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BRVB ࡢ VP3 㸪 NSP3 ࠾ࡼࡧ NSP4 㑇ఏᏊࡢሷᇶ㓄ิ࡞ࡽࡧ࡟ྛ⺮ⓑࡢ

ࣀ㓟㓄ิࡢ┦ྠᛶࡢ᳨ウ

BRVB ࡢ VP3㸪NSP3࠾ࡼࡧ NSP4 ࡢ᏶඲㛗CDS ࡢሷᇶ࠾ࡼࡧ࢔࣑ࣀ㓟㓄

ิࢆ௚ࡢRVBᰴ࡜ẚ㍑ࡋࡓࠋBRVBࢆ௚ࡢRVB࡜ẚ㍑ࡋࡓ࡜ࡁ㸪VP3㸪NSP3

࠾ࡼࡧ NSP4 㡿ᇦࡢ┦ྠᛶࡣሷᇶ㓄ิ࡛ 63.3–64.9%㸪65.9–68.2%࠾ࡼࡧ 52.6–56.2%㸪࢔࣑ࣀ㓟㓄ิ࡛60.9–63.0%㸪56.4–60.3%࠾ࡼࡧ36.1–39.3%࡛࠶

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ࡗࡓ㸦⾲ 3-2㹼3-4㸧ࠋVP3㸪NSP3 ࠾ࡼࡧ NSP4 㡿ᇦࡢࣛࢵࢺ㸪ࣄࢺ࠾ࡼࡧ㇜

RVBࡢ㑇ఏᏊᆺࢆM1-T1-E1㸪M2-T2-M2࠾ࡼࡧM4-T4-E4࡜ࡋ㸪BRVBࡢ㑇

ఏᏊᆺࢆ M3-T3-E3 ࡜௬࡟࿨ྡࡋࡓࠋᆺู࡟᭱ࡶ㐺ᙜ࡞࢝ࢵࢺ࢜ࣇ್ࢆ㸪┦ྠ

ᛶศᕸᅗ࡟࠾࠸࡚㑇ఏᏊᆺእ┦ྠᛶࡢฟ⌧㢖ᗘࢆ㑇ఏᏊෆ┦ྠᛶࡢฟ⌧㢖ᗘ࡛

㝖ࡋࡓ್ࡀ㸯ࢆୗᅇࡿ఩⨨࡟࠾ࡅࡿ┦ྠᛶࡢ್࡜ࡋࡓࠋࡇࢀ࡟ᚑࡗ࡚ィ⟬ࡍࡿ

࡜㸪ሷᇶ㓄ิ࠾ࡼࡧ࢔࣑ࣀ㓟ࡢ࢝ࢵࢺ࢜ࣇ್ࡣ㸪VP3࡛ࡑࢀࡒࢀ70%࠾ࡼࡧ73%㸪 NSP3࡛78%࠾ࡼࡧ74%㸪NSP4࡛70%࠾ࡼࡧ69%࡛࠶ࡗࡓ㸦ᅗ3-5㹼3-7㸧ࠋ

4. ඲㑇ఏᏊࢭࢢ࣓ࣥࢺ࡟࠾ࡅࡿศᏊ⣔⤫ᶞゎᯒ

᏶඲㛗ࡢ VP3㸪NSP3࠾ࡼࡧ NSP4ࡢ࢔࣑ࣀ㓟㓄ิ࡟ࡼࡿ⣔⤫ᶞゎᯒ࡛ࡣ㸪 BRVBࡣM3㸪T3࠾ࡼࡧE3࡜௬࡟࿨ྡࡋࡓࢢ࣮ࣝࣉࢆᙧᡂࡋ㸪ࣄࢺ㸦M2-T2-E2㸧㸪

㇜㸦M4-T4-E4㸧࠾ࡼࡧࣛࢵࢺ㸦M1-T1-E1㸧ࡢ RVB ࡜ࡣ㞳ࢀ࡚ࢡࣛࢫࢱ࣮ࢆ

ᙧᡂࡋࡓ㸦ᅗ3-8 a, b, c㸧ࠋ

ḟ࡟㒊ศⓗ࡞VP7㸪VP4㸪VP6㸪VP1㸪VP2㸪NSP1㸪NSP2࠾ࡼࡧNSP5࢔

࣑ࣀ㓟㓄ิࢆ⏝࠸࡚⣔⤫ᶞゎᯒࢆ⾜ࡗࡓࠋVP7 ࡟ࡼࡿ⣔⤫ᶞゎᯒ࡛ࡣ㸪ᡃࡀᅜ ࡢBRVBᰴࡣ⡿ᅜࡢBRVB࡜࢔࣑ࣀ㓟㓄ิ࡛95.8㸫99.2%ࡢ┦ྠᛶࢆ♧ࡋ㸪࡜

ࡶ࡟G3㑇ఏᏊᆺ࡟ศ㢮ࡉࢀࡓࠋ࢖ࣥࢻࡢBRVBࡣ㸪ᡃࡀᅜࡢBRVB࡜࢔࣑ࣀ

㓟㓄ิ࡛85.0㹼88.8㸣ࡢ┦ྠᛶࢆ♧ࡋࡓࡀ㸪ᡃࡀᅜࡢBRVB࡜ࡣᑡࡋ␗࡞ࡗࡓ ศᒱࢆ♧ࡋ㸪G5㑇ఏᏊᆺࢆᙧᡂࡋࡓ㸦ᅗ3-9 A㸪⾲3-5㸧ࠋNSP2࡟ࡘ࠸࡚ࡣ㸪 ᡃࡀᅜࡢBRVBࡣ࢖ࣥࢻࡢBRVB࡜ࡣ࢔࣑ࣀ㓟࡛ࣞ࣋ࣝ89.1-92.0㸣㸪㇜RVB

࡜ࡣ 86.2-89.1㸣ࡢ┦ྠᛶࢆ♧ࡋ㸪࢖ࣥࢻࡢ BRVB࠾ࡼࡧ㇜ RVB ࡜࡜ࡶ࡟N2

㑇ఏᏊᆺ࡟ศ㢮ࡉࢀࡓ㸦ᅗ3-9 G㸪⾲3-11㸧ࠋNSP5࡛ࡣ㸪ᡃࡀᅜࡢBRVBࡣ࢖

ࣥࢻࡢBRVB࡜࢔࣑ࣀ㓟㓄ิ࡛85.6-88.6㸣ࡢ┦ྠᛶࢆ♧ࡋ㸪࡜ࡶ࡟H6㑇ఏᏊ ᆺ࡟ศ㢮ࡉࢀࡓ㸦ᅗ3-9 H㸪⾲3-12㸧ࠋVP6࡛ࡣ㸪ᡃࡀᅜࡢBRVBࡣ࢖ࣥࢻࡢ

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BRVB ࡜ࡣ 92.1-95.2㸣ࡢ┦ྠᛶࢆ♧ࡋ㸪ഹ࠿࡟ศᒱࡀ␗࡞ࡿࡀ㸪࡜ࡶ࡟ I3 ࢡ

ࣛࢫࢱ࣮ࢆᙧᡂࡋࡓ㸦ᅗ3-9 C㸪⾲3-7㸧ࠋNSP1࣌ࣉࢳࢻ2࡟ࡘ࠸࡚ࡣ㸪ᡃࡀᅜ ࡢBRVBࡣ⊂❧ࡋࡓࢡࣛࢫࢱ࣮ࢆᙧᡂࡋࡓࡀ㸪ࣄࢶࢪࡢRVB࡜67.9–68.8%ࡢ

࢔࣑ࣀ㓟┦ྠᛶࢆ♧ࡋ㸪࡜ࡶ࡟ I3 㑇ఏᏊᆺࢆᙧᡂࡋࡓ㸦ᅗ 3-9 F㸪⾲ 3-10㸧ࠋ NSP1࣌ࣉࢳࢻ1࡟ࡘ࠸࡚ࡣG-2006ࡢࡳࡢ㓄ิࡋ࠿ᚓࡽࢀࡎ㸪ࡇࡢ㓄ิࡣࣄࢶ

ࢪ RVB ࡢࡑࢀ࡜ 75.2%ࡢ࢔࣑ࣀ㓟㓄ิ┦ྠᛶࢆ♧ࡋࡓ㸦ࢹ࣮ࢱᮍᥖ㍕㸧ࠋ㑇ఏ

Ꮚᆺࡀᥦၐࡉࢀ࡚࠸࡞࠸ VP4㸪VP1 ࠾ࡼࡧ VP2 ࡟ࡘ࠸࡚ࡣ㸪ᡃࡀᅜࡢ BRVB ࡣ࢖ࣥࢻࡢ BRVB ࡜࢔࣑ࣀ㓟࡛ࣞ࣋ࣝࡑࢀࡒࢀ 90.0–91.7%㸪85.4–87.6%࠾ࡼ

ࡧ94.9–97.1%ࡢ┦ྠᛶࢆ♧ࡋ㸪ྠࡌࢡࣛࢫࢱ࣮࡟ᒓࡋࡓ㸦ᅗ3-9 B, D, E㸪⾲

3-6, 3-8, 3-9㸧ࠋ

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⪃ ᐹ

ᮏ❶࡟࠾ࡅࡿゎᯒ࡛ࡣ࣓ࢱࢤࣀ࣒ࡢᡭἲࢆ⏝࠸ࡓࢩ࣮ࢡ࢚ࣥࢫἲ࡟ࡼࡾ㸪⣅

౽࠿ࡽ┤᥋ᢳฟࡋࡓ RNA ࢆ౑ࡗ࡚ BRVB ࢤࣀ࣒㓄ิࢆỴᐃࡋࡓࠋ⣅౽୰࡟ࡣ ᐟ୺⏤᮶ࡸ⣽⳦࡞࡝ࡢ㑇ఏᏊࡀከࡃྵࡲࢀ࡚࠸ࡿࡇ࡜࠿ࡽ㸪ࡑࢀࡽ࡟⏤᮶ࡍࡿ1 ᮏ㙐RNAࢆ㝖ཤࡍࡿࡓࡵS1ࢾࢡࣞ࢔࣮ࢮࢆ౑⏝ࡋࡓࠋS1ࢾࢡࣞ࢔࣮ࢮࡣ࣓ࢱ ࢤࣀ࣒࡟ࡼࡿࢹ࢕࣮ࣉࢩ࣮ࢡ࢚ࣥࢫࢆ┠ⓗ࡜ࡋࡓゎᯒ⏝ࡢࣛ࢖ࣈ࣮ࣛࣜࢆస〇 ࡍࡿ㝿࡟᭷ຠ࡛࠶ࡿࡇ࡜ࡀド᫂ࡉࢀ࡚࠸ࡿ㸦78㸧ࠋࡇࡢᡭἲࢆᛂ⏝ࡍࡿࡇ࡜࡛

VP3㸪VP7ࠊNSP1㸪NSP3㸪NSP4 ࠾ࡼࡧ NSP5 ࡢ᏶඲㛗ࡢ CDS 㓄ิࢆᚓࡿ

ࡇ࡜ࡀ࡛ࡁࡓࠋ୍᪉㸪VP4㸪VP6㸪VP1㸪VP2 ࠾ࡼࡧ NSP2 ࡣ㒊ศⓗ࡞㓄ิࡋ

࠿ᚓࡿࡇ࡜ࡀ࡛ࡁ࡞࠿ࡗࡓࠋࡇࢀࡣ࢘࢖ࣝࢫ㑇ఏᏊࡢ㔞ࡀ༑ศ࡛࡞࠿ࡗࡓࡇ࡜

ࡸࢩ࣮ࢡ࢚ࣥࢫࡋ࡟ࡃ࠸㓄ิࢆ᭷ࡋ࡚࠸ࡿࡇ࡜࡟㉳ᅉࡋ࡚࠸ࡿ࡜⪃࠼ࡽࢀࡓࠋ ࡋ࠿ࡋࡇࢀࡽࡢࢭࢢ࣓ࣥࢺࡶ⣔⤫ᶞゎᯒ࡟౪ࡍࡿ࡟༑ศ࡞㑇ఏᏊ㛗ࡀᚓࡽࢀ㸪 11 ࢭࢢ࣓ࣥࢺ඲࡚࡟ࡘ࠸࡚ศᏊ⣔⤫ᶞゎᯒࢆ⾜࠺ࡇ࡜ࡀྍ⬟࡛࠶ࡗࡓࠋBRVB ࡢVP3㸪NSP3࠾ࡼࡧNSP4㑇ఏᏊࡢ᝟ሗࡣDDBJ/EMBL/GenBankࢹ࣮ࢱ࣋

࣮ࢫ࡟Ⓩ㘓ࡀ࡞࠿ࡗࡓࡇ࡜࠿ࡽ㸪ᮏ❶ࡢ◊✲࡛᫂ࡽ࠿࡟ࡉࢀࡓࡇࢀࡽࡢ㓄ิࡣ ୡ⏺ึࡢሗ࿌࡛࠶ࡿࠋ

ࣟࢱ࢘࢖ࣝࢫࡢ VP3 ࡣ N7-MTase ࡸ GTase/RTPase ࡞࡝ከᶵ⬟ᛶࡢ࢟ࣕࢵ

ࣆࣥࢢ㓝⣲ᶵ⬟ࢆ᭷ࡍࡿ㸦65㸧ࠋBRVBࡢVP3ࡢ࢔࣑ࣀ㓟㓄ิࡣ60.9–63.0%࡜㸪

௚ࡢRVB࡜ࡣప࠸┦ྠᛶࢆ♧ࡋࡓࡀ㸪N7-MTaseࡸGTase/RTPaseࢻ࣓࢖ࣥࢆ

☜ㄆࡍࡿࡇ࡜ࡀ࡛ࡁࡓࠋࡍ࡞ࢃࡕࠊ௒ᅇゎᯒࡋࡓBRVBᰴࡢVP3ࡶࠊ࢘࢖ࣝࢫ mRNA ྜᡂ᫬ࡢ࢟ࣕࢵࣆࣥࢢ཯ᛂ࡟㔜せ࡞ᶵ⬟ࢆ᭷ࡍࡿࡇ࡜ࡀᙉࡃ♧၀ࡉࢀࡓࠋ

ࣟࢱ࢘࢖ࣝࢫࡢNSP3ࡢNᮎ➃㡿ᇦࡣࣟࢱ࢘࢖ࣝࢫmRNAࡢ3’UTR࡟Ꮡᅾ ࡍࡿ≉ᐃࡢሷᇶ㓄ิ࡜⤖ྜࡍࡿ୍᪉㸪ࡑࡢ C ᮎ➃㡿ᇦࡣ⩻ヂ㛤ጞᅉᏊ࡜ࡋ࡚▱

50

ࡽࢀࡿeIF4G࡜⤖ྜࡍࡿ㸦16㸧ࠋNSP3ࡢࡇࢀࡽࡢ⤖ྜάᛶࡣ࢘࢖ࣝࢫmRNA

ࡢ⩻ヂ࡟ᙳ㡪ࡍࡿ㸦96㸧ࠋBRVBࡢNSP3࡟࠾ࡅࡿRNA⤖ྜ㡿ᇦࡣಖᏑࡉࢀ࡚

࠸ࡓࡀ㸪eIF4G ⤖ྜ㡿ᇦࡣ௚ࡢRVBࡸRVAࡢྠ㡿ᇦ࡜ࡣ࠿࡞ࡾ␗࡞ࡗ࡚࠸ࡓ (16)ࠋBLASTゎᯒ࡛ࡣBRVBࡢNSP3࡟᭱ࡶ┦ྠᛶࡀ㧗࠿ࡗࡓࡢࡣ㇜RVBࡢ RVB/Pig-wt/VNM/14250/2012/GXP[X] ᰴ࡛࠶ࡗࡓ(70)ࠋࡋ࠿ࡋࡇࡢᰴࡢ㓄ิ

㸦࢔ࢡࢭࢵࢩࣙࣥ␒ྕ KX362419㸧ࡣ 242 ሷᇶᑐ࡜▷ࡍࡂࡿࡓࡵ㸪⣔⤫ᶞゎᯒ

࡟ຍ࠼ࡿࡇ࡜ࡣ࡛ࡁ࡞࠿ࡗࡓ㸦ࢹ࣮ࢱᮍᥖ㍕㸧ࠋ

RVAࡢNSP4ࡣ᭱ึ㸪⭠⟶ࡢ⣽⬊ෆࡢ࢝ࣝࢩ࣒࢘ࣞ࣋ࣝࢆኚ໬ࡉࡏࡿࡇ࡜࡟

ࡼࡾஙࡢࡳ࣐࢘ࢫ࡟ᖺ㱋౫Ꮡᛶࡢୗ⑩⑕ࢆᘬࡁ㉳ࡇࡍ࢘࢖ࣝࢫᛶ࢚ࣥࢸࣟࢺ࢟

ࢩࣥ࡜ࡋ࡚ሗ࿌ࡉࢀࡓ㸦7,18,38㸧ࠋBRVB ࡢ NSP4 ࢔࣑ࣀ㓟㓄ิࡣࣄࢺ㸪㇜࠾

ࡼࡧࣛࢵࢺ RVB ࡢࡑࢀ࡜ࡣ┦ྠᛶࡀ 36.1–39.3%࡜ⴭࡋࡃప࠿ࡗࡓࠋRVA ࡢ NSP4 ࢔ ࣑ࣀ 㓟 㓄ิ 113–135 ఩࡟┦ᙜࡍࡿ࢚ࣥࢸࣟࢺ࢟ࢩࣥࢻ࣓࢖ࣥࡣ caveolin-1࡜⤖ྜࡍࡿ㸦6㸧ࠋࣄࢺRVB CAL-1ᰴࡢNSP࢔࣑ࣀ㓟㓄ิ106–127

఩ࡢ࢚ࣥࢸࣟࢺ࢟ࢩࣥᵝࢻ࣓࢖ࣥࡣ㸪RVAࡢ࢚ࣥࢸࣟࢺ࢟ࢩࣥࢻ࣓࢖ࣥ࡜27%

ࡢ┦ྠᛶ࡛࠶ࡿ㸦41㸧ࠋࡋ࠿ࡋ࡞ࡀࡽ௒ᅇゎᯒࡋࡓ BRVB ࡢ NSP4 ࡟ࡣ࢚ࣥࢸ

ࣟࢺ࢟ࢩࣥᵝࢻ࣓࢖ࣥ࡟┦ᙜࡍࡿ࢔࣑ࣀ㓟㓄ิࡣぢࡘ࠿ࡽ࡞࠿ࡗࡓࠋࡼࡗ࡚㸪

௒ᅇ᳨ฟࡉࢀࡓBRVBࡣNSP4ࡢ࢚ࣥࢸࣟࢺ࢟ࢩࣥάᛶ࡟౫Ꮡࡍࡿࡇ࡜࡞ࡃୗ

⑩ࢆⓎ⑕ࡉࡏࡿ⬟ຊࢆ᭷ࡍࡿࡇ࡜ࡀ♧၀ࡉࢀࡓࠋRVAࡢNSP4ࡢNᮎ➃㡿ᇦ࡟

࠶ࡿ⣙20࢔࣑ࣀ㓟࠿ࡽᵓᡂࡉࢀࡿ3࢝ᡤࡢ␯Ỉᛶࢻ࣓࢖ࣥh1-h3ࡢh2ࡣ㸪࢘

࢖ࣝࢫࡢᙧែᙧᡂࡍࡿ㝿࡟⭷㏱㐣ᛶࢆㄪᩚࡍࡿᙺ๭ࢆᢸࡗ࡚࠸ࡿ㸦41㸧ࠋSOSUI

ࣉࣟࢢ࣒ࣛ࡟ࡼࡾ㸪௒ᅇゎᯒࡋࡓ IS-1 ࠾ࡼࡧ᪤▱ࡢࣄࢺ RVB ࡢ ADRV ᰴ࡟

NSP4 ࡢ⭷㈏㏻㡿ᇦࡀᏑᅾࡍࡿࡇ࡜ࡀド᫂ࡉࢀࡓࠋࡲࡓ㸪∵ RVA ࡢ NCDV ᰴ

࡟࠾࠸࡚ࡶ MTHMM ࣉࣟࢢ࣒ࣛ࡟ࡼࡾ NSP4 ࡟⭷㈏㏻㡿ᇦࡀ♧ࡉࢀࡓࠋࡍ࡞

ࢃࡕ㸪SOSUI࡜MTHMMࣉࣟࢢ࣒ࣛࡢ㛫࡛᏶඲࡟୍⮴ࡋࡓ⤖ᯝࡣᚓࡽࢀ࡞࠿

51

ࡗࡓࡶࡢࡢ㸪㢮ఝࡋࡓ␯Ỉࣉࣟࣇ࢓࢖ࣝࣃࢱ࣮ࣥࡀBRVB㸪ࣄࢺRVB࠾ࡼࡧ∵

RVA࡟☜ㄆࡉࢀࡓࠋ

⌧ᅾ㸪RVB࡟ࡣ21✀ࡢVP7 G㑇ఏᏊᆺ㸦51㸧㸪13✀ࡢVP6 I㑇ఏᏊᆺ㸦54㸧㸪 7✀ࡢNSP1 A㑇ఏᏊᆺ㸦82㸧㸪4✀ࡢNSP2 N㑇ఏᏊᆺ㸦84㸧࠾ࡼࡧ6✀ࡢ NSP5 H㑇ఏᏊᆺࡀሗ࿌ࡉࢀ࡚࠸ࡿ(83,84)ࠋࡲࡓNSP1ࡣ࣌ࣉࢳࢻ1࠾ࡼࡧ࣌

ࣉࢳࢻ2࡟ศ࠿ࢀ࡚࠾ࡾ㸪ࡑࢀࡒࢀ7ࡘࡢA㑇ఏᏊᆺ࡟ศ㢮ࡉࢀ࡚࠸ࡿ㸦82㸧ࠋ ᮏ❶࡛ࡢ⣔⤫ᶞゎᯒࡢ⤖ᯝ㸪ᡃࡀᅜࡢBRVBࡣG3㸪I3㸪N2࠾ࡼࡧH6㑇ఏᏊ ᆺࢆ♧ࡍࡇ࡜ࡀࢃ࠿ࡗࡓࠋࡲࡓ㸪࢖ࣥࢻࡢBRVBࡣG5㸪I3㸪N2࠾ࡼࡧH6࡛

࠶ࡗࡓࡇ࡜ࡼࡾࠊBRVBࡢVP7࡟ࡣᑡ࡞ࡃ࡜ࡶG3㸪G5ࡢ2ࡘࡢ㑇ఏᏊᆺࡀᏑ ᅾࡍࡿࡇ࡜ࡀ᫂ࡽ࠿࡜࡞ࡗࡓࠋ

VP6 ࡢ࢔࣑ࣀ㓟㓄ิ࡟ᇶ࡙ࡃ⣔⤫ᶞ࡛ࡣ㸪ᡃࡀᅜࡢ BRVB ࡣ I3 㑇ఏᏊᆺ࡟

ศ㢮ࡉࢀࡓࡀ㸪࢖ࣥࢻࡢBRVB࡜ࡣᑡࡋ␗࡞ࡿศᒱࢆ♧ࡋࡓࠋࡋ࠿ࡋ㸪VP6ࡢ

⣔⤫ゎᯒ࡟⏝࠸ࡓ࢔࣑ࣀ㓟㓄ิࡣ 126 ࢔࣑ࣀ㓟࡛࠶ࡾ㸪㑇ఏᏊᆺูࢆ⾜࠺ࡢ࡟

ࡣ୙༑ศ࡜ᛮࢃࢀࡓࡓࡵ㸪⣔⤫ᶞࡢసᡂ࡟༑ศ࡞㸪ࡼࡾ㛗࠸࢔࣑ࣀ㓟㓄ิࢆᚓ

࡚෌ゎᯒࡍ࡭ࡁ࡜⪃࠼ࡽࢀࡓࠋ

NSP5࡟ࡘ࠸࡚ࡣ㸪ᡃࡀᅜࡢBRVBࡣ࢖ࣥࢻࡢBRVB࡜␗࡞ࡿࢡࣛࢫࢱ࣮ࢆ

ᙧᡂࡋࡓࡀ㸪᪤ሗ࡟࠶ࡿNSP5ᆺูࡢ࢝ࢵࢺ࢜ࣇ್ࡣሷᇶ࡛ࣞ࣋ࣝ78%࡛࠶ࡾ

㸦84㸧㸪ࡑࢀࢆᑡࡋୖᅇࡗࡓࡓࡵ㸪࢖ࣥࢻࡢBRVB࡜࡜ࡶ࡟H6㑇ఏᏊᆺ࡟ᒓࡍ

ࡿ࡜ุ᩿ࡉࢀࡓࠋNSP1ࡢ㑇ఏᏊᆺูࡣ㸪࣌ࣉࢳࢻ1࡛ࡣ࢔࣑ࣀ㓟࡛ࣞ࣋ࣝ72㸣

࠾ࡼࡧ࣌ࣉࢳࢻ2࡛ࡣ࢔࣑ࣀ㓟࡛ࣞ࣋ࣝ80㸣ࡢ࢝ࢵࢺ࢜ࣇ್ࡀタᐃࡉࢀ࡚࠸ࡿ

ࡀ㸦82㸧㸪ᡃࡀᅜࡢBRVBࡣ࣌ࣉࢳࢻ2࡛ࣄࢶࢪࡢRVBࡢA3㑇ఏᏊᆺ࡜67.9 㹼68.8%ࡢ┦ྠᛶࢆ♧ࡋࡓࠋࡋ࠿ࡋ㸪BRVB G-2006ᰴࡣࣄࢶࢪᰴ࡜࣌ࣉࢳࢻ1

࡛ 75.2%ࡢ┦ྠᛶࢆ♧ࡋࡓࠋࡇࡢࡇ࡜࠿ࡽ㸪ᡃࡀᅜࡢBRVB ࡣA3 㑇ఏᏊᆺ࡟

ศ㢮ࡉࢀࡿࡢ࠿ࡶࡋࢀ࡞࠸ࠋᮏ❶࡛ࡣࠊゎᯒࡋࡓBRVBᰴࡢVP3ࠊNSP3࠾ࡼ

52

ࡧ NSP4 ࡢ࢔࣑ࣀ㓟㓄ิ࡟ᇶ࡙ࡁࠊࡑࢀࡒࢀࢆ M1-M4ࠊT1-T4 ࠾ࡼࡧ E1-E4 ࡢ㑇ఏᏊᆺ࡟௬࡟ศ㢮ࡋࡓࠋࡑࡢୖ࡛ࠊྠศ㢮࡟ᇶ࡙ࡁࠊྛ㑇ఏᏊᆺࢆ༊ูࡍ

ࡿ┦ྠᛶࡢᇶ‽࡜࡞ࡿ࢝ࢵࢺ࢜ࣇ್ࢆୖグࡢྛ⺮ⓑ㉁࡟ࡘ࠸࡚ᥦၐࡋࡓࠋࡋ࠿

ࡋ⌧ᅾࡢ࡜ࡇࢁࢹ࣮ࢱ࣮࣋ࢫ࠿ࡽධᡭ࡛ࡁࡿVP3㸪NSP3࠾ࡼࡧ NSP4ࡢ㑇ఏ Ꮚ㓄ิࡢᩘࡣᑡ࡞ࡃ㸪࢝ࢵࢺ࢜ࣇ್ࢆᐃࡵࡿ࡟ࡣ୙༑ศ࡜⪃࠼ࡽࢀࡿࡓࡵ㸪௒

ᚋከࡃࡢ㑇ఏᏊࢹ࣮ࢱࡢ⵳✚ࡀᚲせ࡜⪃࠼ࡽࢀࡓࠋ

௒ᅇ㸪BRVB ࡢ VP3㸪NSP3࠾ࡼࡧ NSP4 ࡣ௚ࡢ RVBࡢ㑇ఏᏊ㓄ิ࡜ࡣ࠿

࡞ࡾ␗࡞ࡿࡇ࡜ࡀุ᫂ࡋࡓࠋࡋ࠿ࡋ࡞ࡀࡽ㸪ࡑࢀࡒࢀࡢ࢘࢖ࣝࢫ⺮ⓑ㉁࡟࠾ࡅ

ࡿ≉ᚩⓗ࡞ࣔࢳ࣮ࣇࡸศᏊ⏕≀Ꮫⓗ࡞ᛶ㉁ࡀ㸪RVB ࡢᰴ㛫࡟ಖᏑࡉࢀ࡚࠸ࡿࡇ

࡜ࡀド᫂ࡉࢀࡓࠋࡲࡓ㸪ᡃࡀᅜ࠾ࡼࡧ࢖ࣥࢻࡢ BRVB ࡣ, ඲࡚ࡢ㑇ఏᏊࢭࢢ࣓

ࣥࢺ࡟࠾࠸࡚௚ࡢRVB࡜ࡣ␗࡞ࡿ⣔⤫ࢢ࣮ࣝࣉࢆᙧᡂࡍࡿࡇ࡜ࡀࢃ࠿ࡗࡓࠋ௒

ᅇ㸪᪂ࡓ࡟ᚓࡽࢀࡓBRVBࡢ㑇ఏᏊ᝟ሗࡣ㸪RVBࡢศᏊ⏕≀Ꮫⓗ࡟ゎ᫂ࡍࡿୖ

࡛᭷⏝࡛࠶ࡿ࡜⪃࠼ࡽࢀࡓࠋ

53

Accession No. Strain Region

LC185660 RVB/Cow-wt/JPN/IS-1/1999/G3P[X] VP3 LC185661 RVB/Cow-wt/JPN/IS-2/2002/G3P[X] VP3 LC185662 RVB/Cow-wt/JPN/D-2002/2002/G3P[X] VP3 LC185663 RVB/Cow-wt/JPN/E-2002/2002/G3P[X] VP3 LC185664 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] VP3 LC185665 RVB/Cow-wt/JPN/E-2005/2005/G3P[X] VP3 LC185666 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] VP3 LC185667 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] VP3 LC185668 RVB/Cow-wt/JPN/P-2011/2011/G3P[X] VP3 LC185669 RVB/Cow-wt/JPN/IS-1/1999/G3P[X] NSP3 LC185670 RVB/Cow-wt/JPN/IS-2/2002/G3P[X] NSP3 LC185671 RVB/Cow-wt/JPN/D-2002/2002/G3P[X] NSP3 LC185672 RVB/Cow-wt/JPN/E-2002/2002/G3P[X] NSP3 LC185673 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] NSP3 LC185674 RVB/Cow-wt/JPN/E-2005/2005/G3P[X] NSP3 LC185675 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] NSP3 LC185676 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] NSP3 LC185677 RVB/Cow-wt/JPN/P-2011/2011/G3P[X] NSP3 LC185678 RVB/Cow-wt/JPN/IS-1/1999/G3P[X] NSP4 LC185679 RVB/Cow-wt/JPN/IS-2/2002/G3P[X] NSP4 LC185680 RVB/Cow-wt/JPN/D-2002/2002/G3P[X] NSP4 LC185681 RVB/Cow-wt/JPN/E-2002/2002/G3P[X] NSP4 LC185682 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] NSP4 LC185683 RVB/Cow-wt/JPN/E-2005/2005/G3P[X] NSP4 LC185684 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] NSP4 LC185685 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] NSP4 LC185686 RVB/Cow-wt/JPN/P-2011/2011/G3P[X] NSP4 LC185687 RVB/Cow-wt/JPN/IS-1/1999/G3P[X] VP7 LC185688 RVB/Cow-wt/JPN/IS-2/2002/G3P[X] VP7 LC185689 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] VP7 LC185690 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] VP7 LC185691 RVB/Cow-wt/JPN/P-2011/2011/G3P[X] VP7 LC185692 RVB/Cow-wt/JPN/IS-1/1999/G3P[X] VP4 LC185693 RVB/Cow-wt/JPN/IS-2/2002/G3P[X] VP4 LC185694 RVB/Cow-wt/JPN/D-2002/2002/G3P[X] VP4 LC185695 RVB/Cow-wt/JPN/E-2002/2002/G3P[X] VP4 LC185696 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] VP4 LC185697 RVB/Cow-wt/JPN/E-2005/2005/G3P[X] VP4 LC185698 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] VP4 LC185699 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] VP4 LC185700 RVB/Cow-wt/JPN/P-2011/2011/G3P[X] VP4 LC185701 RVB/Cow-wt/JPN/IS-1/1999/G3P[X] VP6 LC185702 RVB/Cow-wt/JPN/IS-2/2002/G3P[X] VP6 LC185703 RVB/Cow-wt/JPN/IS-1/1999/G3P[X] VP1 LC185704 RVB/Cow-wt/JPN/IS-2/2002/G3P[X] VP1 LC185705 RVB/Cow-wt/JPN/IS-2/2002/G3P[X] VP2 LC185706 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] VP2 LC185707 RVB/Cow-wt/JPN/IS-1/1999/G3P[X] NSP1 LC185708 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] NSP1 LC185709 RVB/Cow-wt/JPN/IS-1/1999/G3P[X] NSP2 LC185710 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] NSP2 LC185711 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] NSP5 LC185712 RVB/Cow-wt/JPN/P-2011/2011/G3P[X] NSP5

ᅗ⾲ ⾲

3-1 ▼ᕝ┴᳨࡛ฟࡉࢀࡓBRVBࡢྛ㑇ఏᏊࢭࢢ࣓ࣥࢺ࡜

DDBJ/EMBL/GenBank࢔ࢡࢭࢵࢩࣙࣥ␒ྕ

54

150 RVB/Cow-wt/JPN/IS-1/1999/G3P[X} M S K L I E F SD L G I E V T N R EQ L F K I SN N T S SY E T I R P S KD I ED Y I R E S SH Y V V I D KR R N E P F V E V F T Q L F P S S I VY N H K EGY K SG GC R H L L N N V L H V SN Y MH T Y C N S N T R N L A PD GW T I D K AD E F D D P I GD Y I L R SM VN D C SM EN K AQH R H K RVB/Cow-wt/JPN/IS-2/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/E-2002/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/P-2011/2011]/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉AI S䞉 䞉 䞉E䞉 䞉 䞉L䞉 䞉K V䞉 䞉FV䞉K䞉TG䞉 䞉 䞉 䞉 䞉QN䞉T䞉 䞉 䞉T䞉 䞉R䞉T䞉D E V S KL D T䞉 䞉 䞉TV I S N N E䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉D䞉I䞉 䞉 䞉 䞉SH䞉 䞉SL䞉R䞉D Q SW L䞉T䞉 䞉S V S EC䞉G L䞉 䞉H V䞉 䞉Q䞉I KR I䞉 䞉T䞉 䞉K Q T䞉G䞉I KN L RVB/Human-wt/CHN/WH-1/2002/G2P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉AI S䞉 䞉 䞉D䞉 䞉 䞉L䞉 䞉D䞉 䞉 䞉F K V䞉K䞉T A AK䞉 䞉 䞉G T䞉T䞉 䞉 䞉V LR䞉D䞉D E T ADLT䞉 䞉 䞉TV I F N C E䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉T䞉 䞉ID G䞉F S䞉G VN S I Q䞉W IR䞉 䞉 䞉V䞉 䞉C䞉R L䞉 䞉H V䞉 䞉 䞉 䞉I K L L䞉D S䞉 䞉S Q T VR EMN KM RVB/Human-wt/SEN/MRC-DPRU4680/2010/G2P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉AI S䞉 䞉 䞉D䞉 䞉 䞉L䞉 䞉D䞉 䞉 䞉F K IK䞉T A AK䞉 䞉 䞉G T䞉T䞉 䞉 䞉V LR䞉D䞉D E T ADLT䞉 䞉 䞉TV I F N C E䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉ID G䞉F S䞉G VN S I Q䞉W IR䞉 䞉I V䞉 䞉C䞉R L䞉 䞉H V䞉 䞉 䞉 䞉I K L L䞉D S䞉 䞉S Q T I R EMN KM RVB/Human-wt/IND/IDH-084/2007/G2P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉AI S䞉 䞉 䞉D䞉 䞉 䞉L䞉 䞉D䞉 䞉 䞉F K IK䞉T A AK䞉 䞉 䞉G T䞉T䞉 䞉 䞉V LR䞉D䞉D E T ADL A T䞉 䞉 䞉TV I F N C E䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉ID G䞉F S䞉G VN S I Q䞉W IR䞉 䞉I V䞉 䞉C䞉R L䞉 䞉H V䞉 䞉 䞉 䞉I K L L䞉D S䞉 䞉S Q T I R EMN KM RVB/Human-wt/IC-008/2008/G2P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉AI S䞉 䞉 䞉D䞉 䞉 䞉L䞉 䞉D䞉 䞉 䞉F K IK䞉T A AK䞉 䞉 䞉G T䞉T䞉 䞉 䞉V LR䞉D䞉D E T ADLT䞉 䞉 䞉TV I F N C E䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉ID G䞉F S䞉G VN S I Q䞉W IR䞉 䞉I V䞉 䞉C䞉R L䞉 䞉H V䞉 䞉 䞉 䞉I K L L䞉D S䞉 䞉S Q T I R EMN K V RVB/Human-wt/MMR/MMR-B1/2007/G2P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉AI S䞉 䞉 䞉D䞉 䞉 䞉L䞉 䞉D䞉 䞉 䞉F K IK䞉T A AK䞉 䞉 䞉G T䞉T䞉 䞉 䞉V LR䞉D䞉D E T ADLT䞉 䞉 䞉TV I F N C E䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉ID G䞉F S䞉G VN S I Q䞉W IR䞉 䞉I V䞉 䞉C䞉R L䞉 䞉H V䞉 䞉 䞉 䞉I K L L䞉D S䞉 䞉S Q T I R EMN KM RVB/Human-wt/IND/CAL-1/1998/G2P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉AI S䞉 䞉 䞉D䞉 䞉 䞉L䞉 䞉D䞉 䞉 䞉F K IK䞉T A AK䞉 䞉 䞉G T䞉T䞉 䞉 䞉V LR䞉D䞉D E T ADLT䞉 䞉 䞉TV I F N C E䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉ID G䞉F S䞉G VN S I Q䞉W IR䞉 䞉I V䞉 䞉C䞉R L䞉 䞉H V䞉 䞉 䞉 䞉I K L L䞉D S䞉 䞉S Q T I R EMN N M RVB/Human-wt/BGD/Bang373/2000/G2P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉AI S䞉 䞉 䞉D䞉 䞉 䞉L䞉 䞉D䞉 䞉 䞉F K IK䞉T V AK䞉 䞉 䞉G T䞉T䞉 䞉 䞉V LR䞉D䞉D E T ADLT䞉 䞉 䞉TV I F N C E䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉ID G䞉F S䞉G VN S I Q䞉W IR䞉 䞉I V䞉 䞉C䞉R L䞉 䞉H V䞉 䞉 䞉 䞉I K L L䞉D S䞉 䞉S Q T I R EMN KM RVB/Human-wt/GBD/Bang117/2002/G2P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉AI S䞉 䞉 䞉D䞉 䞉 䞉L䞉 䞉D䞉 䞉 䞉F K IK䞉T V AK䞉 䞉 䞉G T䞉T䞉 䞉 䞉V LR䞉D䞉D E T ADLT䞉 䞉 䞉TV I F N C E䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉ID G䞉F S䞉G VN S I Q䞉W IR䞉 䞉I V䞉 䞉C䞉R L䞉 䞉H V䞉 䞉 䞉 䞉I K L L䞉D S䞉 䞉S Q T I R EMN KM RVB/Rat-hhpUSA/IDIR/1984/G1P[X] 䞉 䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉 䞉A䞉I S䞉 䞉 䞉D䞉 䞉 䞉L䞉 䞉 䞉V䞉 䞉FI V䞉 䞉T T N䞉K䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉T䞉 䞉R䞉D䞉 䞉E AGDLT䞉 䞉 䞉TV I S D S E䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉D䞉 䞉 䞉SL GR D V Q S I T䞉T䞉 䞉N VL CKY䞉 䞉P V䞉 䞉 䞉V䞉K L I䞉 䞉E䞉 䞉C Q T VR䞉M䞉N S

* * * * * * . * * * * . * : : * * * : * * * : * * . . * * : : : . * : * : * * * * : * * * . : * . * * : . : : * * * : * : : * : * * * * * * * * * * * * * * : * . * . : * . . . * * * : . : . * : * * : * * : . : * : * * : . : . : 300 RVB/Cow-wt/JPN/IS-1/1999/G3P[X} PN G VY P K L L N I N E F F I K T L KR M I T P V S N VD F Q S Y H Y L N Q R R Q I G T L VR N T I F E L I AN N NWN VN Y I G P E F E S F R N I C E L L L N R N Y T G K I R F F T F N S S I QH D Y K KQ L S S F H R E K VDWD Y VH R Y R L K F EN A F C H L F L H H I I K SR SY S I I Y VN T RVB/Cow-wt/JPN/IS-2/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/E-2002/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/P-2011/2011]/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉 䞉 䞉D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X] K P䞉T䞉 䞉 䞉 䞉E G V䞉D V䞉KN F䞉T E I T䞉 䞉Q T H䞉 䞉 䞉 䞉 䞉 䞉N䞉M V䞉 䞉K䞉 䞉 䞉Y V䞉 䞉K䞉V䞉Y䞉 䞉R K Y䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KD䞉I L䞉 䞉T DE䞉 䞉 䞉 䞉F I T Y䞉LT䞉K K䞉N䞉R E K E R E Y N S Q䞉A䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉L F N䞉V AQQ䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉S RVB/Human-wt/CHN/WH-1/2002/G2P[X] K P䞉T䞉 䞉 䞉M E R V E䞉E䞉T N F䞉 䞉K䞉T䞉 䞉Q K E I䞉 䞉 䞉 䞉 䞉N䞉A V䞉 䞉 䞉 䞉 䞉 䞉Y V䞉 䞉K䞉VS䞉 䞉K S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K E䞉I N䞉 䞉T D䞉 䞉 䞉 䞉 䞉 䞉F V T䞉 䞉 䞉 䞉 䞉 䞉K S䞉N䞉 䞉H R V K E Y L N K䞉AG䞉N䞉 䞉 䞉 䞉M䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉 䞉 䞉I Y N䞉VM䞉Q䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 RVB/Human-wt/SEN/MRC-DPRU4680/2010/G2P[X] R P䞉T䞉 䞉R M E R V E䞉E䞉T N F䞉 䞉K䞉T䞉 䞉Q K E I䞉 䞉 䞉 䞉 䞉N䞉A V䞉 䞉 䞉 䞉 䞉 䞉Y V䞉 䞉K䞉VS䞉 䞉K S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K E䞉I N䞉 䞉T D K䞉 䞉 䞉 䞉 䞉F V T䞉 䞉 䞉 䞉 䞉 䞉K K䞉S䞉 䞉H H I R E Y L N E䞉AG䞉N䞉 䞉 䞉 䞉M䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉 䞉 䞉I Y N䞉V A䞉Q䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 RVB/Human-wt/IND/IDH-084/2007/G2P[X] R P䞉T䞉 䞉R M E R V E䞉E䞉T N F䞉 䞉K䞉T䞉 䞉Q K E I䞉 䞉 䞉 䞉 䞉N䞉A V䞉 䞉 䞉 䞉 䞉 䞉Y V䞉 䞉K䞉VS䞉 䞉K S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K E䞉I N䞉 䞉T D K䞉 䞉 䞉 䞉 䞉F V T䞉 䞉 䞉 䞉 䞉 䞉K K䞉S䞉 䞉H H I R E Y L N E䞉AG䞉N䞉 䞉 䞉 䞉M䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉 䞉 䞉I Y N䞉V A䞉Q䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 RVB/Human-wt/IC-008/2008/G2P[X] R P䞉T䞉 䞉R M E R V E䞉E䞉T N F䞉 䞉K䞉T䞉 䞉Q K E I䞉 䞉 䞉 䞉 䞉N䞉A V䞉 䞉 䞉 䞉 䞉 䞉Y V䞉 䞉K䞉VS䞉 䞉K S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K E䞉I N䞉 䞉T D K䞉 䞉 䞉 䞉 䞉F V T䞉 䞉 䞉 䞉 䞉 䞉K K䞉S䞉 䞉H H I R E Y L N E䞉AG䞉N䞉 䞉 䞉 䞉M䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉 䞉 䞉I Y N䞉V A䞉Q䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 RVB/Human-wt/MMR/MMR-B1/2007/G2P[X] R P䞉T䞉 䞉R M E R V E䞉E䞉T N F䞉 䞉K䞉T䞉 䞉Q K E I䞉 䞉 䞉 䞉 䞉N䞉A V䞉 䞉 䞉 䞉 䞉 䞉Y V䞉 䞉K䞉VS䞉 䞉K S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K E䞉I N䞉 䞉T D K䞉 䞉 䞉 䞉 䞉F V T䞉 䞉 䞉 䞉 䞉 䞉K K䞉S䞉 䞉H H I R E Y L N E䞉AG䞉N䞉 䞉 䞉 䞉M䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉 䞉 䞉I Y N䞉V A䞉Q䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 RVB/Human-wt/IND/CAL-1/1998/G2P[X] R P䞉T䞉 䞉R M E R V E䞉E䞉T N F䞉 䞉K䞉T䞉 䞉Q K E I䞉 䞉 䞉 䞉 䞉N䞉A V䞉 䞉 䞉 䞉 䞉 䞉Y V䞉 䞉K䞉VS䞉 䞉K S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K E䞉I N䞉 䞉I D K䞉 䞉 䞉 䞉 䞉F V T䞉 䞉 䞉 䞉 䞉 䞉K K䞉S䞉 䞉H Y I R E Y L N K䞉AG䞉N䞉 䞉 䞉 䞉M䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉 䞉 䞉L Y N䞉V A䞉Q䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 RVB/Human-wt/BGD/Bang373/2000/G2P[X] R P䞉T䞉 䞉R M E R V E䞉E䞉T N F䞉 䞉K䞉T䞉 䞉Q K E I䞉 䞉 䞉 䞉 䞉N䞉A V䞉 䞉 䞉 䞉 䞉 䞉Y V䞉 䞉K䞉VS䞉 䞉K S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K E䞉I N䞉 䞉T D K䞉 䞉 䞉 䞉 䞉F I T䞉 䞉 䞉 䞉 䞉 䞉K K䞉S䞉 䞉H H I R E Y L N K䞉AG䞉N䞉 䞉 䞉 䞉M䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉 䞉 䞉I Y N䞉V A䞉Q䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 RVB/Human-wt/GBD/Bang117/2002/G2P[X] R P䞉T䞉 䞉R M E R V E䞉E䞉T N F䞉 䞉K䞉T䞉 䞉Q K E I䞉 䞉 䞉 䞉 䞉N䞉A V䞉 䞉 䞉 䞉 䞉 䞉Y V䞉 䞉K䞉VS䞉 䞉K S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K E䞉I N䞉 䞉T D K䞉 䞉 䞉 䞉 䞉F I T䞉 䞉 䞉 䞉 䞉 䞉K K䞉S䞉 䞉H H I R E Y L N K䞉AG䞉N䞉 䞉 䞉 䞉M䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉 䞉 䞉I Y N䞉V A䞉Q䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 RVB/Rat-hhpUSA/IDIR/1984/G1P[X] R P䞉T䞉 䞉A V K S V䞉E K䞉T E F䞉 䞉H䞉T䞉 䞉Q T A I䞉 䞉 䞉 䞉 䞉N䞉L T䞉 䞉 䞉 䞉 䞉SY V T䞉K T V䞉R䞉 䞉S A䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K E䞉I K䞉 䞉T D K S䞉 䞉 䞉 䞉F I PY䞉 䞉 䞉T䞉R D䞉D䞉N K T E R QY S S S M A䞉 䞉N F䞉 䞉 䞉W䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉Q䞉L F R䞉V I KQ䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉A

* . * * . : : : : * : * . : * * . : * * * * * : * * * . * * . : . * : * : * * * . * * * * * * * * * * * * * * . : * * * : . : * * * * : : * : * : * . * . * . . : : * . . * : : * * * * * : * * * * * : * : : * : : . * * * * * * * * : 450 RVB/Cow-wt/JPN/IS-1/1999/G3P[X} L Y N I GNW T E A Y PW L N I R V VD H I P V I L N N S V V F G F M L S T N AC S F S V N VD S D R V VY S P K PY D D EN N VW T V S I L G EN I G V P SN E A E R I A A K KN G L P N Y I Y GG V K F D V E A L D F N Y V T VG L Y S L SN V I N S P E L I K A T L SY D H I F T F P T Y S EG DWR RVB/Cow-wt/JPN/IS-2/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/E-2002/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉P䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/P-2011/2011]/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X] 䞉 䞉 䞉V䞉 䞉 䞉V T䞉 䞉S䞉 䞉 䞉I N䞉 䞉 䞉 䞉L䞉T䞉Q K䞉 䞉 䞉I䞉 䞉 䞉L䞉 䞉S K D䞉 䞉 䞉 䞉 䞉 䞉T L䞉 䞉I䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉D S䞉Y䞉 䞉 䞉 䞉 䞉M䞉 䞉F䞉 䞉C L䞉D䞉E S䞉 䞉 䞉E䞉T䞉N䞉 䞉 䞉 䞉V F䞉 䞉 䞉P F T A䞉 䞉 䞉 䞉L䞉R I䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉S F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉D䞉N䞉 䞉 RVB/Human-wt/CHN/WH-1/2002/G2P[X] 䞉 䞉 䞉V䞉 䞉 䞉 䞉K䞉 䞉E䞉 䞉D I N䞉 䞉 䞉R L䞉 䞉 䞉Q K䞉 䞉I䞉 䞉 䞉Y䞉 䞉 䞉S K E䞉 䞉 䞉 䞉I䞉 䞉E䞉 䞉L䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉D S䞉M䞉 䞉 䞉 䞉 䞉M䞉D K LT VY䞉 䞉D D䞉 䞉 䞉 䞉 䞉A䞉N䞉 䞉 䞉 䞉V F䞉 䞉 䞉P F T A A䞉 䞉R䞉D䞉 䞉N I A䞉 䞉 䞉 䞉 䞉 䞉S V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉K䞉 䞉 䞉 䞉 RVB/Human-wt/SEN/MRC-DPRU4680/2010/G2P[X] 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉D I N䞉 䞉 䞉 䞉L䞉 䞉 䞉Q K䞉 䞉I I䞉 䞉Y䞉 䞉 䞉S K E䞉 䞉 䞉 䞉 䞉 䞉 䞉E䞉 䞉L䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉D S䞉 䞉 䞉 䞉 䞉 䞉 䞉M䞉N K LT T Y䞉 䞉D D䞉 䞉 䞉V䞉T䞉N䞉 䞉 䞉 䞉V F䞉 䞉 䞉P F T A A䞉 䞉K䞉D䞉I N I A䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉K䞉 䞉 䞉 䞉 RVB/Human-wt/IND/IDH-084/2007/G2P[X] 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉D I N䞉 䞉 䞉 䞉L䞉 䞉 䞉Q K䞉 䞉I I䞉 䞉Y䞉 䞉 䞉S K E䞉 䞉 䞉 䞉 䞉 䞉 䞉E䞉 䞉L䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉D S䞉 䞉 䞉 䞉 䞉 䞉 䞉M䞉N K LT T Y䞉 䞉D D䞉 䞉 䞉V䞉T䞉N䞉 䞉 䞉 䞉V F䞉 䞉 䞉P F T A A䞉 䞉K䞉D䞉I N I A䞉 䞉 䞉 䞉 䞉 䞉S V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉P䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉K䞉 䞉 䞉 䞉 RVB/Human-wt/IC-008/2008/G2P[X] 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉D I N䞉 䞉 䞉 䞉L䞉 䞉 䞉Q K䞉 䞉I I䞉 䞉Y䞉 䞉 䞉S K E䞉 䞉 䞉 䞉 䞉 䞉 䞉E䞉 䞉L䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉D S䞉 䞉 䞉 䞉 䞉 䞉 䞉M䞉N K LT T Y䞉 䞉D D䞉 䞉 䞉V䞉T䞉N䞉 䞉 䞉 䞉V F䞉 䞉 䞉P F T A A䞉 䞉K䞉D䞉I N I A䞉 䞉 䞉 䞉 䞉 䞉S V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉K䞉 䞉 䞉 䞉 RVB/Human-wt/MMR/MMR-B1/2007/G2P[X] 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉D I N䞉 䞉 䞉 䞉L䞉 䞉 䞉Q K䞉 䞉I I䞉 䞉Y䞉 䞉 䞉S K E䞉 䞉 䞉 䞉 䞉 䞉 䞉E䞉 䞉L䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉D S䞉 䞉 䞉 䞉 䞉 䞉 䞉M䞉N K LT T Y䞉 䞉D D䞉 䞉 䞉V䞉T䞉N䞉 䞉 䞉 䞉V F䞉 䞉 䞉P F T A A䞉 䞉K䞉D䞉I N I A䞉 䞉 䞉 䞉 䞉 䞉S V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉K䞉 䞉 䞉 䞉 RVB/Human-wt/IND/CAL-1/1998/G2P[X] 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉D I N䞉 䞉 䞉 䞉L䞉 䞉 䞉Q K䞉 䞉I I䞉 䞉Y䞉 䞉 䞉S K E䞉 䞉 䞉 䞉 䞉 䞉A E䞉 䞉L䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉D S䞉 䞉 䞉 䞉 䞉 䞉 䞉M䞉N K LT T Y䞉 䞉D D䞉 䞉 䞉V䞉T䞉N䞉 䞉 䞉 䞉V F䞉 䞉 䞉P F T A A䞉 䞉K䞉D䞉I N I A䞉 䞉 䞉 䞉 䞉 䞉S V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉K䞉 䞉 䞉 䞉 RVB/Human-wt/BGD/Bang373/2000/G2P[X] 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉D I N䞉 䞉 䞉 䞉L䞉 䞉 䞉Q K䞉 䞉I I䞉 䞉Y䞉 䞉 䞉S K E䞉 䞉 䞉 䞉 䞉 䞉A E䞉 䞉L䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉D S䞉 䞉 䞉 䞉 䞉 䞉 䞉M䞉N K LT T Y䞉 䞉D D䞉 䞉 䞉V䞉T䞉N䞉 䞉 䞉 䞉V F䞉 䞉 䞉P F T A A䞉 䞉K䞉D䞉I N I A䞉 䞉 䞉 䞉 䞉 䞉S V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉K䞉 䞉 䞉 䞉 RVB/Human-wt/GBD/Bang117/2002/G2P[X] 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉D I N䞉 䞉 䞉 䞉L䞉 䞉 䞉Q K䞉 䞉I I䞉 䞉Y䞉 䞉 䞉S K E䞉 䞉 䞉 䞉 䞉 䞉A E䞉 䞉L䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉D S䞉 䞉 䞉 䞉 䞉 䞉 䞉M䞉N K LT T Y䞉 䞉D D䞉 䞉 䞉V䞉T䞉N䞉 䞉 䞉 䞉V F䞉 䞉 䞉P F T A A䞉 䞉K䞉D䞉I N I A䞉 䞉 䞉 䞉 䞉 䞉S V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉K䞉 䞉 䞉 䞉 RVB/Rat-hhpUSA/IDIR/1984/G1P[X] 䞉 䞉 䞉V䞉 䞉 䞉V S G䞉P䞉 䞉Q I N䞉 䞉 䞉 䞉L䞉 䞉 䞉Q K䞉 䞉V V䞉 䞉 䞉L䞉S S S D䞉 䞉 䞉 䞉A䞉 䞉E T E T䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 䞉 䞉G䞉 䞉 䞉 䞉 䞉 䞉 䞉M䞉Q K䞉 䞉A K䞉 䞉D Q S䞉 䞉 䞉A䞉M䞉N䞉A䞉 䞉V F䞉 䞉 䞉P F T A E䞉 䞉Q䞉N䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉 䞉 䞉T V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉 䞉N䞉N䞉 䞉

* * * : * * * . . * * * : * . * * * . : * . * : * * : : * * : : * : : : * * * * : * . . : * * * * * . * * * * : . * * * * * * : * : : * : : * * * * * . * : * * : : * * * * . * * : : : . : . * * * * * * . * * * * * * * * . * * * : * * * * * * : * * . * : * * 600 RVB/Cow-wt/JPN/IS-1/1999/G3P[X} L E I E K T N K I F I T T QR Q F K F N DW I I D A K N L S L EMD T E V V S E S V F L Q L G KD R A F I SD R Y QH M V A F R F KQ KN Y Y SD R Y M SH L G I R Q P S I F N R D R F L T S R L S A Y I D R Q L T L N SD L S L I E KN H F SG F S GH L I A V E KY F H A L T Y T M S PMRW AN R A L RVB/Cow-wt/JPN/IS-2/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/E-2002/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/P-2011/2011]/G3P[X] 䞉 䞉I䞉 䞉 䞉D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉L N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉F䞉R䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X] D䞉R PH D䞉 䞉 䞉 䞉 䞉 䞉E K䞉L RE䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉N A䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉FPH䞉 䞉 䞉 䞉T䞉M F䞉 䞉L I S䞉 䞉 䞉 䞉 䞉E䞉F F䞉 䞉Q䞉 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉QY䞉S䞉 䞉 䞉N䞉 䞉M S䞉 䞉 䞉 䞉 䞉G I䞉 䞉T V䞉K䞉 䞉N䞉A䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉L䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Human-wt/CHN/WH-1/2002/G2P[X] D䞉R P T T D N䞉 䞉V䞉 䞉E K䞉L RE䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉F䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉M䞉 䞉 䞉L I S䞉 䞉 䞉 䞉 䞉K䞉F F䞉 䞉Q K F䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉I Y䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉S T䞉 䞉 䞉 䞉 䞉KQ䞉N䞉E䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉R䞉 䞉 䞉 RVB/Human-wt/SEN/MRC-DPRU4680/2010/G2P[X] D䞉R A T T D D䞉 䞉V䞉 䞉E K䞉L RE䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉F䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉M䞉 䞉 䞉L I S䞉 䞉 䞉 䞉 䞉K䞉F F䞉 䞉Q K F䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉I Y䞉S䞉 䞉 䞉N䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉S T䞉 䞉 䞉 䞉 䞉KQ䞉N䞉E䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 RVB/Human-wt/IND/IDH-084/2007/G2P[X] D䞉R A T T D D䞉 䞉V䞉 䞉E K䞉L RE䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉F䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉M䞉 䞉 䞉L I S䞉 䞉 䞉 䞉 䞉K䞉F F䞉 䞉Q K F䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉I Y䞉S䞉 䞉 䞉N䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉S T䞉 䞉 䞉 䞉 䞉KQ䞉N䞉E䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 RVB/Human-wt/IC-008/2008/G2P[X] D䞉R A T T D D䞉 䞉V䞉 䞉E K䞉L RE䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉F䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉M䞉 䞉 䞉L I S䞉 䞉 䞉 䞉 䞉K䞉F F䞉 䞉Q K F䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉I Y䞉S䞉 䞉 䞉N䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉S T䞉 䞉 䞉 䞉 䞉KQ䞉N䞉E䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 RVB/Human-wt/MMR/MMR-B1/2007/G2P[X] D䞉R A T T D D䞉 䞉V䞉 䞉E K䞉L RE䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉F䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉M䞉 䞉 䞉L I S䞉 䞉 䞉 䞉 䞉K䞉F F䞉 䞉Q K F䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉I Y䞉S䞉 䞉 䞉N䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉S T䞉 䞉 䞉 䞉 䞉KQ䞉N䞉E䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 RVB/Human-wt/IND/CAL-1/1998/G2P[X] D䞉R A T T D D䞉 䞉V䞉 䞉E K䞉L RE䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉F䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉M䞉 䞉 䞉L I S䞉 䞉 䞉 䞉 䞉K䞉F F䞉 䞉Q K F䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉I Y䞉S䞉 䞉 䞉N䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉S T䞉 䞉 䞉 䞉 䞉KQ䞉N䞉E䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 RVB/Human-wt/BGD/Bang373/2000/G2P[X] D䞉R A T T D D䞉 䞉V䞉 䞉E K䞉L RE䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉F䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉M䞉 䞉 䞉L I S䞉 䞉 䞉 䞉 䞉K䞉F F䞉 䞉Q K F䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉I Y䞉S䞉 䞉 䞉N䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉S T䞉 䞉 䞉 䞉 䞉KQ䞉N䞉E䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 RVB/Human-wt/GBD/Bang117/2002/G2P[X] D䞉R A T T D D䞉 䞉V䞉 䞉E K䞉L RE䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉F䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉M䞉 䞉 䞉L I S䞉 䞉 䞉 䞉 䞉K䞉F F䞉 䞉Q K F䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉I Y䞉S䞉 䞉 䞉N䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉S T䞉 䞉 䞉 䞉 䞉KQ䞉N䞉E䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 RVB/Rat-hhpUSA/IDIR/1984/G1P[X] D䞉R R MH D D䞉 䞉VG Q E K䞉L RE䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉F䞉SH䞉 䞉 䞉 䞉 䞉 䞉M䞉 䞉 䞉L I S䞉 䞉 䞉 䞉 䞉N䞉F F䞉 䞉Q K M䞉 䞉L䞉 䞉 䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉I Y䞉S䞉 䞉 䞉N䞉 䞉MN䞉 䞉 䞉 䞉 䞉N S䞉 䞉 䞉K䞉R Q䞉N䞉A䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉

* . : . * * : : : : . * : . * : * * : * * * * * * : * * : : . * * * * * : * * * * : * * * * * : * : * * : : : * * * * * : * : : * * . : * * : * * * * * * * . * * * : * : * * * . * * : . * * * * * . * * : : * : : * : * * * * * * * * * * : * * * * * * . * * * * * : * * * . * * * 750 RVB/Cow-wt/JPN/IS-1/1999/G3P[X} SD A I Y K K T D RW T N A A G ER H S I Q D F R N T Y AY L GD S I N P I F R SN L V T N KY A D K P T H C I Q L L S K KN H I T L Q L T T T S P D R AM P L V I K A V E G T R F KY L G R N K F G T MD V A L Y Q T D GM T QY E I V N I L R T MD I PC QQ T R P Y I MH M T I KD Q SD I P S T I V RVB/Cow-wt/JPN/IS-2/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/E-2002/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/P-2011/2011]/G3P[X] 䞉 䞉 䞉F䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉R䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉L䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉Q䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X] 䞉 䞉 䞉 䞉 䞉R䞉H䞉N F S䞉 䞉V䞉 䞉 䞉 䞉T L D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉L KY K䞉 䞉N䞉 䞉 䞉 䞉 䞉EKY A䞉M䞉MC GN䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KDS N L L T E I GN LH䞉L N I䞉 䞉 䞉 䞉K䞉S䞉 䞉S I N L䞉 䞉 䞉 䞉 䞉K E䞉I䞉 䞉K L S䞉L䞉 䞉S F䞉 䞉A䞉 䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉L䞉 䞉D KN䞉 䞉N I䞉 䞉 RVB/Human-wt/CHN/WH-1/2002/G2P[X] 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉V F S䞉 䞉ID䞉 䞉 䞉L H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉F䞉 䞉D䞉K F A䞉M䞉SC G V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KNQ IL Q E I GN䞉 䞉H䞉M V IK䞉 䞉 䞉 䞉D䞉 䞉S I K䞉I䞉 䞉 䞉 䞉K E䞉I䞉 䞉K䞉C䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T EC N V䞉D V V I RVB/Human-wt/SEN/MRC-DPRU4680/2010/G2P[X] 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉S F S䞉 䞉ID䞉 䞉 䞉L H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉F䞉 䞉D䞉K F A䞉TSC G A䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KNQ IL Q E I GN T I HM V IK䞉D K䞉D䞉 䞉N I K䞉I䞉 䞉 䞉 䞉KD䞉I䞉 䞉K䞉C䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉T E䞉N V䞉D V䞉I RVB/Human-wt/IND/IDH-084/2007/G2P[X] 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉S F S䞉 䞉ID䞉 䞉 䞉L H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉F䞉 䞉D䞉K F A䞉TSC G V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KNQ IL Q E I GN T I HM V IK䞉D K䞉D䞉 䞉N I K䞉I䞉 䞉 䞉 䞉KD䞉I䞉 䞉K䞉C䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉T E䞉N V䞉D V䞉I RVB/Human-wt/IC-008/2008/G2P[X] 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉S F S䞉 䞉ID䞉 䞉 䞉L H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉F䞉 䞉D䞉K F A䞉TSC G V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KNQ IL Q E I GN T I HM V IK䞉D K䞉D䞉 䞉N I K䞉V䞉 䞉 䞉 䞉KD䞉I䞉 䞉K䞉C䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉T E䞉N V䞉D V䞉I RVB/Human-wt/MMR/MMR-B1/2007/G2P[X] 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉S F S䞉 䞉ID䞉 䞉 䞉L H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉F䞉 䞉D䞉K F A䞉TSC G V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KNQ IL Q E I GN T I HM V IK䞉D K䞉D䞉 䞉N I K䞉I䞉 䞉 䞉 䞉KD䞉I䞉 䞉K䞉C䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T E䞉N V䞉D V䞉I RVB/Human-wt/IND/CAL-1/1998/G2P[X] 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉S F S䞉 䞉ID䞉 䞉 䞉L H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉F䞉 䞉D䞉K F A䞉TSC G V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KNQ IL Q E I GN T I HM V IK䞉D K䞉D䞉 䞉N I RI䞉 䞉 䞉 䞉K E䞉I䞉 䞉K䞉C䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉T ER N V䞉D V䞉I RVB/Human-wt/BGD/Bang373/2000/G2P[X] 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉S F S䞉 䞉ID䞉 䞉 䞉L H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉F䞉 䞉D䞉K F A䞉TSC G V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KNQ IL QD I GN T I HM V IK䞉D K䞉D䞉 䞉N I K䞉I䞉 䞉 䞉 䞉K E䞉I䞉 䞉K䞉C䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉T ER N V䞉D V䞉I RVB/Human-wt/GBD/Bang117/2002/G2P[X] 䞉 䞉 䞉 䞉 䞉S䞉 䞉 䞉S F S䞉 䞉ID䞉 䞉 䞉L H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉F䞉 䞉D䞉K F A䞉TSC G V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KNQ IL QD I GN T I HM V IK䞉D K䞉D䞉 䞉N I K䞉I䞉 䞉 䞉 䞉K E䞉I䞉 䞉K䞉C䞉 䞉 䞉 䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉V䞉T ER N V䞉D V䞉I RVB/Rat-hhpUSA/IDIR/1984/G1P[X] 䞉 䞉 䞉VS䞉 䞉 䞉I F S䞉 䞉IE䞉 䞉 䞉 䞉D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉T V䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉S䞉S䞉A L M䞉N C R T C䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉KDE V䞉 䞉T K V K S V I H䞉M K L䞉F䞉 䞉F䞉R䞉 䞉N VRA䞉 䞉 䞉 䞉T S䞉I䞉 䞉K䞉S䞉 䞉 䞉 䞉S F N䞉 䞉 䞉 䞉 䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉L R D D S K P䞉D V䞉L

* * * : * * * : : * * * : * * : : * * . * * * * * * * : : : * * * : . : * * . * * : * * . * . . . . * . . * * * * * * * * . . * . : : : : * : * * . . * . * * . : : * * * * . * * * : : * : * * : : : * . * * * * * * * * * * : : : : * . : : 763

RVB/Cow-wt/JPN/IS-1/1999/G3P[X} AN S N N V K V K E I R C RVB/Cow-wt/JPN/IS-2/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/E-2002/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2005/2005/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/D-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/G-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow-wt/JPN/P-2011/2011]/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X]H R KD I K I䞉 䞉 䞉KH RVB/Human-wt/CHN/WH-1/2002/G2P[X] I䞉R䞉D L T I䞉 䞉 䞉KR RVB/Human-wt/SEN/MRC-DPRU4680/2010/G2P[X] I䞉R䞉D L I I䞉 䞉 䞉KR RVB/Human-wt/IND/IDH-084/2007/G2P[X] I䞉R䞉D L I I䞉 䞉 䞉KR RVB/Human-wt/IC-008/2008/G2P[X] I䞉R䞉D L I I䞉 䞉 䞉KR RVB/Human-wt/MMR/MMR-B1/2007/G2P[X] I䞉R䞉D L I I䞉 䞉 䞉KR RVB/Human-wt/IND/CAL-1/1998/G2P[X] I䞉R䞉D L I I䞉 䞉 䞉KR RVB/Human-wt/BGD/Bang373/2000/G2P[X] I䞉R䞉D L I I䞉 䞉 䞉KR RVB/Human-wt/GBD/Bang117/2002/G2P[X] I䞉R䞉D L I I䞉 䞉 䞉KR RVB/Rat-hhpUSA/IDIR/1984/G1P[X] VE䞉D䞉Y S䞉 䞉 䞉KR : : : : : * * * .

GxxxE(S/T) motif

SGHΦ motif LΩxL(S/T)NxxN motif

ᅗ3-1 BRVB㸪ࣄࢺ࠾ࡼࡧ㇜ࣟࢱ࢘࢖ࣝࢫBࡢVP3࢔࣑ࣀ㓟㓄ิࡢẚ㍑

ࢻࢵࢺࡣୖẁ࡜ྠࡌࡇ࡜ࢆ♧ࡍ

55

RVB/Cow -w t/JPN/IS-1/1999/G3P[X] MA L NA V A S I L G SV L S K HE I DDQ L K I I E DF I HA MK DCGMML DNWRDA Y Y K L R I P K PMT G T T MA I Q L K NME T EV L RL RHE SWK NG E T MK DRL L Q S F DV G K K NGY T V L L P K T RNA EV V L L NS T RVB/Cow -w t/JPN/IS-2/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/D-2002/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/E-2002/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/D-2005/2005/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/E-2005/2005/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/D-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/G-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉Y䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/P-2011/2011/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉R䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X] 䞉 䞉 䞉 䞉V䞉V䞉V䞉 䞉 䞉 䞉 䞉D䞉F Q䞉 䞉NS K E䞉 䞉DE䞉K S䞉L E䞉 䞉 䞉I K I䞉E䞉 䞉 䞉 䞉 䞉 䞉Q E䞉 䞉 䞉 䞉 䞉 䞉S S䞉S䞉 䞉 䞉 䞉 䞉 䞉 䞉F䞉V䞉 䞉 䞉Q䞉 䞉NRA䞉NE䞉A G R䞉E䞉 䞉MS䞉 䞉E I S S䞉K䞉H䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉Q L䞉 䞉A䞉 䞉V RVB/Human-w t/CHN/WH-1/2002/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉RNCG䞉NE I S RV T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉A䞉 䞉 䞉 RVB/Human-w t/CHN/ADRV/1982/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉RNCG䞉NE I S RV T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉A䞉 䞉 䞉 RVB/Human-w t/IND/CAL/1998/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CG䞉NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/9222/2010/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NRA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/MMR/MMR-B1/2007/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/NIV-076222/2007/G3P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/IDH-084/2007/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/NIV-1048101/2010/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/IC-008/2008/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/SEN/MRC-DPRU4680/2010/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/NIV-0924341/2009/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/NIV-0948756/2009/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/BGD/Bang373/2000/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CG䞉NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/BGD/Bang117/2002/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/NIV-0632252/2006/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/NIV-957971/1995/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/NIV-005623/2000/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Human-w t/IND/NIV-04623/2004/G2P[X] 䞉 䞉 䞉D䞉L䞉 䞉 䞉 䞉E T䞉 䞉R䞉CGV NE I S R䞉T T K䞉E E䞉L D䞉 䞉 䞉 䞉KV䞉D䞉 䞉E䞉 䞉 䞉 䞉E䞉F䞉 䞉R䞉 䞉A䞉 䞉 䞉 䞉S䞉I M䞉F䞉I䞉N䞉Q䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉 䞉S䞉 䞉E I䞉N䞉D䞉H䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I L䞉V䞉 䞉 䞉 RVB/Rat-hhp/USA/IDIR/1984/G1P[X] L09722 䞉 䞉 䞉 䞉 䞉L䞉 䞉 䞉 䞉E T䞉 䞉V E CNV P䞉V A RV MNR䞉E D䞉L Y䞉 䞉 䞉 䞉R I E D䞉 䞉 䞉 䞉 䞉 䞉NE䞉L䞉 䞉R䞉 䞉S䞉 䞉 䞉S T䞉 䞉M䞉L䞉I䞉N䞉H䞉 䞉NKA䞉A E䞉A DR䞉F䞉 䞉MS䞉 䞉 䞉L S T䞉 䞉 䞉H䞉I䞉V䞉 䞉 䞉 䞉 䞉 䞉 䞉I I䞉A䞉 䞉S

* * * : . : . * : * : * * : : : : : . * * : . * * * : : * : * * : * * * : * : * * * : . * : * * * : * : * * * . * * : : * : * * * * : . * * : : . * . * : * : * : * * * * * * : : : * * * .

RVB/Cow -w t/JPN/IS-1/1999/G3P[X] I DMK L NP F P S DV V DDL I K K NS E L E NS I KV I Q - E EA NT K - - - MRY Q E E L I E E RDR I MS EMSQQ I T ML K K K I T F L RE NHK E RL E L I NQ HHE E E K K I Y K RE I S E L RL E NG T L RVB/Cow -w t/JPN/IS-2/2002/G3P[X] V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉- D䞉 䞉 䞉 䞉 䞉- - - I K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉Q䞉D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/D-2002/2002/G3P[X] V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉- D䞉T䞉 䞉 䞉- - - I K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉Q䞉D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/E-2002/2002/G3P[X] V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉- D䞉T䞉 䞉 䞉- - - I K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉Q䞉D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/D-2005/2005/G3P[X] V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉- D䞉 䞉 䞉 䞉 䞉- - - I K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉Q䞉D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/E-2005/2005/G3P[X] V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉- D䞉 䞉 䞉 䞉 䞉- - - I K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉Q䞉D䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/D-2006/2006/G3P[X] V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉- D䞉 䞉 䞉 䞉 䞉- - - I K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉Q䞉D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/G-2006/2006/G3P[X] V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉- D䞉 䞉 䞉 䞉 䞉- - - I K䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉G䞉N䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉I䞉 䞉Q䞉D䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Cow -w t/JPN/P-2011/2011/G3P[X] V䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉K䞉 䞉 䞉 䞉 䞉 䞉A䞉 䞉-䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉R䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X] S䞉F R䞉S NNH䞉 䞉 䞉I䞉 䞉 䞉 䞉 䞉R䞉D䞉 䞉T KQ䞉E E L K - CNQ I S S - - - - L E SG DF NMPY Q L QV V SA KV N䞉L S DL L KV CEG E CL K䞉Q䞉RL A䞉Q E䞉E T E D䞉I A A䞉 䞉T䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉L䞉 䞉 䞉 䞉NR䞉NT F S KGM GVB/Human-w t/CHN/WH-1/2002/G2P[X] S䞉L䞉 䞉S S䞉 䞉 䞉EA䞉A K䞉A E E䞉E KMRKQ䞉Q HL R -䞉QQ T S䞉S T T T L CEA L E SMT E R䞉K L I䞉RE K䞉T V R䞉MF L䞉CDK T NQ R䞉R䞉Q䞉Q I CE䞉EA T D䞉 䞉V䞉V䞉S䞉 䞉R䞉 䞉I L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/CHN/ADRV/1982/G2P[X] S䞉L䞉 䞉S S䞉 䞉 䞉EA䞉A K䞉A E E䞉E KMRKQ䞉E HL R -䞉QQ T S䞉S T A T L CEA L E NMT E R䞉K L I䞉RE K䞉T V R䞉MF L䞉CDK T NQ R䞉R䞉Q䞉Q I CE䞉EA T D䞉 䞉V䞉V䞉S䞉 䞉R䞉 䞉I L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/CAL/1998/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E K EQ䞉RKQ䞉 䞉HL R - DQQ T S䞉P T A I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/9222/2010/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉E䞉 䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T T I L CEA L E NMT E K䞉K L I䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/MMR/MMR-B1/2007/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉E K䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T T I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/NIV-076222/2007/G3P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉E K䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T T I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/IDH-084/2007/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉E K䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T T I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/NIV-1048101/2010/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉E K䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T T I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKML L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/IC-008/2008/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉E K䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T T I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 RVB/Human-w t/SEN/MRC-DPRU4680/2010/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉E K䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T T I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/NIV-0924341/2009/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉E K䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T T I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/NIV-0948756/2009/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉E K䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T T I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/BGD/Bang373/2000/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉EQ䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T A I L CEA L E DMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/BGD/Bang117/2002/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉I E E䞉EQ䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T A I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉K I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/NIV-0632252/2006/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉E K䞉RKQ䞉 䞉HV R -䞉QQ T S䞉P T A I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/NIV-957971/1995/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉EQ䞉HKQ䞉 䞉HL R -䞉QQ T S䞉P T A I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/NIV-005623/2000/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉EQ䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T A I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 GVB/Human-w t/IND/NIV-04623/2004/G2P[X] S䞉L䞉 䞉 䞉S䞉 䞉 䞉EA䞉T R䞉V E E䞉EQ䞉RKQ䞉 䞉HL R -䞉QQ T S䞉P T A I L CEA L E NMT E K䞉K L V䞉RE K䞉T V NKMF L䞉C䞉R I NQ R䞉 䞉 䞉Q䞉QMCE䞉EA T D䞉 䞉A䞉V䞉S䞉 䞉R䞉 䞉T L䞉M䞉 䞉 䞉 䞉Y R䞉QM䞉 䞉V䞉 䞉 RVB/Rat-hhp/USA/IDIR/1984/G1P[X] L09722 S EV䞉 䞉S DH䞉 䞉QA䞉 䞉R䞉V䞉 䞉 䞉HA䞉T E KV A Q L E SA KQ DDNS T I I L CKA V E DMS E K L䞉KA䞉K E S S DL E K L L L䞉CNRS NS R䞉 䞉RQ L E I CE䞉E T T D䞉 䞉A A V䞉A䞉 䞉K䞉 䞉T E䞉MR䞉 䞉 䞉Y R䞉Q I V䞉A Q䞉

. : . * . * : : * : : . : : : . . : : . : : . : . . : : : . * . * . : : * : : * : : * * * * * * . * * * * . . :

RVB/Cow -w t/JPN/IS-1/1999/G3P[X] NL T I S T L NKQ F DN MS L DHMK D I QV L I E H V S DV E E RCA K L S S E NV V L RK EQ DE H I RL I KG L A DKA NL QV D I S -RVB/Cow -w t/JPN/IS-2/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 -RVB/Cow -w t/JPN/D-2002/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 -RVB/Cow -w t/JPN/E-2002/2002/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 -RVB/Cow -w t/D-2005/2005/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉M - - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 -RVB/Cow -w t/JPN/E-2005/2005/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉M - - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 -RVB/Cow -w t/JPN/D-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 -RVB/Cow -w t/JPN/G-2006/2006/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 -RVB/Cow -w t/JPN/P-2011/2011/G3P[X] 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉- - - -䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 䞉 -RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X] L EY NNF䞉E T䞉HE RNV K I I SG L A A RA NL L V䞉D - - - S䞉S D䞉EA DDS K E K T E -GVB/Human-w t/CHN/WH-1/2002/G2P[X] REQ䞉DS I GQ E L䞉HS NR I V RG L A NRA G L V V䞉EV DSG NE T S D䞉S DA䞉 䞉HDDHE NS E䞉DL E DMMDPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML QMRE EME R䞉HE䞉ME I L NL NL DL D I -GVB/Human-w t/CHN/ADRV/1982/G2P[X] K EQ䞉DS I EQ E L䞉HS NR I V RG L A NRA G L V V䞉EV DSG NE T S DL S DD䞉 䞉HDDHE NS E䞉DL E DMMDPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML QMRE EME R䞉HE䞉ME I L NL NL DL D I -GVB/Human-w t/IND/CAL/1998/G2P[X] K EQ䞉DS I G䞉E L䞉HS NR I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NS E䞉DL E DMMNPG E䞉 䞉Q䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/IND/9222/2010/G2P[X] K EQ䞉DS I G䞉E L䞉Y S NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NG E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/MMR/MMR-B1/2007/G2P[X] K EQ䞉DS I GQ E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S EQ S DE䞉 䞉HDDHE NG E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME R䞉HE䞉ME I L NL NF DL DM -GVB/Human-w t/IND/NIV-076222/2007/G3P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S EQ S DE䞉 䞉HDDHE NG E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/IND/IDH-084/2007/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NG E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉I E I L NL NF DL DM -GVB/Human-w t/IND/NIV-1048101/2010/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NG E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/IND/IC-008/2008/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NG E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -RVB/Human-w t/SEN/MRC-DPRU4680/2010/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NG E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/IND/NIV-0924341/2009/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NG E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/IND/NIV-0948756/2009/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NG E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/BGD/Bang373/2000/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDNHE NS E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/BGD/Bang117/2002/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDNHE NS E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/IND/NIV-0632252/2006/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NS E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/IND/NIV-957971/1995/G2P[X] K EQ䞉DS I G䞉E L䞉HS NR I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NS E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -GVB/Human-w t/IND/NIV-005623/2000/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NRE䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉MG I L NL NF DL DM -GVB/Human-w t/IND/NIV-04623/2004/G2P[X] K EQ䞉DS I G䞉E L䞉HS NK I I RG L A NRA G L V V䞉EV DSG NE T S DQ S DE䞉 䞉HDDHE NS E䞉DL E DMMNPG E䞉 䞉R䞉P RGG E NP RRQA䞉ML EMRE EME RF HE䞉ME I L NL NF DL DM -RVB/Rat-hhp/USA/IDIR/1984/G1P[X] L09722 䞉DRNL V䞉E E䞉HG HHL DM I RG L A A RA G L V V䞉EG DSG F E T E NS DDD䞉S E䞉 䞉DDNG - F D I E DMME PG D䞉 䞉Q RP RGG E NP RRQA䞉L RQMDNE S E䞉G DE L D䞉DDM I E PG E DE QRP RGGRNP HQA RL I QMQDE L E NMI E DL HI L DE E F DNA L F L L

. : : : : : * . . : . . * *

H1 H2 H3 H4 H5 H6

S1 S2 H7 S3 H8 H8

ᅗ3-2 BRVB㸪ࣄࢺ࠾ࡼࡧ㇜ࣟࢱ࢘࢖ࣝࢫBࡢNSP3࢔࣑ࣀ㓟㓄ิࡢẚ㍑

ࢻࢵࢺࡣୖẁ࡜ྠࡌࡇ࡜ࢆ♧ࡍ

RNA⤖ྜࢻ࣓࢖ࣥ㸪Ș-helices (H1㹼H8)㸪ș-strands㸦S1㹼S3㸧࠾ࡼࡧ┿᰾⏕

≀ࡢ⩻ヂ㜼ᐖᅉᏊ4G⤖ྜ㡿ᇦࡣ㉥㸪㟷㸪⥳࠾ࡼࡧⲔⰍࡢᶓ⥺࡛♧ࡍ

56

110 RVB/Cow-wt/JPN/IS-1/1999/G3P[X} MAE----INESISNVLNVFETLQHETIHNVIKNTASSKLLTNCALSLLSIVTVALARNKVKLPITNKIRSNIRHIAESIVWKAETTIREIVNDVISKNDIFKDIIH RVB/Cow-wt/JPN/IS-2/2002/G3P[X]䞉䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Cow-wt/JPN/D-2002/2002/G3P[X]䞉䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉L䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Cow-wt/JPN/D-2005/2005/G3P[X]䞉䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Cow-wt/JPN/E-2005/2005/G3P[X]䞉䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Cow-wt/JPN/D-2006/2006/G3P[X]䞉䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Cow-wt/JPN/P-2011/2011/G3P[X]䞉䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉C䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X]䞉䞉䞉DSKMEQVIMDTLYDHI䞉䞉FAGNTT䞉䞉HKDTSSSPQI䞉䞉GAFTFGTLLTIILKKGINLLSLK䞉䞉䞉TYL䞉䞉MLAW䞉䞉䞉Q䞉䞉KDTERTLQQHEVIRLNT RVB/Human-wt/CHN/WH-1/2002/G2P[X]䞉䞉䞉SSDMQELFVQATYEEIKLASSVN䞉䞉QRETSSSSPQI䞉䞉GTMTLGALLTLIVKKGTKFLTVQ䞉䞉VAYL䞉䞉MLW䞉䞉NQVKTCDELHQRKVIEKLQC RVB/Human-wt/CHN/ADRV/1982/G2P[X]䞉䞉䞉SSDMQELFVQATYEEIKLASSVN䞉䞉QRETSSSSPQI䞉䞉GTMTLGALLTLIVKKGTKFLTVQ䞉䞉VAYL䞉䞉MVW䞉䞉NQVKTCDELHQRKVIEKLQC RVB/Human-wt/IND/NIV-04622/2004/G3P[X]䞉䞉䞉SNNMQDLFVQATYEEIKLAGSVN䞉䞉QRETSSSSPQI䞉䞉GAMTLGALLTLIVKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQIVKTCDELHQRKVIEKLQC RVB/Human-wt/SEN/MRC-DPRU4680/2010/G2P[X]䞉䞉䞉SSNMQDLFVQATYEEIKLAGSVN䞉䞉QRETSSSSPQI䞉䞉GVMTLGALLTLIVKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQIVKTCDELHQRKVIEKLQC RVB/Human-wt/IND/JBP/K/2008/G3P[X]䞉䞉䞉SSNMQDLFVQATYEEIKLAGSVN䞉䞉QRETSSSSPQI䞉䞉GAMTLGALLATLIVKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQIVKTCDELHQRKVIEKLQC RVB/Human-wt/MMR/MMR-B1/2007/G2P[X]䞉䞉䞉SSNMQDLFVQATYEEIKLAGSVN䞉䞉QRETSSSSPQI䞉䞉GAMTLGALLTLIVKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQIVKTCDELHQRKVIEKLQC RVB/Human-wt/IND/NIV-094456/2009/G2P[X]䞉䞉䞉SSNMQDLFVQATYEEIKLAGSVN䞉䞉QRETSSSSPQI䞉䞉GAMTLGALLTLIVKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQIVKTCDELHQRKVIEKLQC RVB/Human-wt/IND/IDH-084/2007/G2P[X]䞉䞉䞉SSNMQDLFVQATYEEIKLAGSVN䞉䞉QRETSSSSPQI䞉䞉GAMTLGALLTLIVKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQIVKTCDELHQRKVIEKLQC RVB/Human-wt/IND/CAL-1/1998/G2P[X]䞉䞉䞉SSNMQDLFVQATYEEIKLAGSVN䞉䞉QRETSSSSPQI䞉䞉GTMTLGALLTLIMKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQVKTCDELHQRKVIEKLQC RVB/Human-wt/BGD/Bang373/2000/G2P[X]䞉䞉䞉SSNMQDLFVQATYEEIKLAGSVN䞉䞉QRETSSSSPQI䞉䞉GTMTLGALLTLIVKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQVKTCDELHQRKVIEKLQC RVB/Human-wt/BGD/Bang117/2002/G2P[X]䞉䞉䞉SSNMQDLFVQATYEEIKLASSVN䞉䞉QRETSSSSPQI䞉䞉GTMTLGALLTLIVKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQVKTCDELHQRKVIEKLQC RVB/Human-wt/BGD/Bang544/2001/G2P[X]䞉䞉䞉SSNMQDLFVQATYEEIKLAGSVN䞉䞉QRETSSSSPQI䞉䞉GTMTLGALLTLIVKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQVKTCDELHQRKVIEKLQC RVB/Human-wt/IND/NIV-005625/2000/G2P[X]䞉䞉䞉SSNMQDLFVQATYEEIKLAGSVN䞉䞉QRETSSSSPQI䞉䞉GTMTLGALLTLIVKKGTKFLTMQ䞉䞉IVYL䞉䞉MLW䞉䞉NQVKTCDELHQRKVIEKLQC RVB/Rat-hhp/USA/IDIR/1984/G1P[X]䞉䞉䞉NNEMQQLFVQAAYEEIKLADSVD䞉䞉QRESSNSSPQ䞉䞉䞉䞉GALTVTALFTLMVKGTQFLQKFQ䞉䞉VVHLSMLW䞉䞉SQVKQLCDELNQHEVLQKLQC ***::.:*:..***.:.*..::..*:**.*::::::.:.:*:*.*:.**::::*:.**::.::.::..:::: 219 RVB/Cow-wt/JPN/IS-1/1999/G3P[X} LTE---EVERIKQSISKIKGMDVTKEIFDLCEQKMRAIDDKIDDVQKSCERRIKDYDWKIAALASQNLRPVEARVNIQNQTAEVINDDEMTKNNIVRQARTKLNSKRQL RVB/Cow-wt/JPN/IS-2/2002/G3P[X]䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉I䞉䞉HSV䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Cow-wt/JPN/D-2002/2002/G3P[X]䞉䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉I䞉䞉HSV䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Cow-wt/JPN/D-2005/2005/G3P[X]䞉䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉I䞉䞉HSV䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Cow-wt/JPN/E-2005/2005/G3P[X]䞉䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉I䞉䞉HSV䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Cow-wt/JPN/D-2006/2006/G3P[X]䞉䞉䞉---䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉I䞉䞉HSV䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉S䞉䞉䞉䞉䞉 RVB/Cow-wt/JPN/P-2011/2011/G3P[X]䞉䞉䞉---䞉䞉䞉䞉V䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉I䞉䞉HSV䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉䞉 RVB/Pig-tc/USA/LS00011_Ohio/XXXX/GXP[X]IDRILKDIKMYDEHVAL䞉䞉䞉䞉䞉LSM䞉䞉䞉䞉䞉VD䞉䞉NVKIE䞉䞉䞉䞉䞉䞉䞉R䞉䞉䞉䞉䞉䞉S䞉䞉TLHPIQQTAVKNETVAANEIKIVDEVAQTRNKPIKRAP-RRVB/Human-wt/CHN/WH-1/2002/G2P[X]DKMCSEIKLRYDMEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQLPHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/CHN/ADRV/1982/G2P[X]DKMCSEIKLRYDMEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉䞉䞉䞉ID䞉䞉SVKEER䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQLPHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/IND/NIV-04622/2004/G3P[X]DKMCLEIKLRYDIEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQFPTHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/SEN/MRC-DPRU4680/2010/G2P[X]DKMCLEIKLRYDIEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQFPHIDI䞉䞉HIDDAENQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/IND/JBP/K/2008/G3P[X]DKMCLEIKLRYDIEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQFPHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/MMR/MMR-B1/2007/G2P[X]DKMCLEIKLRYDIEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQFPHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/IND/NIV-094456/2009/G2P[X]DKMCLEIKLRYDIEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQFPHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/IND/IDH-084/2007/G2P[X]DKMCLEIKLRYDIEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQFPHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/IND/CAL-1/1998/G2P[X]DKMCLEIKLRYDIEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHTTQQFPPHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/BGD/Bang373/2000/G2P[X]DKMCLEIKLRYDIERFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQFPTHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/BGD/Bang117/2002/G2P[X]DKMCLEIKLRYDIEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQFPTHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/BGD/Bang544/2001/G2P[X]DKMCLEIKLRYDIEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQFPTHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Human-wt/IND/NIV-005625/2000/G2P[X]DKMCLEIKLRYDIEHFQ䞉䞉䞉䞉䞉䞉䞉LIQM䞉䞉R䞉䞉ID䞉䞉SVKEE䞉䞉䞉D䞉䞉䞉R䞉䞉䞉䞉䞉䞉䞉TTAHPTQQFPTHIDI䞉䞉HIDDAETQIIQQHMNK䞉䞉䞉V䞉䞉䞉䞉RNRRVB/Rat-hhp/USA/IDIR/1984/G1P[X]DQLCEDVKLRYNVEHI䞉䞉LISNLISLTR䞉䞉AD䞉䞉GRIRDVERSCDRKIRDYDWKLAALTANPVHQIAAHVDMISHENEAEAQDIQQHNK䞉䞉䞉VMSR-R:::*.:..:...**:::*::.::*.****.*:.:::.**:***.******:*::...*::::....:...*

Enterotoxin-like sequence of human RVB Hydrophobic region 2 (bovine RVB) (Tansmembrane region)

57

ᅗ3-4 SOSUI

ࣉࣟࢢ࣒ࣛ㸦A㸧࠾ࡼࡧ

TMHMM

ࣉࣟࢢ࣒ࣛ㸦B㸧ࢆ⏝࠸ࡓྛ✀

RVB

࠾ࡼࡧ

RVA

ᰴࡢ NSP4 ࢔࣑ࣀ㓟㓄ิࢆ⏝࠸ࡓぶỈᛶ/␯Ỉᛶࣉࣟࣇ࢓࢖ࣝ࠾ࡼࡧ⭷㈏㏻㡿

ᇦࡢ᥎ᐃ

58

⾲4-1

VP3

䛻䛚䛡䜛ሷᇶ㓄ิ䠄ୗ䠅䛚䜘䜃䜰䝭䝜㓟㓄ิ䠄ୖ䠅䛾┦ྠᛶ䠄䠂䠅

94.0-100 73.0

97.5-99.5 70.0

95.0-100 92.1-100

⾲4-2

NSP3䛻䛚䛡䜛ሷᇶ㓄ิ䠄ୗ䠅䛚䜘䜃䜰䝭䝜㓟㓄ิ䠄ୖ䠅䛾┦ྠᛶ䠄䠂䠅

92.3-100 74.0

92.8-99.8 78.0

96.2-100 96.1-100

4-3

NSP4䛻䛚䛡䜛ሷᇶ㓄ิ䠄ୗ䠅䛚䜘䜃䜰䝭䝜㓟㓄ิ䠄ୖ䠅䛾┦ྠᛶ䠄䠂䠅

93.6-100 69.0

90.5-99.8 70.0

96.8-100 96.5-100

䡲䡫䢀䡱䢈್

(%)

M4 68.8 68.7-69.0 63.3-63.9

M1 72.6-72.9 60.9-62.0 71.0

M2 68.3-68.9 61.6-63.0 72.0-72.3

M1 (

䝷䝑䝖

) M2 (

䝠䝖

) M3 (

) M4 (

)

E3 52.6-53.0 53.8-56.1 38.4-38.8

E2 68.6-69.7 37.9-39.3 55.7-57.5

M2 (䝠䝖) M3 (∵) M4 (㇜)

M3 63.7-63.8 63.9-64.9

T3

70.8 68.7-69.7

E4 61.4 63.9-65.6

䡲䡫䢀䡱䢈್

(%)

E1 65.8-67.1 36.1-37.4 47.5

M1 (䝷䝑䝖)

䡲䡫䢀䡱䢈್

M1 (䝷䝑䝖) M2 (䝠䝖) M3 (∵) M4 (㇜) (%)

59.7-61.3 57.7-60.3

70.3-72.3 56.4-56.9 62.3

61.2-62.1

57.9-58.2

55.3-56.2 T1

T2

T4 65.9-66.4

67.4-67.9 66.9-68.2 76.0-76.5

3-3

3-4

3-2

59

㸦㸣㸧

㸦㸣㸧

ᅗ3-5 VP3ࡢሷᇶ㓄ิ(A)࠾ࡼࡧ࢔࣑ࣀ㓟㓄ิ(B)࡟࠾ࡅࡿ┦ྠᛶศᕸᅗ

㸦┦ྠᛶ㢖ᗘ㸦㸣㸧 㸧

㸦┦ྠᛶ㢖ᗘ㸦㸣㸧 㸧

VP3

ሷᇶࣞ࣋ࣝࡢ࢝ࢵࢺ࢜ࣇ್㸸70㸣

VP3

࢔࣑ࣀ㓟ࣞ࣋ࣝࡢ࢝ࢵࢺ࢜ࣇ್㸸73㸣

60

㸦㸣㸧

㸦㸣㸧

ᅗ3-6 NSP3ࡢሷᇶ㓄ิ(A)࠾ࡼࡧ࢔࣑ࣀ㓟㓄ิ(B)࡟࠾ࡅࡿ┦ྠᛶศᕸᅗ

㸦┦ྠᛶ㢖ᗘ㸦㸣㸧 㸧

㸦┦ྠᛶ㢖ᗘ㸦㸣㸧 㸧

61

ᅗ3-7 NSP4ࡢሷᇶ㓄ิ(A)࠾ࡼࡧ࢔࣑ࣀ㓟㓄ิ(B)࡟࠾ࡅࡿ┦ྠᛶศᕸᅗ

㸦㸣㸧

㸦㸣㸧 㸦 ┦ ྠ ᛶ 㢖 ᗘ 㸦 㸣 㸧㸧

㸦 ┦ ྠ ᛶ 㢖 ᗘ 㸦 㸣 㸧㸧

62

ᅗ3-8 VP3 (a)㸪NSP3 (b)࠾ࡼࡧNSP4 (c)࡟࠾ࡅࡿ࢔࣑ࣀ㓟࡟ᇶ࡙ࡃศᏊ⣔⤫ᶞ

ࣈ࣮ࢶࢫࢺࣛࢵࣉ್ࡣྛࢢ࣮ࣝࣉࡢศᒱ࡟♧ࡍ

ࣟࢱ࢘࢖ࣝࢫBࡢ㑇ఏᏊᆺࢆྑഃ࡟♧ࡍ ᮏ❶࡛ゎᯒࢆ⾜ࡗࡓᰴࢆە࡛♧ࡍ

63 ᅗ3-8 㸦⥆ࡁ㸧

64

ᅗ3-9. VP7 (A)㸪VP4 (B)㸪VP6 (C)㸪VP1 (D)㸪VP2 (E)㸪NSP1 (F)㸪NSP2 (G)

࠾ࡼࡧNSP5 (H)࡟࠾ࡅࡿ࢔࣑ࣀ㓟࡟ᇶ࡙ࡃศᏊ⣔⤫ᶞ

ࣈ࣮ࢶࢫࢺࣛࢵࣉ್ࡣྛࢢ࣮ࣝࣉࡢศᒱ࡟♧ࡍ

ࣟࢱ࢘࢖ࣝࢫBࡢ㑇ఏᏊᆺࢆྑഃ࡟♧ࡍ ᮏ❶࡛ゎᯒࢆ⾜ࡗࡓᰴࢆە࡛♧ࡍ

65

3-5

VP7

䛾ሷᇶ㓄ิ䠄 ୗ䠅 䛚 䜘 䜃 䜰 䝭 䝜 㓟䠄 ୖ䠅 䛻 䛚 䛡 䜛 ┦ྠᛶ

G1 (

䝷䝑 䝖

)G2 (

䝠䝖

)G3 (

∵䚷᪥ᮏ䞉 䜰䝯 䝸 䜹

)G4 (

)G5 (

∵䚷 䜲 䞁 䝗

)G6 (

)G7 (

)G8 (

) G149.2-50.450.0-51.750.051.352.552.551.2 G256.9-57.359.5-63.260.7-61.661.6-62.863.2-63.664.1-64.563.2 G357.8-59.760.3-62.558.3-61.285.0-88.866.9-68.669.0-71.965.3-67.8 G455.761.4-62.861.6-64.360.763.662.059.5 G557.1-57.263.8-64.377.1-78.964.9-65.268.672.3-72.768.2 G658.165.2-65.866.0-68.263.468.5-68.6 G758.563.9-64.764.5-68.263.467.9-68.0 64.3-66.066.7-68.962.368.9-69.0

71.570.7 69.369.8 67.668.5G858.8

66

⾲ 3- 6 䚷 VP4 䛾ሷᇶ㓄ิ䠄 ୗ䠅 䛚 䜘 䜃 䜰 䝭 䝜 㓟䠄 ୖ䠅 䛻 䛚 䛡 䜛 ┦ྠᛶ 61. 0- 63. 7 61. 5- 64. 6 64. 7- 69. 1 64. 3- 68. 4

64. 7- 73. 3 ㇜

∵䠄 䜲 䞁 䝗 䠅 63. 1- 62. 7 63. 0- 64. 0 80. 1- 82. 0

64. 7- 65. 4 64. 0- 64. 8 90. 0- 91. 7 65. 2- 74. 0

60. 7- 65. 0 ∵䠄 ᪥ᮏ䠅

䝠䝖 63. 6- 64. 2 64. 5- 66. 0 63. 3- 64. 8

䝷䝑 䝖 64. 5- 65. 5 67. 4- 67. 6 65. 7- 66. 7 60. 0- 62. 9

∵䠄 ᪥ᮏ䠅 ∵䠄 䜲 䞁 䝗 䠅 ㇜ 䝷䝑 䝖 䝠 䝖

67

⾲ 㻟 㻙㻣䚷 㼂 㻼 㻢䛾 ሷ ᇶ 㓄 ิ 䠄ୗ 䠅䛚 䜘 䜃 䜰 䝭 䝜 㓟 䠄ୖ 䠅䛻 䛚 䛡 䜛 ┦ ྠ ᛶ I4 69. 1- 69. 8 71. 4- 74. 1 73. 5- 74. 9 73. 8- 76. 2

I3 ( 䜲䞁 䝗 ) 70. 4- 73. 8 69. 8- 71. 2 78. 8- 81. 0 86. 5- 88. 9

I3 ( ᪥ᮏ ) 69. 6- 71. 7 68. 8- 71. 4 92. 1- 95. 2 85. 7- 87. 3

I2 74. 3- 74. 9 82. 5- 85. 7 82. 5- 86. 5 79. 4- 82. 5

I1 89. 7- 92. 1 85. 7- 86. 5 86. 5- 88. 1 81. 8

I1 ( 䝷䝑 䝖 ) I2 ( 䝠䝖 ) I3 ( ᪥ᮏ䚷 ∵ ) I3 ( ∵䚷 䜲 䞁 䝗 ) I4 䠄㇜ 䠅

68

⾲㻟 㻙 㻤 䚷 㼂 㻼 㻝 ሷ ᇶ㓄ิ䠄 ୗ䠅 䛚 䜘 䜃 䜰 䝭 䝜 㓟䠄 ୖ䠅 䛻 䛚 䛡 䜛 ┦ྠᛶ 62. 2- 66. 5 61. 1- 70. 0 62. 8- 71. 0 65. 2- 74. 2

62. 4- 63. 1 65. 4- 66. 9 74. 2- 76. 6 69. 1- 86. 0 ㇜

60. 9- 61. 8 65. 2- 67. 6 85. 4- 87. 6 67. 4- 82. 6 ∵䠄 䜲 䞁 䝗 䠅 67. 2- 68. 5 68. 0- 70. 8 66. 9- 69. 7 69. 1- 75. 8 ∵䠄 ᪥ᮏ䠅

75. 3- 76. 4 65. 7- 68. 0 66. 9- 68. 0 69. 1- 76. 4 䝠䝖

䝷䝑 䝖

䝷䝑 䝖 䝠 䝖 ∵䠄 ᪥ᮏ䠅 ∵䠄 䜲 䞁 䝗 䠅 ㇜

69

⾲ 㻟 㻙㻥䚷 㼂 㻼 㻞䛾 ሷ ᇶ 㓄 ิ 䠄ୗ 䠅䛚 䜘 䜃 䜰 䝭 䝜 㓟 䠄ୖ 䠅䛻 䛚 䛡 䜛 ┦ ྠ ᛶ ㇜ 69. 5- 73. 7 69. 5- 74. 6 69. 5- 78. 1 69. 0- 77. 8

∵䠄 䜲 䞁 䝗 䠅 71. 5- 72. 0 68. 8- 71. 0 80. 5- 82. 0 72. 8- 92. 7

∵䠄 ᪥ᮏ䠅 70. 2- 70. 7 68. 8- 72. 0 94. 9- 97. 1 74. 3- 91. 2

䝠䝖 77. 3- 78. 5 75. 0- 76. 5 75. 7- 77. 2 71. 3- 86. 8

䝷䝑 䝖 90. 4- 91. 2 75 .7 76 .5 73. 5- 84. 6

䝷䝑 䝖 䝠 䝖 ∵䠄 ᪥ᮏ䠅 ∵䠄 䜲 䞁 䝗 䠅 ㇜

70

⾲ 㻟㻙㻝㻜䚷 㻺 㻿 㻼 㻝 㻌䝨 䝥 䝏 䝗 㻙㻞 䛾ሷᇶ㓄ิ䠄 ୗ䠅 䛚 䜘 䜃 䜰 䝭 䝜 㓟䠄 ୖ䠅 䛻 䛚 䛡 䜛 ┦ྠᛶ

70.8-77.3 A753.1-53.551.8-55.458.1-58.759.8-60.959.6-61.567.6-69.070.8-72.4

73.970.8-72.7 A653.453.5-54.562.260.1-61.060.9-61.369.7

70.0-70.859.0-59.356.2-56.555.6-56.8 A554.051.4-52.361.061.0-60.661.8-62.2 52.5-54.4 A452.2-52.652.3-52.967.5-68.2

51.9-52.451.1-52.168.6-69.270.5-72.856.8-57.554.4-55.3 66.5-67.156.554.451.2-51.9 A3 (

᪥ᮏ

) 43.3-45.2 A3 (

䝠䝒 䝆

)51.753.067.9–68.8%

64.5-64.740.6-40.943.3-44.643.3-44.044.6-45.243.7-44.3A2 41.543.3-44.643.7-44.247.447.445.2-45.8

A5 (

)A6 (

)A7 (

) A163.2-63.6

A1 (

䝷䝑 䝖

)A2 (

䝠䝖

)A3 (

䝠䝒 䝆

)A3 (

∵䚷 ᪥ᮏ

)A4 (

∵䚷 䜲 䞁 䝗

)

71

⾲ 㻟㻙㻝㻝䚷 㻺 㻿 㻼 㻞 䛾ሷᇶ㓄ิ䠄 ୗ䠅 䛚 䜘 䜃 䜰 䝭 䝜 㓟䠄 ୖ䠅 䛻 䛚 䛡 䜛 ┦ྠᛶ N4 70. 2- 73. 3 68. 3- 71. 2 65. 6- 68. 0 67. 6- 70. 4 64. 9- 66. 4

N365.6-67.674.8-76.073.373.8-74.3

69. 6- 70. 3

N2 (

)66.4-70.977.2-81.074.3-75.580.4-81.9

72. 5- 75. 4

N2 (

䜲䞁 䝗

)65.1-66.880.1-81.784.8-89.178.3-79.0

71. 7- 73. 9

N2 (

∵䚷 ᪥ᮏ

)65.4-69.089.1-92.086.2-89.179.7-81.9

71. 7- 74. 6

N167.4-73.268.1-73.267.4-73.967.4-69.6

71. 7- 81. 9

N4 (

)N1 (

䝷䝑 䝖 䚸 䝠 䝖

)N2 (

∵䚷 ᪥ᮏ

)N2 (

䜲䞁 䝗 䚷 ∵

)N2 (

)N3 (

)

72

⾲ 㻟 㻙㻝㻞䚷 㻺 㻿 㻼 㻡 䛾 ሷ ᇶ 㓄 ิ 䠄 ୗ 䠅 䛚 䜘 䜃 䜰 䝭 䝜 㓟 䠄 ୖ 䠅 䛻 䛚 䛡 䜛 ┦ ྠ ᛶ

82.0-83.2 H6 (

䝠䝒 䝆

)47.946.1-47.147.5-47.948.5-49.668.9-71.078.4-79.676.9-77.3

85.6-88.682.0-83.2 H6 (

∵䚷 䜲 䞁 䝗

)50.6-51.348.8-51.848.7-49.648.1-50.268.9-73.476.3-79.0

50.0-50.648.2-49.950.4-51.548.5-51.167.7-72.6

70.8-73.270.2-70.8 H6 (

∵䚷 ᪥ᮏ

)

H548.3-51.148.8-52.251.3-53.649.2-51.572.0-73.2 61.7-63.667.6-69.140.3-42.339.4-41.142.9-46.338.9-41.7 42.3-42.939.4-40.6 H464.8-65.9

H365.3-68.063.6-66.165.1-66.541.1-44.642.9-44.0 58.9-62.954.9-58.941.1-43.443.4-44.643.4-45.140.0-40.6

45.7-46.342.3 H268.0-68.8

H165.7-68.062.3-64.662.9-63.644.0-45.743.4-44.0 H5 (

)H6 (

∵䚷 ᪥ᮏ

)H6 (

∵䚷 䜲 䞁 䝗

)H6 (

䝠䝒 䝆

)H1 (

䝷䝑 䝖

)H2 (

䝠䝖

)H3 (

)H4 (

)

73

⥲ ᣓ

BRVB ࡣ㸪ᡃࡀᅜ࡛ࡣ BCoV࡟ḟࡄᡂ∵ࡢୗ⑩⑕ࡢཎᅉ࡜ࡉࢀ࡞ࡀࡽࡶ㸪࢘

࢖ࣝࢫᏛⓗ࡞▱ぢࡣ༑ศ࡟ሗ࿌ࡉࢀ࡚࡞࠿ࡗࡓࠋࡑࡇ࡛ᮏ◊✲࡛ࡣ㸪BRVB ࡢ 㜵␿࡟㈨ࡍࡿࡓࡵ㸪ࡇࢀࡲ࡛▱ࡽࢀ࡚࠸࡞࠿ࡗࡓBRVB ࡢ␿Ꮫ࠾ࡼࡧศᏊ⏕≀

Ꮫⓗᛶ≧࡟ࡘ࠸࡚◊✲ࢆᐇ᪋ࡋࡓࠋ

➨୍❶࡛ࡣ BRVBࡢᡂ∵࡬ࡢឤᰁ࡟ࡼࡾ㸪ୗ⑩⑕≧௨እ࡟ࡶἪங㔞పୗࢆᘬ ࡁ㉳ࡇࡍࡇ࡜ࢆୡ⏺࡛ึࡵ࡚᫂ࡽ࠿࡟ࡋࡓࠋࡉࡽ࡟㸪᳨ฟ࢘࢖ࣝࢫࡢ㑇ఏᏊゎ

ᯒ࡛ࡣ, RNA-PAGE࠿ࡽࢩ࣮ࣙࢺࢤࣀ࣒ࣃࢱ࣮ࣥࢆ♧ࡍࡇ࡜ࡀุ᫂ࡋ㸪ᡃࡀᅜ

᭱ึࡢBRVBᰴ࡛࠶ࡿ Nemuroᰴࡢࣟࣥࢢࢤࣀ࣒ࣃࢱ࣮ࣥ࡜␗࡞ࡿࡇ࡜࠿ࡽ㸪

ᡃࡀᅜ࡟ࡣᑡ࡞ࡃ࡜ࡶ཮᪉ࡢᰴࡀᏑᅾࡋ࡚࠸ࡿࡇ࡜ࢆ᫂ࡽ࠿࡟ࡋࡓࠋ

➨஧❶࡛ࡣ㸪኱つᶍ㓗㎰ᆅᖏ࡟࠾࠸࡚㸪3ᖺ㛫㝸࡛2ᅇࡢBRVB࡟ࡼࡿ㞟ᅋ ᛶୗ⑩ࡀⓎ⏕ࡋࡓ஦౛ࡼࡾ㸪BRVB࡟ࡼࡿୗ⑩⑕ࡢⓎ⏕ࡣ∵⩌ࡢච␿ࣞ࣋ࣝ࡟ᙳ 㡪ࡉࢀ㸪ྠࡌ㎰ሙෆ࡟࠾࠸࡚ࡶ∵⩌ࡢᢠయࣞ࣋ࣝࡀపୗࡍࡿ࡜෌ឤᰁࡀ㉳ࡇࡾ㸪 㞟ᅋᛶࡢୗ⑩⑕ࢆ⧞ࡾ㏉ࡍࡇ࡜ࢆ᫂ࡽ࠿࡟ࡋࡓࠋࡲࡓಶయ࡛ࣞ࣋ࣝࡣ㸪BRVB ࡣ෌ឤᰁࡍࡿࡀ㸪෌ឤᰁ࡛ࡣࡉ࡯࡝㔜⠜࡞⑕≧ࢆ♧ࡉ࡞࠸ࡇ࡜ࢆ᫂ࡽ࠿࡟ࡋࡓࠋ

➨୕❶࡛ࡣ㸪ࡇࢀࡲ࡛࡟▼ᕝ┴᳨࡛ฟࡉࢀࡓBRVB 9ᰴࡢ඲㑇ఏᏊ㓄ิࢆỴ ᐃࡋ㸪VP3㸪NSP4࠾ࡼࡧNSP5ࡀ௚ࡢࣟࢱ࢘࢖ࣝࢫ Bࡢ㑇ఏᏊ㓄ิ࡜ࡣ࠿࡞

ࡾ␗࡞ࡾ㸪᪂ࡋ࠸㑇ఏᏊᆺ࡛࠶ࡿࡇ࡜ࢆぢ࠸ࡔࡋࡓࠋࡲࡓᡃࡀᅜ࠾ࡼࡧ࢖ࣥࢻ ࡢBRVBࡣ඲࡚ࡢ㑇ఏᏊࢭࢢ࣓ࣥࢺ࡟࠾࠸࡚௚ࡢࣟࢱ࢘࢖ࣝࢫB࡜ࡣ␗࡞ࡗࡓ ࢢ࣮ࣝࣉࢆᙧᡂࡍࡿࡇ࡜ࡀุ᫂ࡋ㸪RVB ࢆศᏊ⏕≀Ꮫⓗ࡟ゎ᫂ࡍࡿୖ࡛᭷⏝࡞

᝟ሗࢆᚓࡿࡇ࡜ࡀ࡛ࡁࡓࠋ

ᮏ◊✲࡛ࡣ㸪BRVB ࡢឤᰁ஦౛ࢆ㏻ࡋ㸪ヲ⣽࡞␿Ꮫㄪᰝ࡜ศᏊ⏕≀Ꮫⓗゎᯒ

ࢆ⾜࠺ࡇ࡜࡟ࡼࡾ㸪᪂ࡓ࡞▱ぢࡀᩘከࡃᚓࡽࢀࡓ࡜ඹ࡟㸪⩌ච␿ࡢ㔜せᛶࡀ᫂

ࡽ࠿࡜࡞ࡗࡓࠋ

74

ᑗ᮶ⓗ࡟ࡣBRVB࡟ᑐࡍࡿ࣡ࢡࢳࣥ㛤Ⓨࡀ࡞ࡉࢀ㸪BRVB࡟ࡼࡿ⤒῭ⓗ⿕ᐖ ࡀ᭱ᑠ㝈࡟࡜࡝ࡵࡽࢀࡿࡼ࠺࡟࡞ࡿࡇ࡜ࢆᮇᚅࡋࡓ࠸ࠋ

75 ㅰ ㎡

ᮏㄽᩥసᡂ࡟࠶ࡓࡾࠊᩘከࡃࡢࡈᣦᑟ࡜ࡈຓゝࢆ㈷ࡾࡲࡋࡓ㸪ᮾி㎰ᕤ኱Ꮫ

㎰Ꮫ㒊㝃ᒓᅜ㝿ᐙ␆ឤᰁ⑕㜵␿◊✲ᩍ⫱ࢭࣥࢱ࣮ Ỉ㇂ဴஓᩍᤵ࡞ࡽࡧ࡟▼ᕝ

┴❧኱Ꮫ⏕≀㈨※⎔ቃᏛ㒊⏕⏘⛉Ꮫ⛉ື≀⦾ṪᏛ◊✲ᐊ 㛗஭ㄔᩍᤵ㸪ࡲࡓ◊

✲ࡢ㐙⾜࡟㝿ࡋ㸪㛗ᖺ࡟ࢃࡓࡾࡈᣦᑟࡈຓゝࢆ㈷ࡾࡲࡋࡓඖ㎰ᴗ࣭㣗ရ⏘ᴗᢏ

⾡⥲ྜ◊✲ᶵᵓ ື≀⾨⏕◊✲㒊㛛 ໭ᾏ㐨◊✲ᣐⅬ㛗 ᜏග⿱ඛ⏕࡟῝ࡃᚚ

♩⏦ࡋୖࡆࡲࡍࠋ

ࡲࡓᮏㄽᩥⲡ✏࡟࠶ࡓࡾ㸪᭷┈࡞ࡈຓゝ࡜ࡈᰯ㜀ࢆ㈷ࡾࡲࡋࡓ㸪ᖏᗈ␆⏘኱

Ꮫᛂ⏝⋇་Ꮫ⣔ ᑠᕝᬕᏊᩍᤵ㸪ᒾᡭ኱Ꮫᐇ㦂ື≀Ꮫ◊✲ᐊ ྂᕷ㐩ဢᩍᤵࠊ ᮾி㎰ᕤ኱Ꮫ㎰Ꮫ㒊ඹྠ⋇་Ꮫ⛉⋇་ఏᰁ⑓Ꮫ◊✲ᐊ ⓑ஭῟㈨ᩍᤵࠊᒱ㜧኱

Ꮫே⋇ඹ㏻ឤᰁ⑕Ꮫ◊✲ᐊ ఀ⸨┤ே෸ᩍᤵ࡟ᚰࡼࡾឤㅰ⏦ࡋୖࡆࡲࡍࠋ

ࡲࡓᮏ◊✲࡟ከ኱࡞ࡿࡈ⌮ゎ࡜ࡈᨭ᥼ࢆ㈷ࡾࡲࡋࡓ▼ᕝ┴㎰ᯘỈ⏘⥲ྜࢭࣥ

ࢱ࣮␆⏘ヨ㦂ሙ ᆏཱྀᨻಙሙ㛗㸪᪩ᕝ⿱஧๪ሙ㛗㸪ᅵᒇ࠸࡙ࡳᢏ⾡㛤Ⓨ㒊㛗㸪

໭ඖ㤶⳯Ꮚ୺௵ᢏᖌࡉࡽ࡟᳨ᰝᮦᩱࡢ᥇ྲྀ࡟ࡈ༠ຊ࠸ࡓࡔ࠸ࡓ▼ᕝ┴ෆࡢ㎰ᐙ ࡢⓙᵝ࡟ឤㅰ࠸ࡓࡋࡲࡍࠋ

᭱ᚋ࡟ᮏㄽᩥࡢసᡂ࡟࠶ࡓࡾ㸪ẕぶᴗࢆ௦⾜ࡋ࡚ࡃࢀࡓኵ࡜ᛂ᥼ࡋ࡚ࡃࢀࡓ ឡፉ࡟ឤㅰࡢពࢆ♧ࡋ㸪ㅰ㎡࡜ࡉࡏ࡚࠸ࡓࡔࡁࡲࡍࠋ

76 ཧ⪃ᩥ⊩

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