࠊ㞝࡛ࡣ㞤ࢆዷፎࡉࡏࡿ≧ែࠊ㞤࡛ࡣ㞝࡜஺㓄ࡍࡿࡇ࡜࡛ዷፎࡋ

࠺ࡿ≧ែ࡟ࡲ࡛ᡂ㛗ࡋࡓ⏕Ṫ⏕⌮Ꮫ≧ែࢆゝ࠺ࠋ᫓ᶵⓎືࡣ⏕ṪჾࡢⓎ⫱ࡀ㛤 ጞࡉࢀࠊ⏕Ṫᶵ⬟ࡢ୍㒊ࡀⓎ⌧ࡍࡿ≧ែ࡜ᐃ⩏ࡉࢀ࡚࠸ࡿࠋ㞝࡛ࡣࠊ⢭ᕢࡢᛴ

⃭࡞Ⓨ⫱ࡸ⢭Ꮚᙧᡂ⬟ࡢ㛤ጞࠊ㞤࡛ࡣ༸ᕢ࡛༸⬊ࡀᛴ⃭࡟Ⓨ⫱ࡋ࡚ึ᤼༸ࡀㄏ

㉳ࡉࢀࡿ᫬ᮇࢆព࿡ࡍࡿࠋ᫓ᶵⓎືࡢ᫬ᮇࡣẼೃࠊᏘ⠇ࠊᰤ㣴≧ែ࡞࡝ᵝࠎ࡞

せᅉ࡟ࡼࡾኚ໬ࡋࠊရ✀ࡸ㑇ఏⓗせᅉ࡜࠸ࡗࡓಶయᕪࡶ⏕ࡌࡸࡍ࠸ࠋ≉࡟࣐࢘

ࡣࠊ඾ᆺⓗ࡞㛗᪥Ꮨ⠇⦾Ṫື≀࡛࠶ࡾᏘ⠇࡟ࡼࡿ᪥↷ࡢ㛗ࡉ࡟ᙳ㡪ࢆཷࡅࡿࠋ 㞤࣐࢘ࡢ᫓ᶵⓎືࡣࠊ㹼 ࢝᭶㱋࡛㉳ࡇࡾࠊᛶᡂ⇍ࡣ 㹼 ࠿᭶㱋࡛࠶

ࡿࠋ㞝࣐࢘ࡣࠊ㹼 ࠿᭶㱋࡛⢭Ꮚ⏕⏘ࡀࡣࡌࡲࡾ ࢝᭶๓ᚋ࡛ᛶᡂ⇍࡟㐩 ࡍࡿ㸦᳃ࡽ㸧ࠋ

㏆ᖺࠊࢃࡀᅜࡢ➇㉮㤿⏕⏘࡛⦾Ṫࢩ࣮ࢬࣥࡢ᪩ᮇ໬ࡀ㐍ࢇ࡛࠾ࡾࠊࠊ ᭶

⏕ࡲࢀࡢᏊ㤿ࡶぢࡽࢀࡿࡼ࠺࡟࡞ࡗ࡚ࡁࡓࠋࢃࡀᅜࡢ⮬↛⎔ቃୗ࡛ࡣࢧࣛࣈࣞ

ࢵࢻࡢᏘ⠇⦾Ṫᛶ࡟ࡼࡾࠊ㏻ᖖࠊ㞤࣐࢘ࡣ 㹼 ᭶㡭࡟᤼༸ࡀጞࡲࡿࡓࡵࠊࠊ ᭶⏕ࡲࢀࡢᏊ㤿ࢆ⏘ࡴࡇ࡜ࡣ㞴ࡋ࠸ࠋࡑࡇ࡛ࠊ࣐࢘⏕⏘∾ሙ࡛ࡣࠊ᪥↷᫬㛫 ࡢ▷࠸ ᭶࠿ࡽ㤿ᡣෆ࡟᫨ ᫬㛫ࠊኪ ᫬㛫ࡢ↷᫂⎔ቃࢆసฟࡋࠊேⅭ

ⓗ࡟᪥↷᫬㛫ࢆᘏ㛗ࡍࡿග⥺ฎ⌮ἲࢆ⏝࠸ࡿࡇ࡜࡛ࠊ㞤࣐࢘ࡢࢩ࣮ࢬࣥึᅇ᤼

༸ࡢ᪩ᮇ໬ࢆㄏⓎࡋ࡚ ࠊ ᭶࡟᤼༸ࡍࡿฎ⨨ࡀ⏝࠸ࡽࢀ࡚࠸ࡿࠋ

ࡍ࡞ࢃࡕࠊ㛗᪥Ꮨ⠇⦾Ṫື≀࡛࠶ࡿ࣐࡛࢘ࡣࠊ᪥↷᫬㛫ࡀ༙᪥ࢆୖᅇࡿ㛗᪥

ᮇ࡟ᛶ⭢ᶵ⬟ࡀ㈿ά໬ࡉࢀࠊ஺㓄ྍ⬟࡞≧ែ࡟࡞ࡿࠋ▷᪥ᮇ࡟㤿ᡣෆࢆ↷᫂ࡍ

ࡿࡇ࡜࡟ࡼࡗ࡚ேⅭⓗ࡟㛗᪥ᮇࢆసࡾฟࡍග⥺ฎ⌮ἲࡣࠊ࣐࢘ࡢᛶ⭢ᶵ⬟ࢆ᪩

ᮇ࡟㈿ά໬ࡋࠊ᤼༸ࢆಁ㐍ࡍࡿࡇ࡜࡟฼⏝ࡉࢀ࡚࠸ࡿࠋ㸦%XUNKDUGWࠊ

93

)UHHGPDQHWDO㸧ࡲࡓࠊ⫱ᡂ୰ࡢࢧࣛࣈࣞࢵࢻࡢ࠾࠸࡚ࡶග⥺ฎ⌮ἲ࡟

ࡼࡾࠊ᫓ᶵⓎືࡢ᪩ᮇ໬ࡸẟⰏࡢኚ໬㸦.XQLLHWDOࠊ㸧ࠊ➽⫗ࡢቑᙉస⏝

㸦ᒸ㔝ࠊ㸧ࡀ☜ㄆࡉࢀ࡚࠸ࡿࠋ

ᮏ◊✲࡛ࡣࠊ㞤ࢧࣛࣈࣞࢵࢻ࡟ ṓࡢ෤࠿ࡽ ṓࡢ᫓࡟࠿ࡅ࡚ග⥺ฎ⌮ἲࢆ

ᐇ᪋ࡋࠊ⏕యࡢෆศἪⓗኚ໬ࠊᛶ⭢ࡢኚ໬ཬࡧ⿕ẟࡢኚ໬࡟ࡘ࠸᳨࡚ウࡋࡓࠋ

94

㸴㸫㸰㸬ᮦᩱ࠾ࡼࡧ᪉ἲ 㸴㸫㸰㸬ᮦᩱ࠾ࡼࡧ᪉ἲ㸴㸫㸰㸬ᮦᩱ࠾ࡼࡧ᪉ἲ 㸴㸫㸰㸬ᮦᩱ࠾ࡼࡧ᪉ἲ

㸴㸫㸰㸫㸯㸬౪ヨື≀

㸴㸫㸰㸫㸯㸬౪ヨື≀

㸴㸫㸰㸫㸯㸬౪ヨື≀

㸴㸫㸰㸫㸯㸬౪ヨື≀

໭ᾏ㐨᪥㧗ᆅ༊࡛㣫㣴ࡉࢀ࡚࠸ࡓ ᖺ⏕ࡲࢀࡢࢧࣛࣈࣞࢵࢻ✀㞤㤿 㢌

ࢆ⏝࠸ࡓࠋࡑࡢ࠺ࡕࠊ 㢌࡟ࡘ࠸࡚ࠊ ᖺ ᭶ ᪥࠿ࡽ ᭶ ᪥ࡲ࡛ࡢ⣙

ࣨ᭶㛫ග⥺ฎ⌮ἲࢆᐇ᪋ࡋࡓ㸦௨ୗࠊ(3 ⩌㸧ࠋṧࡾࡢ 㢌ࡣᑐ↷⩌࡜ࡋ࡚ࠊ⮬

↛ගࡢࡳ࡛㣫⫱ࡋࡓࠋ

㸴㸫㸰㸫㸰㸬 㸴㸫㸰㸫㸰㸬 㸴㸫㸰㸫㸰㸬

㸴㸫㸰㸫㸰㸬ග⥺ฎ⌮ග⥺ฎ⌮ග⥺ฎ⌮ἲ࡜ᐇ㦂᧯సග⥺ฎ⌮ἲ࡜ᐇ㦂᧯సἲ࡜ᐇ㦂᧯సἲ࡜ᐇ㦂᧯స

ග⥺ฎ⌮ἲࡣࠊ㤿ᡣෆኳ஭࡟

࣡ࢵࢺࡢⓑⰍ㟁⌫ࢆタ⨨ࡋࠊ᫂ᮇ ᫬㛫ࠊ ᬯᮇ ᫬㛫ࡢ↷᫂᮲௳ࢆࢱ࢖࣐࣮᧯స࡟ࡼࡾసฟࡋࡓࠋẖ㐌

ᅇ᥇⾑ࡍࡿ࡜

࡜ࡶ࡟ࠊ୧⩌ࡑࢀࡒࢀᐇ㦂㛤ጞ ࣨ᭶᫬Ⅼ࡟࠾࠸࡚ẟⰏࡢホ౯ࢆ⾜ࡗࡓࠋෆศ ἪᏛⓗኚ໬ࢆㄪ࡭ࡿࡓࡵ࡟ࠊ᥇⾑ࡋࡓ⾑ᾮࢆ⏝࠸࡚ࠊ㯤యᙧᡂ࣍ࣝࣔࣥ㸦/+㸧ࠊ

༸⬊่⃭࣍ࣝࣔࣥ㸦)6+㸧ࠊࣉࣟࣛࢡࢳࣥࠊ࢖ࣥࣄࣅࣥࠊࢸࢫࢺࢫࢸࣟࣥࠊ࢚ࢫ

ࢺࣛࢪ࣮࢜ࣝࢆࣛࢪ࢜࢖࣒ࣀ࢔ࢵࢭ࢖㸦5,$㸧ἲ࡟࡚ࠊࣉࣟࢪ࢙ࢫࢸࣟࣥࢆ᫬㛫

ศゎ⺯ගච␿ ᐃἲ㸦75),$㸧࡟࡚ ᐃࡋࡓࠋ

㸴㸫㸰㸫㸱㸬⾑ᾮࡢ᥇ྲྀ࠾ࡼࡧฎ⌮

㸴㸫㸰㸫㸱㸬⾑ᾮࡢ᥇ྲྀ࠾ࡼࡧฎ⌮

㸴㸫㸰㸫㸱㸬⾑ᾮࡢ᥇ྲྀ࠾ࡼࡧฎ⌮

㸴㸫㸰㸫㸱㸬⾑ᾮࡢ᥇ྲྀ࠾ࡼࡧฎ⌮

᥇⾑ࡣࠊᐇ㦂ᮇ㛫୰ࠊ㹼 ᭶ࡲ࡛ࡣ᭶ ᅇࠊࡑࢀ௨㝆ࡣẖ㐌ࠊ㢁㟼⬦࠿ࡽ࡬

ࣃࣜࣥฎ⌮┿✵᥇⾑⟶㸦ࢸࣝࣔࠊᮾி㸧ࢆ⏝࠸࡚⾜ࡗࡓࠋ᥇⾑ᚋࡣࠊịୖ࡛ಖ Ꮡࡋࠊ࡛ࡁࡿࡔࡅ㏿ࡸ࠿࡟ Υࠊ*

࡛ ศ㛫㐲ᚰࡋ࡚⾑₢ࢆศ㞳ࡋࡓࠋᚓ

ࡽࢀࡓ⾑₢ࡣࠊྛ✀࣍ࣝࣔࣥ ᐃࡲ࡛㸫Υ࡟࡚෾⤖ಖᏑࡋࡓࠋ

㸴㸫㸰㸫㸲㸬⿕ẟ 㸴㸫㸰㸫㸲㸬⿕ẟ 㸴㸫㸰㸫㸲㸬⿕ẟ

㸴㸫㸰㸫㸲㸬⿕ẟࡢホ౯ࡢホ౯ࡢホ౯ࡢホ౯

ᐇ㦂㛤ጞ ࣨ᭶ࡢ᫬Ⅼ࡟࠾࠸࡚ࠊ

ྡࡢホ౯⪅࡟ࡼࡾ (3 ⩌

㢌ࠊᑐ↷⩌ 㢌

࡟ࡘ࠸࡚┣᳨ⓗ࡟ホ౯ࢆ⾜࠸ࠊࡑࡢホ౯Ⅼࢆẚ㍑ࡋࡓࠋホ౯Ⅼࡣࠊ

ࠕⰋ࠸ࠖࢆ

95

Ⅼࠊ ࠕᬑ㏻ࠖࢆ Ⅼࠊ ࠕᝏ࠸ࠖࢆ Ⅼࡢ ẁ㝵ホ౯࡟ࡼࡾ᥇Ⅼࡋࡓࠋ

㸴㸫㸰㸫㸴㸫㸰㸫

㸴㸫㸰㸫㸴㸫㸰㸫㸳㸳㸳㸳㸬㞤࣐࢘ࡢึᅇ᤼༸㸬㞤࣐࢘ࡢึᅇ᤼༸㸬㞤࣐࢘ࡢึᅇ᤼༸㸬㞤࣐࢘ࡢึᅇ᤼༸

ึᅇ᤼༸᫬ᮇࢆ⾑୰ࣉࣟࢪ࢙ࢫࢸࣟࣥ⃰ᗘࡀ QJPO ௨ୖࢆ♧ࡋࡓ᪥ࡢ

๓࡜タᐃࡋࡓࠋ

㸴㸫㸰㸫㸴㸫㸰㸫

㸴㸫㸰㸫㸴㸫㸰㸫㸴㸴㸴㸬⤫ィฎ⌮㸴㸬⤫ィฎ⌮㸬⤫ィฎ⌮㸬⤫ィฎ⌮

ࡍ࡭࡚ࡢ⤖ᯝࡣࠊ

ᖹᆒsᶆ‽ㄗᕪ࡛♧ࡋࡓࠋ ୧⩌㛫ࡢ࣍ࣝࣔࣥ⃰ᗘࡢẚ㍑ࢆ

ඖ㓄⨨ࡢศᩓศᯒཬࡧ 3RVW+RF ࢸࢫࢺࠊẟⰏࡢホ౯ࡢẚ㍑ࡣ 0DQQ:KLWQH\

8

᳨ᐃࢆ⏝࠸࡚ࠊ᭷ពᕪࢆุᐃࡋࡓࠋ

඲࡚ࡢ⤫ィฎ⌮࡟㛵ࡋ࡚ࠊ᭷ពỈ‽

ୗ࡛᭷ព࡜ุᐃࡋࡓࠋ

96

㸴㸫㸱㸬⤖ᯝ 㸴㸫㸱㸬⤖ᯝ 㸴㸫㸱㸬⤖ᯝ 㸴㸫㸱㸬⤖ᯝ

㸴㸫㸱㸫㸯㸬⾑୰

㸴㸫㸱㸫㸯㸬⾑୰

㸴㸫㸱㸫㸯㸬⾑୰

㸴㸫㸱㸫㸯㸬⾑୰ྛ✀ྛ✀ྛ✀࣍ࣝࣔࣥ⃰ᗘྛ✀࣍ࣝࣔࣥ⃰ᗘ࣍ࣝࣔࣥ⃰ᗘࡢ࣍ࣝࣔࣥ⃰ᗘࡢࡢࡢኚ໬ኚ໬ኚ໬ኚ໬

㸴㸫㸱㸫㸯㸫㸯㸬ࣉࣟࣛࢡࢳࣥ㸦 㸴㸫㸱㸫㸯㸫㸯㸬ࣉࣟࣛࢡࢳࣥ㸦 㸴㸫㸱㸫㸯㸫㸯㸬ࣉࣟࣛࢡࢳࣥ㸦

㸴㸫㸱㸫㸯㸫㸯㸬ࣉࣟࣛࢡࢳࣥ㸦)LJ)LJ)LJ)LJDDD㸧D㸧㸧㸧

ᑐ↷⩌࡛ࡣࠊ ᭶࡟ୖ᪼ࡋࡓᚋ ᭶ࡲ࡛ప್ࢆ⥔ᣢࡋࡓᚋ࡟ࠊୖ᪼ഴྥࢆ♧ࡋ

ࡓࠋග⥺ฎ⌮⩌࡛ࡣࠊ ᭶ࠊ ᭶࡜ᛴ⃭࡟ୖ᪼ࡋࠊࡑࡢᚋࡣࠊୖ᪼࡜పୗࢆ⧞ࡾ

㏉ࡋࡓࠋ ᭶௨㝆ࡣࠊග⥺ฎ⌮⩌ࡀᑐ↷⩌࡜ẚ㍑ࡋ࡚ࠊ㧗࠸ഴྥࢆ♧ࡋࠊ ᭶࡟

᭷ព࡟㧗࠸್ࢆ♧ࡋࡓࠋ

㸴㸫㸱㸫㸯㸫㸰㸬 㸴㸫㸱㸫㸯㸫㸰㸬 㸴㸫㸱㸫㸯㸫㸰㸬

㸴㸫㸱㸫㸯㸫㸰㸬㯤యᙧᡂ࣍ࣝࣔࣥ㸦㯤యᙧᡂ࣍ࣝࣔࣥ㸦㯤యᙧᡂ࣍ࣝࣔࣥ㸦/+㯤యᙧᡂ࣍ࣝࣔࣥ㸦/+/+/+㸧㸦㸧㸦㸧㸦)LJ㸧㸦)LJ)LJE)LJ EEEࠊࠊࠊࠊ)LJ)LJ)LJ)LJDFDFDFDF

᭶࠿ࡽ ᭶ࡲ࡛ࡢᑐ↷⩌࡜ග⥺ฎ⌮⩌ࡢ⾑୰

/+ ⃰ᗘࡢᖹᆒ್ࡢኚ໬ࢆࠊ)LJ E

࡟♧ࡍࠋࡲࡓࠊ

୧⩌ࡢ ᭶࠿ࡽ ᭶ࡲ࡛ࡢಶయࡈ࡜ࡢ⾑୰ /+ ⃰ᗘࢆ )LJ

DF ࡟♧ࡍࠋᑐ↷⩌࡛ࡣࠊ ᭶࠿ࡽ⦆ࡸ࠿࡞ୖ᪼ࢆ♧ࡋࡓࡀࠊග⥺ฎ⌮⩌

࡛ࡣᛴ⃭࡟ୖ᪼ࡋࠊ ᭶ࡲ࡛ග⥺ฎ⌮⩌ࡀᑐ↷⩌࡜ẚ㍑ࡋ࡚ࠊ㧗࠸್ࢆ♧ࡋࡓࠋ

୧⩌࡜ࡶಶయᕪࡀ኱ࡁࡃ᭷ពᕪࡣㄆࡵࡽࢀ࡞࠿ࡗࡓ㸦)LJE㸧ࠋ୧⩌࡜ࡶ⾑

/+ ⃰ᗘ࡟ಶయ㛫ࡢࣂࣛࢶ࢟ࡀ኱ࡁ࠿ࡗࡓࡇ࡜࠿ࡽࠊ ᭶࠿ࡽ ᭶ࡲ࡛ࡢ⾑୰

/+ ⃰ᗘࢆಶయࡈ࡜࡟ࣉࣟࢵࢺࡋࡓ⤖ᯝࠊ ᭶ࡲ࡛ࡣࠊ୧⩌࡜ࡶ࡟ QJPO ࢆ㉸

࠼ࡿ⾑୰ /+ ⃰ᗘࢆ♧ࡍಶయᩘࡣࠊྠᩘ࡛࠶ࡗࡓࡀࠊ ᭶௨㝆ࡣࠊග⥺ฎ⌮⩌ࡀ ከ࠸ഴྥࢆ♧ࡋࡓࠋࡲࡓࠊ ᭶௨㝆ࡣࠊග⥺ฎ⌮⩌࡛ࡣࠊQJPO

௨ୖࢆ♧ࡍಶయᩘࡶᑐ↷⩌࡟ẚ࡭࡚ከࡃࠊග⥺ฎ⌮⩌ࡀᑐ↷⩌࡟ẚ࡭࡚ᖹᆒ⾑

/+ ⃰ᗘࡀ㧗࠸್ࢆ♧ࡋࡓ㸦)LJDF㸧ࠋ

㸴㸫㸱㸫㸯㸫㸱㸬 㸴㸫㸱㸫㸯㸫㸱㸬 㸴㸫㸱㸫㸯㸫㸱㸬

㸴㸫㸱㸫㸯㸫㸱㸬༸⬊่⃭࣍ࣝࣔࣥ㸦༸⬊่⃭࣍ࣝࣔࣥ㸦༸⬊่⃭࣍ࣝࣔࣥ㸦)6+༸⬊่⃭࣍ࣝࣔࣥ㸦)6+)6+)6+㸧㸦㸧㸦㸧㸦)LJ㸧㸦)LJ)LJF)LJ FFF㸧㸧㸧㸧

୧⩌ඹ࡟ࠊ ᭶࠿ࡽ ᭶ࡲ࡛ࠊ ࡯ࡰ୍ᐃࡢᇶᗏ್ࢆ⥔ᣢࡋࠊ ୧⩌㛫࡟ᕪࡣㄆࡵ

97

ࡽࢀ࡞࠿ࡗࡓࠋ

㸴㸫㸱㸫㸯㸫㸲㸬࢖ࣥࣄࣅࣥ㸦 㸴㸫㸱㸫㸯㸫㸲㸬࢖ࣥࣄࣅࣥ㸦 㸴㸫㸱㸫㸯㸫㸲㸬࢖ࣥࣄࣅࣥ㸦

㸴㸫㸱㸫㸯㸫㸲㸬࢖ࣥࣄࣅࣥ㸦)LJ)LJ)LJ)LJGGG㸧G㸧㸧㸧

ᑐ↷⩌࡛ࡣࠊ ᭶࠿ࡽ ᭶ࡲ࡛⦆ࡸ࠿࡞ୖ᪼ഴྥࢆ♧ࡋࡓࡢ࡟ᑐࡋ࡚ࠊග⥺

ฎ⌮⩌࡛ࡣࠊ ᭶࠿ࡽ ᭶ࡲ࡛ࠊ࡯ࡰ୍ᐃࡢ್ࢆ♧ࡋࡓࠋ୧⩌㛫࡟ᕪࡣㄆࡵࡽ

ࢀ࡞࠿ࡗࡓࠋ

㸴㸫㸱㸫㸯㸫㸳㸬ࣉࣟࢪ࢙ࢫࢸࣟࣥ㸦 㸴㸫㸱㸫㸯㸫㸳㸬ࣉࣟࢪ࢙ࢫࢸࣟࣥ㸦 㸴㸫㸱㸫㸯㸫㸳㸬ࣉࣟࢪ࢙ࢫࢸࣟࣥ㸦

㸴㸫㸱㸫㸯㸫㸳㸬ࣉࣟࢪ࢙ࢫࢸࣟࣥ㸦)LJ)LJ)LJ)LJHHHHࠊࠊࠊࠊ)LJ)LJ)LJ)LJEGEGEG㸧EG㸧㸧㸧

᭶࠿ࡽ ᭶ࡲ࡛ࡢᑐ↷⩌࡜ග⥺ฎ⌮⩌ࡢ⾑୰ࣉࣟࢪ࢙ࢫࢸࣟࣥ⃰ᗘࡢᖹᆒ

್ࡢኚ໬ࢆࠊ)LJH ࡟♧ࡋࡓࠋࡲࡓࠊ୧⩌ࡢ ᭶࠿ࡽ ᭶ࡲ࡛ࡢಶయࡈ

࡜ࡢ⾑୰ࣉࣟࢪ࢙ࢫࢸࣟࣥ⃰ᗘࢆ )LJEG ࡟♧ࡋࡓࠋ⾑୰ࣉࣟࢪ࢙ࢫࢸ

ࣟࣥ⃰ᗘࡣࠊᑐ↷⩌࡛ࡣࠊ ᭶ࡲ࡛ప್ࢆ♧ࡋࠊ ᭶࠿ࡽ ᭶࡟᫂ࡽ࠿࡞ୖ᪼ࢆ

♧ࡋࡓࠋ୍᪉ࠊග⥺ฎ⌮⩌࡛ࡣࠊ ᭶࠿ࡽୖ᪼ࡋࠊ ᭶ࡲ࡛ᑐ↷⩌࡜ẚ㍑ࡋ࡚㧗

࠸್ࢆ♧ࡋࡓ㸦)LJH㸧ࠋ ᭶࠿ࡽ ᭶ࡲ࡛ࡢ⾑୰ࣉࣟࢪ࢙ࢫࢸࣟࣥ⃰ᗘ

ࢆಶయࡈ࡜࡟ࣉࣟࢵࢺࡋࡓ⤖ᯝࠊᑐ↷⩌࡛ࡣࠊ ᭶ࡲ࡛ࡣࠊప್ࢆ♧ࡋࠊ ᭶࠿

ࡽ㧗್ࢆ♧ࡍಶయࡀㄆࡵࡽࢀࡓࠋ୍᪉ࠊග⥺ฎ⌮⩌࡛ࡣࠊ ᭶࠿ࡽ㧗್ࢆ♧ࡍಶ యࡀฟ⌧ࡋࡓ㸦)LJEG㸧ࠋ

㸴㸫㸱㸫㸯㸫㸴㸬࢚ࢫࢺࣛࢪ࣮࢜ࣝ㸦 㸴㸫㸱㸫㸯㸫㸴㸬࢚ࢫࢺࣛࢪ࣮࢜ࣝ㸦 㸴㸫㸱㸫㸯㸫㸴㸬࢚ࢫࢺࣛࢪ࣮࢜ࣝ㸦

㸴㸫㸱㸫㸯㸫㸴㸬࢚ࢫࢺࣛࢪ࣮࢜ࣝ㸦)LJ)LJ)LJ)LJIIII㸧㸧㸧㸧

ᑐ↷⩌࡜ග⥺ฎ⌮⩌ඹ࡟ ᭶ᚋ༙࠿ࡽୖ᪼ࢆ♧ࡋࡓࡀࠊ୧⩌㛫࡟ᕪࡣㄆࡵࡽ

ࢀ࡞࠿ࡗࡓࠋ

㸴㸫㸴㸫

㸴㸫㸴㸫㸱㸫㸰㸬⿕ẟ㸱㸫㸰㸬⿕ẟ㸱㸫㸰㸬⿕ẟ㸱㸫㸰㸬⿕ẟࡢホ౯㸦ࡢホ౯㸦ࡢホ౯㸦)LJࡢホ౯㸦)LJ)LJ)LJ)LJ)LJ)LJ㸧)LJ 㸧㸧㸧

ᑐ↷⩌࡜ග⥺ฎ⌮⩌ࡢ௦⾲౛ࡢ෗┿ࢆࠊ)LJ ࡟ࠊホ౯⪅࡟ࡼࡿホ౯⤖ᯝ

ࢆ )LJ ࡟♧ࡋࡓࠋ

ᑐ↷⩌࡛ࡣࠊ෤ẟࡀṧࡗ࡚࠸ࡿࡢ࡟ᑐࡋ࡚ග⥺ฎ⌮⩌ࡣࠊ

98

෤ẟࡀᢤࡅ࡚ኟẟ࡟ኚࡿኚ໬ࡀほᐹࡉࢀࠊ୧⩌㛫࡟᫂ࡽ࠿࡞㐪࠸ࡀㄆࡵࡽࢀࡓࠋ

⿕ẟࡢホ౯ࡣࠊග⥺ฎ⌮⩌ࡢᖹᆒⅬࡀ

Ⅼ㸦s㸧࡛࠶ࡗࡓࡢ࡟ᑐࡋ

࡚ࠊᑐ↷⩌ࡢᖹᆒⅬࡀ

Ⅼ㸦s㸧࡛࠶ࡾࠊග⥺ฎ⌮⩌ࡀᑐ↷⩌࡜ẚ㍑

ࡋ࡚ࠊ᭷ព࡟㧗࠸್ࢆ♧ࡋࡓࠋ

㸴㸫㸱㸫㸱㸬ึᅇ᤼༸

㸴㸫㸱㸫㸱㸬ึᅇ᤼༸

㸴㸫㸱㸫㸱㸬ึᅇ᤼༸

㸴㸫㸱㸫㸱㸬ึᅇ᤼༸᫬ᮇ᫬ᮇ᫬ᮇ㸦᫬ᮇ㸦㸦㸦)LJ)LJ)LJ)LJ㸧㸧㸧㸧

᭶ ᪥࠿ࡽ ᭶ ᪥ࡲ࡛ࡢᐇ㦂ᮇ㛫୰࡟ึᅇ᤼༸ࡢ㉳ࡇࡗࡓ㢌ᩘࡣࠊ

ᑐ↷

⩌࡛ࡣࠊ 㢌୰ 㢌ࠊග⥺ฎ⌮⩌࡛ࡣࠊ 㢌୰㸴㢌࡛࠶ࡗࡓࠋᑐ↷⩌࡛ࡣࠊึ

ᅇ᤼༸ࡀ㸲᭶࡟ࡣ㸴㢌࡟ㄆࡵࡽࢀࡓࡢ࡟ᑐࡋ࡚ࠊග⥺ฎ⌮⩌࡛ࡣࠊ

㸰᭶࡟㸱㢌ࠊ 㸱᭶࡟㸰㢌ࠊ㸳᭶࡟㸯㢌࡟ㄆࡵࡽࢀࠊᑐ↷⩌࡟ẚ࡭࡚ࠊග⥺ฎ⌮⩌࡛᫂ࡽ࠿࡟

ึᅇ᤼༸᫬ᮇࡢ᪩ᮇ໬ࡀㄆࡵࡽࢀࡓࠋ

99

㸴㸫㸲㸬⪃ᐹ 㸴㸫㸲㸬⪃ᐹ㸴㸫㸲㸬⪃ᐹ 㸴㸫㸲㸬⪃ᐹ

ග⥺ฎ⌮࡟ࡼࡾࠊ

/+ࠊ ࣉࣟࣛࢡࢳࣥࡢศἪࡢ᪩ᮇஹ㐍ࡀࡳࡽࢀࡓࡀࠊ)6+

࡟ࡣ ㄆࡵࡽࢀ࡞࠿ࡗࡓࠋࡋ࠿ࡋࠊ༸ᕢ࡬ࡢᙳ㡪ࡀㄆࡵࡽࢀࠊ༸ᕢ࠿ࡽศἪࡉࢀࡿࣉ

ࣟࢪ࢙ࢫࢸࣟࣥࡢ᪩ᮇୖ᪼ࡀㄆࡵࡽࢀࠊึᅇ᤼༸ࡀ᪩ᮇ໬ࡉࢀࡓ࡜᥎ᐹࡉࢀࡓࠋ

࢚ࢫࢺࣛࢪ࣮࢜ࣝࠊ࢖ࣥࣄࣅࣥࡢ᪩ᮇୖ᪼ࡣㄆࡵࡽࢀ࡞࠿ࡗࡓࠋ

ࣉ ࣟ

ࢡ ࢳ

ࣥ ࡣ ࠊ ୗ

య ๓

࠿ ࡽ ศ Ἢ ࡉ ࢀ ࡿ

⺮ ⓑ ㉁ ࣍ ࣝ ࣔ

ࣥ ࡛ ࠶

ࡾ ࠊ 㛗 ᪥ ᮇ ࡟ ศ Ἢ

ࡀ ቑ

ࡍ ࡿ

࣍ ࣝ ࣔ

ࣥ ࡛ ࠶ ࡿ ࠋ ᡂ ⇍ 㞝 㤿 ࠊ

ཤ ໃ

㤿 㸦

'KDNDOHWDO㸧 ཬ ࡧ ฟ ⏕ ᚋ ᖺ ௨ ෆ ࡢ Ꮚ 㤿 㸦 '

KDNDOHWDO 㸧 ࡛ ࡶ 㛗 ᪥ ᮇ ࡟ ศ Ἢ

ࡀ ቑ

ࡍ ࡿ ࡇ ࡜ ࡀ

ሗ ࿌

ࡉ ࢀ ࡚ ࠸ ࡿ ࠋ

ࣉ ࣟ

ࢡ ࢳ

ࢆ ᡂ ⇍ 㞤 㤿 ࡟ ᢞ ୚ ࡍ ࡿ ࡇ ࡜ ࡟ ࡼ ࡗ ࡚ ࠊ ᤼ ༸ ࡢ ᪩ ᮇ ໬ ࡀ ㉳ ࡇ ࡿ ࡜ ሗ ࿌ ࡉ ࢀ ࡚ ࠸ ࡿ 㸦 7 KRPSVRQHWDO㸧 ࡇ ࡜ ࠿ ࡽ ࠊ ᮏ ◊ ✲ ࡟ ࠾ ࡅ ࡿ

ࣉ ࣟ

ࢡ ࢳ

⃰ ᗘ ࡢ ୖ ᪼ ࡀ 㞤 ࡢ ṓ 㤿 ࡟ ㉳ ࡇ ࡗ ࡓ ᤼ ༸ ࡢ ᪩ ᮇ ໬ ࡟ ࡶ 㛵 ୚ ࡋ ࡚

࠸ ࡿ ࡜ ᥎ ᐹ ࡉ ࢀ ࡓ ࠋࡲ ࡓ ࠊⓎ ᝟ ࿘ ᮇ ୰ ࡢ 㞤 ࢘ ࣐ ࡟ ࠾ ࠸ ࡚ ࠊࣉ ࣟ ࣛ ࢡ ࢳ ࣥ ࡀ

᤼ ༸ ๓ ᚋ 㸦 .LQJVHWDO㸧 ࡸ 㯤 య ㏥ ⾜ ࡢ ᫬ ᮇ 㸦 .LQJVHWDO

࡟ ቑ ຍ ࡋ ࠊ༸ ᕢ ⤌ ⧊ ෆ ࡟ ࣉ ࣟ ࣛ ࢡ ࢳ ࣥ ࡸ ࢻ ࣮ ࣃ ࣑ ࣥ ࣞ ࢭ ࣉ ࢱ ࣮ ࡀ Ꮡ ᅾ ࡍ ࡿ ࡇ ࡜ 㸦 .LQJVHWDO㸧 ࠿ ࡽ ࠊ 㞤 ࢘ ࣐ ࡟ ࠾ ࠸ ࡚ ࠊ ࣉ ࣟ ࣛ ࢡ ࢳ ࣥ ࡀ ༸ ᕢ ᶵ ⬟ ࡟ 㔜 せ ࡞ ᙺ ๭ ࢆ ᢸ ࡗ ࡚ ࠸ ࡿ ࡇ ࡜ ࡀ ሗ ࿌ ࡉ ࢀ ࡚ ࠸ ࡿ ࠋ

ග ⥺ ฎ ⌮ ἲ ࢆ ᐇ ᪋ ࡋ ࡓ ṓ 㤿 ࡛ ࡣ ࠊ ᪩ ᮇ ࡟

ࣉ ࣟ

ࢡ ࢳ

⃰ ᗘ

ࡢ ୖ

ࡀ ㄆ ࡵ ࡽ ࢀ ࡓ ࠋ

ࣉ ࣟ

ࢡ ࢳ

ࣥ ࡟ ࡣ

ຠ ᯝ

ࡀ ࠶ ࡿ ࡇ ࡜ ࡀ

ሗ ࿌

ࡉ ࢀ ࡚ ࠾ ࡾ ࠊ࣐࢘࡟࠾࠸࡚ࠊග⥺ฎ⌮㸦.RRLVWUDDQG

*LQWKHUࠊ1DPERHW

DO.XQLLHWDO

6X]XNLHWDO㸧ࠊ ࣉࣟ

ࢡࢳࣥࡢ ᢞ୚

㸦7KRPSVRQHWDO㸧ࡸࢻࣃ࣑ࣥᣕᢠ๣ࡢᢞ୚㸦'RQDGXHDQG7KRPSVRQ 7KRPSVRQ㸧ࡀ⬺ẟຠᯝࢆಁ㐍ࡍࡿ࡜ሗ࿌ࡉࢀ࡚࠸ࡿࠋື ≀ ࡛ ࡣ

࠸ ᫬ ᮇ ࡟ ࡞ ࡿ ࡜

ࡉ ࠿ ࡽ

య ࢆ

ࡿ ࡓ ࡵ ࡟ ẟ ࡀ

ࡧ ࡚ ࠊ

࠿ ࠸ ᫬ ᮇ

100

࡟ ࡞ ࡿ ࡜

ࡧ ࡓ ẟ ࡀ

ẟ ࡍ ࡿ ࡇ ࡜ ࡟ ࡼ ࡗ ࡚

  ᗘ

⎔ ቃ ࡟

㐺 ᛂ

ࡋ ࡚ ࠸ ࡿ ࠋ ᮏ ◊ ✲ ࡟ ࠾ ࠸ ࡚ ࡶ ṓ 㤿 ࡢ⬺ẟ ࡟

ࣉ ࣟ

ࢡ ࢳ

ࣥ ࡀ

㛵 ୚

ࡋ ࡚ ࠸ ࡿ ࡜

᥎ ᐹ

ࡉ ࢀ ࡿ ࠋ ࡍ ࡞ ࢃ ࡕ ࠊ ᮏ ◊ ✲ ࡛ ࠊ ග ⥺ ฎ ⌮ ࢆ ᐇ ᪋ ࡋ ࡓ ࡇ ࡜ ࡟ ࡼ ࡾ ࠊ

ࣉ ࣟ

ࢡ ࢳ

⃰ ᗘ

ࡀ ᪩ ᮇ ࡟ ୖ

ࡋ ࡓ ࡇ ࡜ ࠿ ࡽ

ࣉ ࣟ

ࢡ ࢳ

ࣥ ࡀ ෤ ẟ ࡢ

ẟ ࢆ ಁ ࡋ ࠊ

ẟ ࡟ ኚ ໬ ࡉ ࡏ ࡓ ࡜

᥎ ᐹ

ࡉ ࢀ ࡓ ࠋ

ᮏ◊✲࡟ࡼࡾࠊග⥺ฎ⌮ἲࡣᙜṓ㤿࡟࠾࠸࡚ࡶ᭷⏝࡛࠶ࡿࡇ࡜ࡀึࡵ࡚᫂ࡽ

࠿࡟ࡉࢀࡓࠋ

101 0

5 10 15 20 25

D J F M A

(b)

/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

0 5 10 15

D J F M A

3URJHVWHURQHᤡQJPOᤢ

* (e)

0 1 2 3

D J F M A

(d)

LULQLKLELQᤡQJPOᤢLULQLKLELQᤡQJPOᤢLULQLKLELQᤡQJPOᤢLULQLKLELQᤡQJPOᤢ

0 5 10 15 20 25

D J F M A

(f)

(VWUDGLROᤡSJPOᤢ(VWUDGLROᤡSJPOᤢ(VWUDGLROᤡSJPOᤢ(VWUDGLROᤡSJPOᤢ

0 2 4 6 8

D J F M A

(c)

)6+ᤡQJPOᤢ)6+ᤡQJPOᤢ)6+ᤡQJPOᤢ)6+ᤡQJPOᤢ

0 1 2 3

D J F M A

EP Control

(a)

3URODFWLQᤡQJPOᤢ3URODFWLQᤡQJPOᤢ3URODFWLQᤡQJPOᤢ3URODFWLQᤡQJPOᤢ

**

Fig6-1 Changes in plasma concentrations of prolactin (a), LH (b), FSH (c), ir-inhibin(d), testosterone (e) and estradiol-17β (f) in Thoroughbred fillies in the extended photoperiod (EP) group (○) and the control group (●) from Dcember at zero year old to May at one year old. Results are expressed as means ± SEM. Letters represent first letter of each month. *P<0.05.

102

Extended Photoperiod

Fig. 6-2. Comparison of hair coat condition of representative fillies (A,B)

in the extended photoperiod group (A) and the control group (B) on March at one year old.

Control A

B

103

EP Control 0

1 2 3

Sc or e of h ai r C oa t

*

Fig.6-3 Comparison of score of hair coat condition between

the extended photoperiod (EP) group (■) and the control group (□) at March.

Results are expressed as means ±SEM. *P<0.05.

104

0 10 20 30 40 50

D J F M A

(c)

Month Extended Photoperiod

/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

0 10 20 30 40 50

D J F M A

(a)

Control

Month

/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ/+ᤡQJPOᤢ

0 5 10 15 20 25

D J F M A

Progesterone(ng/ml)

(d)

Month Extended Photoperiod 0

5 10 15 20 25

D J F M A

Progesterone(ng/ml)

(b)

Month Control

Fig. 6-4. Changes in plasma concentrations of LH (a, c) and progesterone

(b, d) in individual Thoroughbred filly in the control (a, b) and the extended

photoperiod group (c, d) from October to April at one year old. Letters

represent first letter of each month.

105

㸵❶㸵❶㸵❶

㸵❶ ⥲ᣓ⥲ᣓ⥲ᣓ⥲ᣓ

࣐࢘࡟ࡣࠊ⦾Ṫ࡟㛵ࡋ࡚ᵝࠎ࡞≉ᚩࡀᏑᅾࡍࡿࠋ⌧ᅾࠊ᪥ᮏ࡟࠾ࡅࡿ㤿⏕⏘

࡟࠾࠸࡚ࠊ➇㉮㤿⏕⏘ࡀ኱㒊ศࢆ༨ࡵ࡚࠸ࡿࠋࡑࡢࡼ࠺࡞⌧≧࡟࠾࠸࡚ࠊ⦾Ṫ

㞀ᐖࡣࠊ⦾Ṫᡂ⦼ࡢపୗࢆࡶࡓࡽࡋࠊ ⤒῭ⓗᦆኻࡣ኱ࡁ࠸ࠋ⦾Ṫ㞀ᐖ࡟࠾࠸࡚ࠊ

ෆศἪᏛⓗ࡞␗ᖖࡀཎᅉ࡜࡞ࡿࡇ࡜ࡣᑡ࡞ࡃ࡞࠸ࠋࡋ࠿ࡋࠊ᪥ᮏࡢࢧࣛࣈࣞࢵ

ࢻ✀ࡢ㞤㤿࡟࠾࠸࡚ࠊヲ⣽࡞ෆศἪᏛⓗゎᯒࡣ⾜ࢃࢀ࡚࠸࡞࠸ࠋࡲࡓࠊ᪥ᮏࡣ

௚ࡢ㤿⏘ᆅࠊ౛࠼ࡤࠊ࢖ࢠࣜࢫࠊࣇࣛࣥࢫࠊ࢔࢖ࣝࣛࣥࢻࠊ࣮࢜ࢫࢺࣛࣜ࢔ࠊ

࢔࣓ࣜ࢝࡞࡝࡜ࡣ␗࡞ࡾࠊ᫂░࡞ᅄᏘࡢኚ໬ࡀᏑᅾࡍࡿࠋࡇࡢ᪥ᮏ≉᭷ࡢᏘ⠇

ኚ໬࡟࠾࠸࡚ࠊࢧࣛࣈࣞࢵࢻ✀ࡢ㞤㤿࡛ࡣ࡝ࡢࡼ࠺࡞ෆศἪᏛⓗኚ໬ࡀㄆࡵࡽ

ࢀࡿ࠿ࡣ᫂ࡽ࠿࡟࡞ࡗ࡚࠸࡞࠸ࠋ

ᮏ◊✲࡛ࡣࠊ᪥ᮏࡢࢧࣛࣈࣞࢵࢻ✀ࡢ㞤㤿࡟࠾࠸࡚ᵝࠎ࡞ࢫࢸ࣮ࢪ࡛ࡢෆศ ἪᏛⓗኚ໬ࢆ᫂ࡽ࠿࡟ࡋࠊ௒ᚋࡢ⦾Ṫᡂ⦼ࡢྥୖࡸ⦾Ṫ㞀ᐖࡢゎ᫂࡟ᐤ୚ࡍࡿ

ࡇ࡜ࢆ┠ⓗ࡜ࡋࡓࠋ

ࡲࡓࠊ㛗᪥Ꮨ⠇⦾Ṫື≀࡛࠶ࡿ⦾Ṫ∷㤿࡟࠾࠸࡚㛗᪥ฎ⌮ࢆ᪋ࡋࠊ ᭶࠿ࡽ

᭶ࡢ▷᪥ᮇ࡛ࡶ⦾Ṫྍ⬟࡞≧ែࢆ⥔ᣢࡋࠊ஺㓄ࡀ⾜ࢃࢀ࡚࠸ࡿࠋ㛗᪥ฎ⌮ࡀ ṓ㤿࡟࠾࠸࡚➽⫗ቑᙉస⏝ࡸ᫓ᶵⓎືࡢ᪩ᮇ໬ࠊẟⰏࡢⰋ໬ࢆᘬࡁ㉳ࡇࡍࡇ࡜

ࡀ᫂ࡽ࠿࡟࡞ࡗ࡚࠸ࡿࠋ

ᮏ◊✲࡛ࡣࠊ᪥ᮏࡢࢧࣛࣈࣞࢵࢻ✀ࡢᙜṓ㤿࡟࠾࠸࡚㛗᪥ฎ⌮ࡀෆศἪᏛ

ⓗ࣭ᙧែᏛⓗ࡟ཬࡰࡍᙳ㡪࡟ࡘ࠸࡚ࡶ᳨ウࡋࡓࠋ

㸵㸫㸯㸬ࢧࣛࣈࣞࢵࢻ✀࡟࠾ࡅࡿዷፎᮇࡢෆศἪᏛⓗኚ໬

㸵㸫㸯㸬ࢧࣛࣈࣞࢵࢻ✀࡟࠾ࡅࡿዷፎᮇࡢෆศἪᏛⓗኚ໬

㸵㸫㸯㸬ࢧࣛࣈࣞࢵࢻ✀࡟࠾ࡅࡿዷፎᮇࡢෆศἪᏛⓗኚ໬

㸵㸫㸯㸬ࢧࣛࣈࣞࢵࢻ✀࡟࠾ࡅࡿዷፎᮇࡢෆศἪᏛⓗኚ໬

࣐࢘ࡢዷፎᮇ㛫ࡣ⣙ ᪥࡛࠶ࡿࠋ㐣ཤࠊከࡃࡢෆศἪᏛⓗゎᯒࡀ⾜ࢃࢀ࡚

ࡁࡓࡀࠊ༢୍ࡢ࣍ࣝࣔࣥኚື࡛࠶ࡗࡓࡾࠊ㝈ᐃࡉࢀࡓᮇ㛫࡛࠶ࡗࡓࡾ࡜ከ✀ࡢ

࣍ࣝࣔࣥኚືࢆࠊዷፎ඲ᮇ㛫ࢆ㏻ࡋ࡚ࠊྠ୍ࢧࣥࣉ࡛ࣝ ᐃ࣭ゎᯒࡋࡓሗ࿌ࡣ

106

関連したドキュメント