阿蘇海における垂下飼育によるハマグリの成長
久田哲二，尾﨑仁，谷本尚史，藤原正夢
京都府農林水産技術センター海洋センター
阿蘇海における垂下飼育によるハマグリの成長 *
久田哲二，尾﨑仁，谷本尚史，藤原正夢
Growth of venerid clams (Meretrix lusoria) during hanging culture in Asokai Lagoon Tetsuji Hisada, Hitoshi Ozaki, Naofumi Tanimoto and Masamu Fujiwara
Venerid clams (Meretrix lusoria) were subjected to hanging culture, and their growth was followed. Furthermore, the obesity level of venerid clams during hanging culture was analyzed. The results indicate that the growth period for this shellfish is April through November, and that its growth is suspended in December through to the following March. The data on the obesity level suggest that the spawning season for cultured venerid clams in Asokai Lagoon is July through September. During the growth period, arrival at the highest daily growth rate was followed by a slower growth period. Maturation seemed to be one of the factors explaining such a course of growth.
ࠠࡢ࠼㧦ࡂࡑࠣޓုਅ㘺⢒ޓᚑ㐳ޓ᳓᷷ޓ㒙⯃ᶏ ࡂࡑࠣMeretrix lusoria ߩో࿖ߩṪ₪㊂ߪ㧘1970 ᐕઍߪ 3,000 㨪 8,000 ࠻ࡦߢផ⒖ߒߡߚ߇㧘ߘࠇ એ㒠ߪỗᷫߒ㧘1980 ᐕઍ߆ࠄ 1990 ᐕઍߦߪᢙ⊖㨪 1,000࠻ ࡦ บ ߣ ߥ ࠅ㧔 ጊ ਅ ࠄ㧘2004㧕㧘2005 ᐕ ߦ ߪ 200࠻ࡦᧂḩߣߥߞߡࠆ㧔ㅺ㧘2009㧕ޕᷫዋߒߚ ࡂࡑࠣṪ₪㊂ࠍ߁ߚ㧘ㄭᐕߢߪࠪ࠽ࡂࡑࠣ 㧔Meretrix petechialis㧕ࠍߪߓߣߔࠆᄖ࿖↥ࡂࡑࠣ ߇ャߐࠇߡࠆ㧔ጊਅࠄ㧘2004㧕ޕ ޓ߹ߚޔࡂࡑࠣ↢↥㊂ࠍ࿁ᓳߐߖࠆߚߦ㧘⒳ ⧣↢↥ߐࠇߚࡂࡑࠣߩᵹ߇ⴕࠊࠇߡࠆ㧔ᩉ㧘 2009㧧ጊࠄ㧘2009㧕ઁ㧘⾗Ḯ߇ᷫዋߒߚవߢߪ㧘 ⒖ᱺᵹ߇ᬌ⸛ߐࠇߡࠆ㧔ၔࠄ㧘1983㧘᧻㊁ࠄ㧘 2008㧕ޕ ޓ╩⠪ࠄߪ㧘੩ㇺᐭߩࡂࡑࠣ↢↥㊂ᄢߩߚߩ ৻ᚻᴺߣߒߡ㧘࠻ࠟࠗFulva multica ߥߤߩẜ⍾ᕈ ੑᨎ⽴㘃㙃ᱺߢታ↪ൻߐࠇߡࠆုਅ㘺⢒㧔↰ਛࠄ㧘 2006㧕ࠍ⹜ߺߚޕߘߩਛߢ㧘࿖ౝߢᢙߩዋߥࡂ ࡑࠣߩᚑ㐳ߦߟߡ⍮ࠍᓧߚߩߢႎ๔ߔࠆޕ 材料および方法 ޓ⹜㛎ߪ㧘㒙⯃ᶏߩችᵤᏒḴዥవ᳓ᷓ 10m ၞߦ⸳ ⟎ߒߚ╱㧔એਅ㧘㒙⯃ᶏߣߔࠆޕ㧕ߢⴕߞߚ㧔Fig.1㧕ޕ ⹜㛎ߦߪ㧘㒙⯃ᶏߩ㊁↰ᎹᴡญߢṪᬺ⠪߇ណ㓸ߒߚ ᄤὼ⽴ࠍ↪㧘Ზ㐳ࠨࠗ࠭ߦ S ߅ࠃ߮ L ߩ 2 ⹜㛎ࠍ⸳ቯߒߚޕ⹜㛎㐿ᆎᤨߩฦ⹜㛎ߩᐔဋᲖ 㐳߅ࠃ߮ኈᢙߪ㧘S ߢߪ 25.8 r 3.1mm㧔ᐔဋ Ზ㐳rᮡḰᏅ㧘એਅห᭽ߣߔࠆޕ㧕߅ࠃ߮ 172 㧘 Lߢߪ 45.8 r 1.9mm ߅ࠃ߮ 90 ߢߞߚޕ⹜ 㛎ᦼ㑆ߪ 2009 ᐕ 4 21 ᣣ߆ࠄ 2011 ᐕ 10 28 ᣣ ߹ߢߣߒߚޕ ޓ⹜㛎ߪ࠻ࠟࠗߩ㙃ᱺᣇᴺ㧔↰ਛࠄ ,2006㧕ߦḰ ߓߡⴕߞߚޕ⹜㛎㐿ᆎᤨߦߪ㧘㘺⢒ኈེߦߪ㧘ࡐ ࡊࡠࡇࡦߩࠦࡦ࠹࠽㧔ౝኸ 50 32 21cm㧘 એਅ㧘ࠦࡦ࠹࠽ߣߔࠆ㧕ࠍ↪ߚ߇㧘2010 ᐕ 8 એ㒠ߪ㧘ࡐࠛ࠴ࡦߩ✂ࠞࠧ㧔ౝኸ 40 40 20cm✂⋡ 0.5mm㊎㊄ࡈࡓ㧕ߦಾࠅᦧ߃ߚޕ ߎࠇߪ㧘⹜㛎ᦼ㑆ਛߦࡈࠫ࠷ࡏ߿ࡎࡗ㘃ࠍਥߣߔ ࠆᄢ㊂ߩ↢‛߇ઃ⌕ߔࠆߎߣߦࠃߞߡࠦࡦ࠹࠽឵ ߿ᷡᬺߦ߁ഭജ߇Ⴧᄢߒߚߚߢࠆޕ㘺⢒ ኈེߩౝㇱߦߪࠕࡦࠬࠨࠗ࠻㧔☸ᓘ 2 㨪 3mm㧕 ࠍෘߐ⚂ 10cm ߦߥࠆࠃ߁ߦᢝߚޕ㘺⢒ኈེߩ 㕙ߦߪ✂⋡ 2cm ߩ✂⬄ࠍߒߡုਅߒߚޕઃ⌕↢‛ߦ ࠃߞߡౝㇱߩᶏ᳓឵߇ᅹߍࠄࠇࠆߎߣࠍ੍㒐ߔࠆ 㧖 1ᧄ⎇ⓥߩ৻ㇱߪ㧘ㄘᨋ᳓↥⋭ߩޟᣂߚߥㄘᨋ᳓↥╷ࠍផㅴߔࠆታ↪ᛛⴚ㐿⊒ᬺᆔ⸤ᬺޠߦࠃࠅታᣉߒߚޕ Asokai Lagoon Mizyazu Bay Experimental point 135㫦10′E 35㫦35′N Tango Peninsula 1km 0 Kunda Peninsula
Fig. 1ޓA map showing the site the experiment was conduced.
ߚ㧘ේೣߣߒߡ 1 ࡩᲤߩ㗫ᐲߢ㘺⢒ኈེߩ឵ ࠍⴕߞߚޕ㘺⢒ኈེߩ឵ᤨߦߪ⚂ 2cm ⷺ⋡ߩ✂ࡈ ࡞ࠗߢࠕࡦࠬࠨࠗ࠻ౝߩࡂࡑࠣએᄖߩ↢‛ࠍน ⢻ߥ㒢ࠅขࠅ㒰߈㧘ኈེౝߦၸⓍߒߚᶋᵆ߿♮╬ࠍ 㒰ߒߚޕߎߩߣ߈㧘ࠕࡦࠬࠨࠗ࠻߇ᷫዋߒߡ ࠇ߫㧘⹜㛎㐿ᆎᤨߣห⒟ᐲ߹ߢలߒߚޕုਅ᳓ᷓߪ㧘 㒙⯃ᶏߦ߅ߌࠆࠕࠨߩ㧔⼱ᧄࠄ㧘2011㧕ࠍෳ ⠨ߦ㧘ᦨ߽ࠕࠨߩᚑ㐳߇⦟߆ߞߚ 3m ᷓߣߒߚޕ ޓේೣߣߒߡ 1 ࡩᲤߦ 50 ߹ߚߪోߩᲖ㐳 ࠍ⸘᷹ߒߚޕ⸘᷹ߪ㧘╱ߢ࠺ࠫ࠲࡞ࡁࠡࠬ㧔ࡒ࠷ ࠻࡛ CD20-PMX㧕ࠍ↪ߡⴕ㧘0.1mm නߢ ⸥㍳ߒߚޕ߹ߚ㧘ቄ▵ߦࠃࠆᚑ㐳ߩᄌേࠍᲧセߔࠆ ߚ㧘ᣣ㑆ᚑ㐳₸㧔Gr㧔㧑 /day㧕㧕ࠍᰴᑼߦࠃࠅ▚ ߒߚޕ 2 10 T2 T1 （L2−L1）／L1 Gr × − = 㧔ߎߩߣ߈㧘L1ߪᤨ㑆 T1ߦ߅ߌࠆᐔဋᲖ㐳㧘L2ߪᤨ 㑆 T2ߦ߅ߌࠆᐔဋᲖ㐳ߢࠆޕᤨ㑆ߩනߪᣣ㧔day㧕 ߢࠆޕ㧕 ޓုਅ㘺⢒ߒߚࡂࡑࠣߩりࠅ⁁ᴫࠍ⺞ᩏߔࠆ ߚ㧘Ṫᬺ⠪߇ห╱ߦ߅ߡ㧘100 / ኈེߩኒᐲ ߢ 2009 ᐕ 4 ߆ࠄုਅ㘺⢒ߒߡߚࠍ⾼ߒ㧘 2009ᐕ 9 ߆ ࠄ 2010 ᐕ 7 ߩ 㑆 ߦ ⸘ 7 ࿁ ߩ ⢈ ḩ ᐲ㧔CF㧕ߩ᷹ቯࠍⴕߞߚޕߎߩߣ߈㧘⢈ḩᐲ▚ߦ ߪࠕࠨߢ↪ࠄࠇߡࠆᰴᑼ㧔㠽⠀㧘ᷓጊ ,1991㧕 ࠍ↪ߚޕ 5 10 SW SH SL STW CF × × × = 㧔ߎߩߣ߈㧘STW ߪエㇱ㊀㊂㧘SL ߪᲖ㐳㧘SH ߪ Ზ㜞㧘SW ߪᲖߢࠆޕ㧕エㇱ㊀㊂ߪ㔚ሶᄤ⒊ࠍ ↪ߡ 0.1g නߢ㧘Ზ㐳㧘Ზ㜞߅ࠃ߮Ზߦߟߡ ߪ࠺ࠫ࠲࡞ࡁࠡࠬ㧔ࡒ࠷࠻࡛ CD20-PMX㧕ࠍ↪ ߡ㧘0.1mm නߢ᷹ቯߒߚޕߥ߅㧘৻ᐲߩ᷹ቯߦ ߪ 30 ߆ࠄ 50 ߩࡂࡑࠣࠍ↪ߚޕ᷹ቯᤨߩᐔ ဋᲖ㐳㧔rᮡḰᏅ㧕ߪ 49.6 r 2.8mm 㨪 59.9 r 1.8mmߩ▸࿐ߢߞߚޕ ޓ㘺⢒᳓ᷓߩ᳓᷷ࠍࡂࡑࠣߩ⸘᷹ᤨߦ᷹ቯߒߚޕ ᷹ ቯ ߦ ߪ㧘2009 ᐕ 4 ߆ ࠄ 2010 ᐕ 2 ߹ ߢ ߪ ု ਅᑼᄙᯏ⢻ࡦࠨ㧔ၳ႐ᚲ , ᳓⾰࠴ࠚ࠶ࠞ U-10߹ ߚ ߪ U-50㧕 ࠍ㧘2010 ᐕ 3 ߆ ࠄ 2011 ᐕ 10ߦߟߡߪ㧘ုਅᑼᄙ㗄⋡᳓⾰ࡦࠨ࠱ࡦ࠺ 㧔ࠕ࠶ࠢ㔚ሶ␠㧘ࠢࡠࡠ࠹࠶ࠢ ACL208-DK㧕ࠍ ↪ߚޕ 結 果 ޓᚑ㐳ޓਔ⹜㛎ߩᐔဋᲖ㐳ߩផ⒖ࠍ Fig.2 ߦ␜ߒ ߚޕ2011 ᐕ 10 28 ᣣߩ⚳ੌᤨߦߪ㧘S ߢ 65.7 r 10mm㧘L ߢ 71.4 r 6.7mm ߢ ߞ ߚޕFig.3 ߦᣣ㑆ᚑ㐳₸㧔Gr㧕ߣ᳓᷷ߩផ⒖ࠍ߹ߣߡ␜ߒߚ 㧔ࠣࡈߢߪᣣ㑆ᚑ㐳₸ߪᦼ㑆ਛ㑆ᣣߦࡊࡠ࠶࠻ߒ ߚޕ㧕ޕోᦼ㑆ࠍㅢߒߚᣣ㑆ᚑ㐳₸ߪ㧘S ߢߪ 0.01 㨪 0.30㧑 /day ߩ▸࿐ߦࠅ㧘ోᦼ㑆ࠍㅢߓߚᐔဋ ୯ߪ 0.09㧑 /day ߢߞߚޕL ߢߪ㧘0.00 㨪 0.10 㧑 /day ߩ▸࿐ߢ㧘ోᦼ㑆ࠍㅢߓߚᐔဋ୯ߪ 0.05㧑 /dayߢ ߞ ߚޕ2009 ᐕ 4 21 ᣣ ߆ ࠄ 2010 ᐕ 4 27 ᣣߩ⚂ 1 ᐕ㑆㧔એਅ㧘1 ᐕ⋡ߣߔࠆޕ㧕ߦ߅ ߡ㧘ᣣ㑆ᚑ㐳₸ߩᦨ㜞୯ࠍ␜ߒߚߩߪ S 㧘L ߣ ߽ߦ㧘4 21 ᣣ㨪 6 24 ᣣߩᦼ㑆ߢ㧘ߎߩᦼ㑆ߩ ᳓᷷ߪ 16.5 㨪 23.0͠ߩ▸࿐ߢߞߚޕᣣ㑆ᚑ㐳₸ ߇ᦨૐ୯ࠍ␜ߒߚߩߪ S 㧘L ߣ߽ߦ㧘1 24 ᣣ 㨪 2 24 ᣣߩᦼ㑆ߢࠅ㧘ߎߩᦼ㑆ߩ᳓᷷ߪ 11.1 ͠ ߢ ߞ ߚޕ2010 ᐕ 4 27 ᣣ 㨪 2011 ᐕ 4 13 ᣣ㧔એਅ㧘2 ᐕ⋡ߣߔࠆޕ㧕ߩ 1 ᐕ㑆ߦߟߡߪ㧘ᣣ 㑆ᚑ㐳₸߇ᦨ㜞୯ࠍ␜ߒߚߩߪ㧘S 㧘L ߣ߽ߦ㧘 10 15 20 25 30 0.10 0.15 0.20 0.25 0.30 㪮 㪸㫋 㪼㫉 㪫 㪼㫄 㫇㪼 㫉㪸 㫋㫌 㫉㪼 㩿㷄 㪀㩷 㪣 㪪 㪮㪅㪫 0 5 0.00 0.05
AM J J A SOND J FMAM J J A SOND J FMAM J J A SO
㪉㪇㪇㪐 㪉㪇㪈㪇 㪉㪇㪈㪈
Growth rate(
%
)
Fig. 3ޓChanges in growth rate (%/day) of Meretrix
lusoria on hanging culture and the water
temperature (°C) at a 3-m depth in Asokai lagoon. ●:L;○:S. The dashed line shows the water temperature. 30 40 50 60 70 80 㪣 㪪 0 10 A M J J A S O N D J F M A M J J A S O N D J F M A M J J A S O 㪉㪇㪇㪐 㪉㪇㪈㪇 㪉㪇㪈㪈 20 A verage shell length(mm)
Fig. 2ޓGrowth cur ves of Meretrix lusoria on hanging culture at a 3-m depth in Asokai lagoon. Filled circles (●) show experiment division L, and open circles (○) show experiment division S. Vertical lines indicate standard deviations.
4 27 ᣣ㨪 6 24 ᣣߩᦼ㑆ߢ㧘ߎߩᦼ㑆ߩ᳓᷷ߪ 13.9㨪 22.9͠ߢߞߚޕᣣ㑆ᚑ㐳₸߇ᦨૐ୯ࠍ␜ߒ ߚߩߪ 1 21 ᣣ㨪 3 11 ᣣߩᦼ㑆ߢ㧘ߎߩᦼ㑆ߩ ᳓᷷ߪ 11.1 㨪 11.4͠ߢߞߚޕ1 ᐕ⋡߅ࠃ߮ 2 ᐕ⋡ ߦߟߡߪ㧘ࡂࡑࠣ߇ᦨ߽ᚑ㐳ߒߚᤨᦼߪ 4 㨪 6 ߩ᳓᷷ᦼߢ㧘ᦨ߽ᚑ㐳ߩṛߔࠆᤨᦼߪ 1 㨪 3ߩᦨૐ᳓᷷ᦼߢߞߚޕ߹ߚ㧘ᣣ㑆ᚑ㐳₸߇ᐕ 㑆ߩᐔဋ୯એ㧔1 ᐕ⋡ߩ S ߢߪ 0.12%/day㧘L ߪ 0.06%/day㧘2 ᐕ ⋡ ߩ S ߪ 0.09%/day㧘L ߪ 0.04%/day એ㧕ߢߞߚᦼ㑆ߪ㧘1 ᐕ⋡ߩ S ߢߪ㧘4 21 ᣣ㨪 6 24 ᣣߩᦼ㑆㧘8 5 ᣣ㨪 10 6 ᣣߩᦼ㑆㧘11 18 ᣣ㨪 12 10 ᣣߩᦼ㑆ߢ ߞ ߚޕL ߢ ߪ 4 21 ᣣ 㨪 8 5 ᣣ ߩ ᦼ 㑆㧘9 1ᣣ㨪 12 10 ᣣ߹ߢߩᦼ㑆㧘2 ᐕ⋡ߩ S ߢߪ 3 24 ᣣ㨪 11 18 ᣣߩᦼ㑆㧘L ߢߪ 3 24 ᣣ㨪 6 23 ᣣ㧘7 22 ᣣ㨪 10 18 ᣣ㧘11 18 ᣣ㨪 12 14 ᣣߩᦼ㑆ߢߞߚޕߎߩࠃ߁ߦ㧘1 ᐕ⋡ߣ 2ᐕ⋡ߦߟߡߪ㧘ᚑ㐳ߔࠆᤨᦼߪ 4 㨪 11 㧘ᚑ㐳 ߇ṛߔࠆᤨᦼߪ㧘᳓᷷ 15͠એਅߩ 12 㨪 3 ߢ ߞߚޕ ޓ৻ᣇ㧘1 ᐕࠍㅢߓߚ࠺࠲ߪߥ߽ߩߩ㧘2011 ᐕ 4 13 ᣣ㨪 10 28 ᣣߩ 7 ࡩ㑆㧔એਅ㧘3 ᐕ⋡ߣ ߔࠆޕ㧕ߩᣣ㑆ᚑ㐳₸ࠍߺࠆߣ㧘S ߦߟߡߪ㧘6 9 ᣣ㨪 7 21 ᣣߩᦼ㑆ߦ㧘L ߦߟߡߪ 7 21ᣣ㨪 8 24 ᣣߩᦼ㑆ߦૐਅߒߚᓟ㧘ౣ߮ߒ ߡߚޕ 1 ᐕ⋡߆ࠄ 3 ᐕ⋡ߩ S 㧘L ߩోߡߢㅢߒߚ ․ᓽߣߒߡߪ㧘ᣣ㑆ᚑ㐳₸ߪ 4 㨪 7 ߩ᳓᷷ᦼ ߦ㜞୯ࠍ␜ߒߚᓟ㧘7 㨪 8 ߦ߆ߌߡ৻ᐲૐਅߒ㧘 ߘߩᓟౣ߮ߒߡߚޕ߹ߚ㧘᳓᷷߇ 15͠ࠍਅ࿁ ࠆߣ㧘ᚑ㐳ߪ߶ߣࠎߤṛߒߡߚޕߥ߅㧘᷹ቯᣣ 㑆ߩᐔဋ᳓᷷ߣᣣ㑆ᚑ㐳₸ߦߪ㧘⍎ߥ㑐ଥߪߺࠄ ࠇߥ߆ߞߚ㧔Fig.4㧕ޕ ޓ⢈ḩᐲߩផ⒖ޓ⢈ḩᐲߩផ⒖ࠍ Fig.5 ߦ␜ߒߚޕ ⢈ḩᐲߪ 2009 ᐕ 9 ߦߪ⹜㛎ᦼ㑆ਛߩᦨૐ୯ 10.4 r 2.3㧔ᐔဋ୯rᮡḰᏅ㧕ߢߞߚ߇㧘ߘߩᓟ ߒߡ 1 ߦߪ 15.1 r 2.7 ߣߥߞߚޕ3 ߆ࠄ 6 ߹ߢߪ 17 ೨ᓟߩ㜞᳓Ḱߢផ⒖ߒ㧘7 ߦߪ 13.7 r 1.3߹ߢૐਅߒߚޕ 考 察 ޓᧄ⎇ⓥߢߪ㧘1 ᐕ⋡߆ࠄ 3 ᐕ⋡߹ߢߩోߡߩ⹜㛎 ߦㅢߒߡ 4 㨪 7 ߦᣣ㑆ᚑ㐳₸ߪ㜞୯ࠍ␜ߒ ߚᓟ㧘7 㨪 8 ߦߪ৻ᐲૐਅߒߡ߅ࠅ㧘ᚑ㐳߇㊰ൻ ߒߡࠆᦼ㑆߇ࠄࠇߚ㧔Fig.3㧕ޕ⢈ḩᐲߩផ⒖ࠍ ߺࠆߣ㧘9 ߦᦨૐ୯ࠍ␜ߒߡߘߩᓟߒ㧘3 㨪 6 ߦᦨ㜞୯ઃㄭߢផ⒖ߒߚᓟ㧘7 ߦߪૐਅߒߡ ߚ㧔Fig.5㧕ޕࡂࡑࠣߩᚑᾫߣり₸㧔エㇱ㊀㊂ /㊀ 100㧦ᧄ⎇ⓥߢߩ⢈ḩᐲߦ⋧ᒰߔࠆ୯㧕ߪࠃ ߊ৻⥌ߔࠆ㧔ၔࠄ㧘1985㧕ߎߣ߆ࠄ㧘㒙⯃ᶏߦ߅ ߌࠆࡂࡑࠣߩ↥ෆᦼߪ 7 㨪 9 ߣផቯߐࠇࠆޕᧄ ⎇ⓥߣห᭽ߩုਅ㘺⢒ࠍⴕߞߚࠕࠨߢߪ㧘ᚑᾫ ↥ෆߦ߁ᚑ㐳ߩṛ߇⏕ߐࠇߡࠆ㧔⮮ේࠄ㧘 2008㧕ޕࡂࡑࠣߦߟߡ߽㧘7 㨪 8 ߩᚑ㐳㊰ൻ ߪᚑᾫ↥ෆ߇৻ߟߩⷐ࿃ߣߥߞߡࠆน⢻ᕈ߇⠨ ߃ࠄࠇߚޕ ޓ᧲੩ḧᄢᐓẟ㧔ਛࠄ㧘2009㧕ߩ㘺⢒ታ㛎ߢ ߪ㧘౻ቄߦߪᚑ㐳߇ṛߒߡ߅ࠅ㧘ૐ᳓᷷ᦼߦߟ ߡߪᧄ⎇ⓥߣห᭽ߩ⚿ᨐߢߞߚޕ৻ᣇ㧘ᦨ㜞ߩᚑ 㐳₸㧔ߚߛߒ㧘ߎߎߢߪⓍᚑ㐳₸㧔Gv㧕ߢࠆޕ㧕 ࠍ␜ߒߚߩߪ㧘8 㧔᳓᷷ 25 㨪 30͠એ㧕ߢࠅ㧘 ࡂࡑࠣߩᚑ㐳ߪ᳓᷷ߣᗧߥ⋧㑐㧔2 ߟߩ⹜㛎 ߢ r2=0.49,r2=0.60㧕߇ߺࠄࠇߚޕߒ߆ߒ㧘ᧄ⎇ⓥߢ ߪ᧲੩ḧߩ㧔ਛࠄ ,2009㧕ߣߪ⇣ߥࠅ㧘7 㨪 9 㪌 㪈㪇 㪈㪌 㪉㪇 㫆㫅㪻㫀㫋㫀㫆㫅㩷 㪽㪸㪺㫋㫆㫉 㪇 㪪 㪦 㪥 㪛 㪡 㪝 㪤 㪘 㪤 㪡 㪡 㪘 㪚㫆 2009 2010
Fig. 5ޓFluctuations of condition factor in Meretrix
lusoria on hanging culture at a 3-m depth in
Asokai lagoon. Vertical lines indicate standard deviations. 0.1 0.15 0.2 0.25 0.3 0.35 Gr owth rat e(%/da y) 0 0.05 0 10 20 30 Water temperature(͠)
Fig. 4ޓRelationship between the water temperature ( °C) and growth rate (%/day) of Meretrix lusoria on hanging culture at a 3-m depth in Asokai lagoon.
The water temperature and growth rate are average values between the start and finish of each observation.
㧔᳓᷷ 25 㨪 30͠㧕ߩᦼ㑆ߩᚑ㐳ߪ㧘4 㨪 6 ߩ ᦼ㑆㧔᳓᷷ 13 㨪 23͠㧕ߦᲧߴߡഠࠆߎߣ߇ᄙ߆ߞ ߚ㧔Fig.3㧕ޕ߹ߚ㧘᳓᷷ߣᣣ㑆ᚑ㐳₸ߩ㑆ߦߪ৻ቯ ߩ㑐ଥ߇ᛠីߢ߈ߥ߆ߞߚ㧔Fig.4㧕ޕ᧲੩ḧߩ㧔ਛ ࠄ㧘2009㧕ߢߪ㧘ଏ⹜⽴ࠍ⹜㛎㐿ᆎᲤߦࠇᦧ߃ ߡ㧘ଏ⹜⽴ߩ㐿ᆎᤨᲖ㐳ߪᏱߦ 15 㨪 20mm ߢߞ ߚޕߎࠇߪ㧘ࡂࡑࠣߩᚑᾫᦨዊࠨࠗ࠭ߢࠆ 17 㨪 20mm㧔Nakamura at el.,2010㧕ߦㄭߊ㧘ᧄ⎇ⓥ ߢ↪ߚߦᲧߴߡዊߐ߆ߞߚޕ৻⥸⊛ߦ㧘േ‛ ߢߪ⧯ᤨᦼߦߪหൻߒߚࠛࡀ࡞ࠡߩᄢㇱಽߪᚑ 㐳ߦ㧘ᚑᾫᐕએ㒠ߪ❥ᱺߦᝄࠅะߌࠄࠇࠆ㧔⩵ᳰ㧘 1999㧕ޕࠕࠨߢߪ㧘ᄢဳߪዊဳߦᲧߴߡ㧘 ៨ขࠛࡀ࡞ࠡࠍᚑ㐳ࠃࠅ߽ᚑᾫߦలᒰߔࠆߚ㧘 ᄢဳߩᚑ㐳ߪዊဳࠃࠅ߽ഠࠆ㧔㠽⠀ᷓጊ㧘 1994㧕ߣߐࠇߡࠆޕߎࠇࠄࠍෳ⠨ߦߔࠆߣ㧘ࡂࡑ ࠣߦߟߡ߽ᚑᾫ߇㑐ਈߔࠆߎߣߦࠃߞߡ㧘 ࠨࠗ࠭ߦࠃࠅ㧘ᦨ߽ᚑ㐳ߔࠆᤨᦼ߿᳓᷷߇⇣ߥࠆߎ ߣ߇੍ᗐߐࠇࠆޕߎࠇ߇㧘ᧄ⎇ⓥߣ᧲੩ḧߩߢ ⇣ߥࠆ⚿ᨐߣߥߞߚⷐ࿃ߩ৻ߟߢߪߥ߆ߣ⠨߃ࠄ ࠇߚޕ 謝 辞 ޓᧄ⎇ⓥࠍⴕ߁ߦߚࠅ㧘੩ㇺᐭṪᬺදห⚵วችᵤ ᡰᚲḴዥߩṪᬺ⠪ߦߪ╱ߩ৻ㇱಽࠍုਅ⹜㛎႐ ᚲߣߒߡឭଏߒߡ㗂ߚޕ߹ߚ㧘㘺⢒ᬺ╬⹜㛎ߦ 㑐ߔࠆ᭽ޘߥදജࠍ⾦ߞߚޕᔃࠃࠅᗵ⻢↳ߒߍ߹ ߔޕ 文 献 ⮮ේᱜᄞ㧘ㄞ⑲ੑ㧘↰ਛ㓷ᐘ㧘৻㧘ਛ㓷ᐘ㧚 2008㧚ုਅࠦࡦ࠹࠽㘺⢒ߦ߅ߌࠆࠕࠨߩᚑ㐳㧚 ੩ㇺᐭᶏᵗ⎇ႎ㧘:49-53㧚 ㅺᵏਭ㧚2009㧚╙ 4 ┨ޓࡂࡑࠣߩ↢‛ቇ㧘╙ 5 ┨ޓᣣᧄฦߦ߅ߌࠆࡂࡑࠣߩ⁁㧚ޟ⢈ᓟࡂ ࡑࠣߩ⾗Ḯ▤ℂߣࡉࡦ࠼ൻޠ㧚81-152㧚ᚑᢥ ၴ㧘᧲੩㧚 ၔ⟵ା㧘ޓᤘ㧘ᮮ᧻⧐ᴦ㧘᧲ᰵੑ㧘↰ᨴඳ┨㧘 ࿖ᱞੱ㧘⍹Ꮉม㧘ᣣᶆ⌀ℂሶ㧚1983㧚ਛ㑆 ⢒ᚑޓࡂࡑࠣߩ⾗Ḯၭ㙃ᛛⴚ㐿⊒⎇ⓥޓᤘ 57ᐕᐲ⎇ⓥ㐿⊒ଦㅴᬺ㧚19-35㧚ᄢಽ⋵ᵻᶏ Ṫᬺ⹜㛎႐㧚 ၔ⟵ା㧘ᮮ᧻⧐ᴦ㧘᧲ᰵੑ㧘1985㧚⒳⧣↢↥ޓ ࡂࡑࠣߩ⾗Ḯၭ㙃ᛛⴚ㐿⊒⎇ⓥޓᤘ 59 ᐕᐲ ⎇ⓥ㐿⊒ଦㅴᬺ㧚1-19㧚ᄢಽ⋵ᵻᶏṪᬺ⹜㛎႐㧚 ⩵ᳰᵏੑ㧘1999㧚╙ 10 ┨ޓᚑ㐳ߣᐕ㧘ޟエേ‛ ቇ⺑ޓਅᏎޠ㧚339-348㧚ࠨࠗࠛࡦ࠹ࠖࠬ࠻␠㧘 ᧲੩㧚 ጊ ผ ᐽ㧘 ᬀ ᧁ ▸ ⴕ㧘 ᓟ ⮮ ⌀ ᮸㧘 ᳓ ᚭ 㥏㧚2009㧚 ࡂ ࡑ ࠣ ߩ ⒳ ⧣ ↢ ↥ ⹜ 㛎㧚 ጟ ጊ ᳓ ⹜ ႎ㧚 :128-131㧚 ਛᐽ↵㧘ዊᴰቁ⟵㧘Ꮢᐽ㧘ᧁᐈ㇌↵㧚2009㧚᧲ ੩ḧߩੱᎿᐓẟߦ߅ߌࠆࡂࡑࠣߩ↢ᱷߣᚑ㐳 ޓࠤࠫታ㛎ߦ߅ߌࠆઁ⒳ߣߩᲧセ㧚᳓ⅣႺቇ ળ㧚㧦549-555㧚
Nakamura Y.,Nakano T.,Yurimoto T.,Maeno Y.,Koizumi A.,Tamaki A. 2010.Reproductive cycle of the venerid clam Meretrix lusoria in Ariake Sound and Tokyo Bay,Japan.Fish.Sci.,: 931-941. ↰ਛ㓷ᐘ㧘⼱ආᔒ㧘⮮ේᱜᄞ㧚2006㧚࠻ࠟࠗ㙃 ᱺߦ㑐ߔࠆ⎇ⓥ㧙Χޓዊဳᄌᒻ⽴ߩߣ㒐ᱛ ᣇᴺ㧚੩ㇺᶏᵗ⎇ႎ㧘:6-10㧚 ⼱ᧄዏผ㧘ਛ㓷ᐘ㧘ਭ↰ືੑ㧘የ㦮ੳ㧘⮮ේᱜ ᄞ㧚2011㧚㒙⯃ᶏߦ߅ߌࠆုਅ㘺⢒ߦࠃࠆࠕ ࠨߩᚑ㐳㧘↢ᱷ㧘⢈ḩᐲ㧚੩ㇺᶏᵗ⎇ႎ㧘 11-23㧚 㠽⠀శ᥍㧘ᷓጊ⟵ᢥ㧚1991㧚㘺⢒ࠕࠨߩᕈᚑᾫㆊ ⒟ߣ↥ෆ⺃⊒㧚ᣣ᳓㧘㧔7㧕:1269-1275㧚 㠽⠀శ᥍㧘ᷓጊ⟵ᢥ㧚1994㧚㘺⢒ࠕࠨߩࠨࠗ࠭ߣ ᚑᾫ㧘↥ෆߩ㑐ଥ㧚ᣣ᳓㧘㧔2㧕:173-178㧚 ጊਅඳ↱㧘⮮ᘕ৻㧘㊄ᢘḮ㧘ㅺᵏਭ㧘㐳↰⧷Ꮖ㧘 ጊᧄ⨃㓶㧘ᳰญሶ㧘᳓㑆㊀㧘ฬޓ⚐㧘㜞 ፉޓ㤀㧚2004㧚ᴉ㤩ߩᐓẟޓࡂࡑࠣࠍㅢߒߡ ࠆࠕࠫࠕߩᶏߣ㘩ߩᧂ᧪㧚㜞ᧁၮ㊄ഥᚑႎ๔ 㓸㧘:85-91㧚 ᩉື㓶㧚2009㧚ࡂࡑࠣṪ႐ౣ↢ޓ᪀ฬᏒ⿒㗇⾐Ṫ ද㧚Ꮊᄢቇᔕ↪ജቇ⎇ⓥᚲᚲႎ㧚 29-31㧚
