• 検索結果がありません。

ᐮ෭ᆅ࡟㐺ࡋࡓ࢖ࢳࢦࡢ⪏⑓ᛶရ✀ࡢ⫱ᡂ࡜⏘ᆅ࡬ࡢ㈉⊩

N/A
N/A
Protected

Academic year: 2021

シェア "ᐮ෭ᆅ࡟㐺ࡋࡓ࢖ࢳࢦࡢ⪏⑓ᛶရ✀ࡢ⫱ᡂ࡜⏘ᆅ࡬ࡢ㈉⊩"

Copied!
7
0
0

読み込み中.... (全文を見る)

全文

(1)

ᛂᛂ⏝◊✲ㄽᩥ

ᐮ෭ᆅ࡟㐺ࡋࡓ࢖ࢳࢦࡢ⪏⑓ᛶရ✀ࡢ⫱ᡂ࡜⏘ᆅ࡬ࡢ㈉⊩

㧗ᶫ᫓ᐿ 㸪ྜྷ⏣ᗣᚨ 㸪⚄⏣ၨ⮧ 㸪ྂᒇᘅග 㸪ᯇᮏ ໅ 㸪㧗஭㝯ḟ

1 ⛅⏣┴❧኱Ꮫ ⏕≀㈨※⛉Ꮫ㒊࢔ࢢࣜࣅࢪࢿࢫᏛ⛉

2 ⛅⏣┴❧኱Ꮫ ⏕≀㈨※⛉Ꮫ㒊⏕≀⏕⏘⛉Ꮫ⛉

3⛅⏣┴❧኱Ꮫ▷ᮇ኱Ꮫ㒊

4⛅⏣┴❧㎰ᴗ▷ᮇ኱Ꮫ

࢖ࢳࢦࡢ㯮ᩬ⑓ࡸⴎ㯤⑓࡟ࡼࡿ⿕ᐖࡀ⛅⏣┴࡞࡝ࡢᐮ෭ᆅࡢ࢖ࢳࢦ᱂ᇵ࡟࠾࠸࡚኱ࡁ࡞ၥ㢟࡜࡞ࡗ࡚࠸ࡓ㸬ࡑࡇ࡛㸪ⴭ⪅ࡽࡣ⫱✀

ࡢ❧ሙ࠿ࡽᮏၥ㢟ࢆゎỴࡋ㸪⏘ᆅ࡟㈉⊩ࡍࡿࡇ࡜ࢆヨࡳࡓ㸬㯮ᩬ⑓ࡣᐮ෭ᆅࡢ୺せရ✀ࡢ୍ࡘ࡛࠶ࡿ͂┒ᒸ

16

ྕ̓࡟≉␗ⓗ࡟Ⓨ⏕

ࡋ࡚࠸ࡓ⑓ᐖ࡛࠶ࡿࡀ㸪ᮏရ✀ࡣ኱ᯝ࡛㣗࿡ࡀࡼࡃ᪥ᣢࡕᛶ࡟ඃࢀ࡚࠸ࡓࡇ࡜࠿ࡽ㸪ࡇࢀࡽࡢㅖ≉ᛶࢆྍ⬟࡞㝈ࡾኻࢃ࡞࠸ࡼ࠺࡟

͂┒ᒸ

16

ྕ̓ࡢయ⣽⬊ኚ␗ಶయࡢ୰࠿ࡽ⪏⑓ᛶಶయࢆ㑅ᢤࡍࡿࡇ࡜࡜ࡋࡓ㸬ࡑࡢ⤖ᯝࠊⱼ㡬⤌⧊ࡢᇵ㣴࡟ࡼࡗ࡚సฟࡋࡓ

2237

ಶ యࡢ࢝ࣜࢡ᳜ࣟࣥ≀࠿ࡽ͂┒ᒸ

16

ྕ̓ࡢᯝᐇ≉ᛶ➼ࢆ᭷ࡋ㯮ᩬ⑓࡟ᑐࡋ࡚⪏ᛶ࡛࠶ࡿ 0$51㸪2 ࠾ࡼࡧ

3

3

⣔⤫ࢆ⋓ᚓ࡛ࡁࡓ㸬 ࡑࡢ୰࡛ẚ㍑ⓗ཰㔞ࡀ㧗࠿ࡗࡓ 0$52 ࢆ͂࢔࢟ࢱ࣮࣋ࣜ̓࡜࿨ྡࡋ࡚✀ⱑⓏ㘓ࡋᬑཬ࡟ྥࡅࡓ㸬ⴎ㯤⑓⪏ᛶရ✀ࡢ⫱ᡂ࡟࠶ࡓࡗ

࡚ࡣ᪤Ꮡရ✀࡜␗࡞ࡿ᪂ရ✀⫱ᡂࢆ┠ᶆ࡜ࡋ㸪 ͂Pajaro̓™࣮͂࣋ࣝࣝࢪࣗ̓ࡢ஺㞧ᐇ⏕

1006

ಶయ࠿ࡽⴎ㯤⑓⪏ᛶ࡛⏕⫱ࡢࡼ࠸

͂WB-A15̓ 㸪͂WB-B22̓㸪 ͂WB-B33̓ࡢ 3 ⣔⤫ࢆ㑅ᢤࡋࡓ㸬ࡇࢀࡽࡢ⣔⤫ࡢ࠺ࡕ཰㔞ࡀᏳᐃࡋ࡚㧗࠿ࡗࡓ͂WB-B33̓ࢆ͂ࡇࡲ

ࡕ࣮࣋ࣜ̓࡜ࡋ࡚✀ⱑⓏ㘓ࡋ⏘ᆅ࡟ᑟධࡋࡓ㸬ࡲࡓࠊ ͂WB-A15̓࡜͂WB-B22̓࡟ࡘ࠸࡚ࡣ✀ⱑⓏ㘓ࡋ࡞࠿ࡗࡓࡀ㸪⏕⏘⪅ࡸᕷሙ

࠿ࡽࡢせᮃࡶ࠶ࡗ࡚ࡑࢀࡒࢀ͂ࡣࡿࡳ̓㸪 ͂ࢽ࣮ࣗࡣࡿࡳ̓ࡢရ✀ྡ࡛୺࡟⛅⏣┴ࢆ୰ᚰ࡟᱂ᇵࡉࢀ࡚࠸ࡿ㸬

࣮࣮࢟࣡ࢻ㸸┒ᒸ

16

ྕ㸪࢖ࢳࢦ㯮ᩬ⑓㸪య⣽⬊ኚ␗㸪࢖ࢳࢦⴎ㯤⑓㸪ᅵተ⑓ᐖ㸪⪏⑓ᛶ

࢖ࢳࢦࡢ͂┒ᒸ16ྕ̓㸦㧗஭㸪᪋ᒣ㸪ⰼᒸཬࡧబ

⸨㸪1979㸧ࡣ኱ᯝ࡛㣗࿡ࡀࡼࡃ᪥ᣢࡕᛶ࡟ඃࢀ࡚࠸

ࡿࡇ࡜࠿ࡽ㸪1980ᖺ㹼1990ᖺ௦ึࡵ࡟࠿ࡅ࡚୺࡟໭

㛵ᮾ௨໭ࡢ㟢ᆅࡸ༙ಁᡂ᱂ᇵࡢ୺ຊရ✀࡛࠶ࡗࡓ 㸦㧗ᶫ㸪2001㸧㸬ࡋ࠿ࡋ㸪ᮏရ✀ࡣ6᭶㹼9᭶ୖ᪪ࡢ 㧗 ᮇ࡟ⱼⴥࡸࣛࣥࢼ࣮࡟Ⓨ⏕ࡍࡿ୙ṇᙧࡢ㯮〓Ⰽ ࡢ⑓ᐖࡀከⓎࡍࡿࡇ࡜࠿ࡽ㸪⏘ᆅ࡟࡜ࡗ࡚኱ࡁ࡞ၥ 㢟࡛࠶ࡗࡓ㸬

ࡑࡢᚋ㸪ࡇࡢ⑓ᐖࡣ㔝⳯Ⲕᴗヨ㦂ሙ┒ᒸᨭሙ㸦⌧

⊂❧⾜ᨻἲேᮾ໭㎰ᴗ◊✲ࢭࣥࢱ࣮㸧ࡢΏ㎶࡜ᱵᕝ 㸦1977㸧࡟ࡼࡗ࡚͂┒ᒸ ྕ̓࡟≉␗ⓗ࡟Ⓨ⏕ࡍࡿ

᪂⑓ᐖ㸪࢖ࢳࢦ㯮ᩬ⑓㸦Alternaria alternata strawberry pathotype㸧࡛࠶ࡿࡇ࡜ࡀ᫂ࡽ࠿࡟ࡉࢀࡓ㸬ᮏ⑓࡟㛵 ࡋ࡚ࡣ⑓⌮Ꮫⓗ࡞❧ሙ࠿ࡽΏ㎶㸪ᱵᕝཬࡧすᮧ

㸦1978㸧ࡢ࡯࠿࡟Maekawa et al㸬㸦1984㸧Namiki et al㸬1986aNamikiet al㸬㸦1986b㸧ࡀᮏ⑓⳦ࡢ ỴᐃᅉᏊ࡛࠶ࡿᐟ୺≉␗ⓗẘ⣲㸦AFẘ⣲㸧ࡸ⳦ࡢᙧ

ែⓗ≉ᚩ࡟ࡘ࠸࡚⢭ຊⓗ࡟◊✲ࢆ⾜ࡗࡓ㸬ࡋ࠿ࡋ㸪 ᮏ⑓࡟ᑐࡋ࡚≉ຠᛶࢆ♧ࡍ⸆๣ࡀ࡞࠿ࡗࡓࡇ࡜࠿ࡽ㸪

⏕⏘⪅࠿ࡽࡣ͂┒ᒸ16ྕ̓࡜ྠ➼ࡢ⏕⫱࠾ࡼࡧᯝᐇ

≉ᛶࢆᣢࡕ㸪ᮏ⑓⪏ᛶࡢ᪂ရ✀⫱ᡂࡀᙉࡃᮃࡲࢀࡓ㸬 ࡇࢀࡽࡢࡇ࡜࠿ࡽ㸪ⴭ⪅ࡽࡣ࢖ࢳࢦ㯮ᩬ⑓࡟⪏ᛶ࡛㸪

㈐௵ⴭ⪅㐃⤡ඛ㸸㧗ᶫ᫓ᐿ ࠛ995-0203 ᒣᙧ┴ᮧᒣᕷ኱Ꮠ኱ᵞ

418 ๓බ❧኱Ꮫἲே⛅⏣┴❧኱Ꮫ⏕≀㈨※⛉Ꮫ㒊࢔ࢢࣜࣅࢪࢿࢫᏛ⛉

E-mail: [email protected]

2006

3

᭶ ⛅⏣┴❧኱Ꮫࡢ㛤ㅮ࡟క࠸㛢Ꮫ

1999

4

᭶⛅⏣┴❧኱Ꮫ▷ᮇ኱Ꮫ㒊࡟ྡ⛠ኚ᭦

(2)

͂┒ᒸ16ྕ̓ࡢඃⰋ࡞ᯝᐇ≉ᛶࡸ཰㔞ᛶࢆኻࢃࡎ࡟

᪂ရ✀ࢆ⫱ᡂࡍࡿࡇ࡜ࢆ┠ⓗ࡟1986ᖺ࠿ࡽ◊✲࡟

╔ᡭࡋࡓ㸬

෗┿1 ͂┒ᒸ16 ྕ̓ࡢ」ⴥ࡟Ⓨ⏕ࡋ ࡓ࢖ࢳࢦ㯮ᩬ⑓㸬

࢖ࢳࢦ㯮ᩬ⑓ࡢ⪏⑓ᛶရ✀⫱ᡂࡢᡂᯝࡀฟጞࡵࡓ 㡭㸦㧗ᶫ㸪㧗஭ཬࡧᯇᮏ㸪1988㸧㸪⛅⏣┴࡞࡝ࡢ㟢ᆅ ᱂ᇵࡸ༙ಁᡂ᱂ᇵ࢖ࢳࢦ࡟᪂ࡓ࡞ၥ㢟ࡀⓎ⏕ࡋࡓ㸬

࢖ࢳࢦࡢ⑓ᐖࡢ୰࡛ࡶ㞴⑓ࡢ୍ࡘ࡜ࡉࢀ࡚⏕⏘⪅࡟

ᜍࢀࡽࢀ࡚࠸ࡿ࢖ࢳࢦⴎ㯤⑓㸦Fusarium oxysporum f㸬 sp㸬fragariae㸧࡟ࡼࡿ⿕ᐖ࡛࠶ࡿ㸬ᮏ⑓ࡣ࢖ࢳࢦ㯮

ᩬ⑓࡜ࡣ␗࡞ࡾ⑓ཎ⳦ࡀᆅୗ㒊࡟Ꮡᅾࡍࡿࡓࡵ㜵㝖 ࡀ㞴ࡋࡃ㸪ࢡࣟࣝࣆࢡࣜࣥ๣࡞࡝ࡢ๻⸆ࢆᅵተ࡟℺

ὀࡋ࡚ẅ⳦ࡍࡿ௨እ࡟᪉ἲࡀ࡞࠸㸬ࡋ࠿ࡶ㸪ࡇࡢ᪉ ἲࢆ⏝࠸࡚ࡶࡑࡢຠᯝࡣ࠶ࡲࡾᮇᚅ࡛ࡁ࡞࠸ሙྜࡀ ከࡃ㸪⩣ᖺࡲࡓࡣ⩣ࠎᖺ࡟⃭Ⓨࡍࡿࢣ࣮ࢫࡀከࠎ࠶

ࡿ㸬

≉࡟㸪ᮏ⑓ࡢ⿕ᐖࡣ⛅⏣┴‮ἑᕷ⛅ࣀᐑ㸦ᪧ㞝຾

⏫⛅ࣀᐑ㸧ࡢ㟢ᆅ࢖ࢳࢦ⏘ᆅ࡟࠾࠸࡚⏒኱࡛࠶ࡗࡓ㸬

⛅ࣀᐑࡣᙜ᫬30ᖺ௨ୖ⥆࠸࡚࠸ࡿ᪥ᮏ࡛ࡶ᭷ᩘࡢ ኟ࡝ࡾࡢ࢖ࢳࢦ⏘ᆅ࡛㸪6㹼7᭶࡟ᮾி㒔୰ኸ༺኎ᕷ ሙ࡟༨ࡵࡿฟⲴ㔞ࡣ඲ᅜ1㹼3఩࡛࠶ࡗࡓ㸦㧗ᶫ㸪 2006㸧ࡇ࡜࠿ࡽ㸪⏘ᆅࡢࡳ࡞ࡽࡎ┴ෆእࡢ༺኎ᕷሙ

➼࠿ࡽࡶ⪏⑓ᛶရ✀ࡢ⫱ᡂࢆษᮃࡍࡿኌࡀᴟࡵ࡚㧗

࠿ࡗࡓ㸬ࡑࡇ࡛㸪ⴭ⪅ࡽࡣᏛෆ࡛◊✲ࢢ࣮ࣝࣉࢆ❧

ࡕୖࡆ㸪⛅⏣┴ࢆ୰ᚰ࡜ࡋࡓᐮ෭ᆅࡢ㟢ᆅ᱂ᇵࡸ༙

ಁᡂ᱂ᇵ࡟ྥࡃရ✀⫱ᡂࢆ┠ⓗ࡟1991ᖺ࠿ࡽ◊✲

ࢆ㛤ጞࡋࡓ㸬

௨ୖ㏙࡭ࡓ ࡘࡢ◊✲ࡣ㸪࠸ࡎࢀࡶ㛗ᮇ㛫ࢆせࡋ ࡓࡀ⪏⑓ᛶရ✀ࡢ㛤Ⓨࡲ࡛⤖ࡧࡘࡅࡿࡇ࡜ࡀ࡛ࡁ㸪

⛅⏣┴ࡢ࢖ࢳࢦ᱂ᇵࡢࡳ࡞ࡽࡎ≉࡟໭᪥ᮏࡢ࢖ࢳࢦ ᱂ᇵ࡟኱ࡁࡃ㈉⊩ࡋࡓࡢ࡛ࡣ࡞࠸࠿࡜⪃࠼࡚࠸ࡿ㸬 ௨ୗ࡟ࡑࡢᴫせࢆ㏙࡭࡚ࡳࡓ࠸㸬

࢖ࢳࢦ㯮ᩬ⑓⪏⑓ᛶရ✀ࡢ⫱ᡂ

࢖ࢳࢦ㯮ᩬ⑓࡜ࢃࡀᅜ࡟࠾ࡅࡿ⨯⑓ᛶရ✀

ⴭ⪅ࡽࡀ㯮ᩬ⑓㸦෗┿1㸧࡟⪏ᛶࢆࡶࡘရ✀⫱ᡂ

࡟ྲྀࡾ⤌ࢇࡔᙜึ㸪ࢃࡀᅜ࡛⫱ᡂࡉࢀࡓရ✀࡛ࡣ͂┒

ᒸ16ྕ̓ࡢ࡯࠿࡟⨯⑓ᛶရ✀ࡣᏑᅾࡋ࡞࠿ࡗࡓ㸬ᮏ

⑓ࡢⓎ⏕ࡣᖹᆒẼ ࡀ15Υ௨ୖ࡟࡞ࡿ6᭶ึࡵ㡭࠿

ࡽቑຍࡍࡿ㸬ࡇࡢ᫬ᮇ࠿ࡽ9᭶୰᪪㡭ࡲ࡛ࡣ࢖ࢳࢦ ⱑࡢ⫱ⱑᮇ࡟࠶ࡓࡿࡇ࡜࠿ࡽⱑ㉁ࡢపୗࢆࡲࡡࡃࡔ

ࡅ࡛࡞ࡃ㸪ᮏᅡ࡟᳜࠼௜ࡅࡿᰴᩘࡢ☜ಖࡀฟ᮶࡞ࡃ

࡞ࡿሙྜࡀከ࠸㸬

࢖ࢳࢦ㯮ᩬ⑓࡟ࡘ࠸࡚ࡣ㸪᭱㏆ࡢ◊✲㸦୕⃝㸪す ᕝཬࡧᰰ᳃㸪2012㸧ࡸ⏕⏘⪅࠿ࡽࡢ᝟ሗ㸦⚾ಙ㸧࡟

ࡼࡗ࡚㸪᪂ࡓ࡟ᅄᏘᡂࡾᛶࡢ࢚͂ࢵࢳ࢚ࢫ-138̓࡜

͂ࡍࡎ࠶࠿ࡡ̓ࡢ2ရ✀ࡀ⨯⑓ᛶ࡛࠶ࡿࡇ࡜ࡀ☜ㄆ ࡉࢀ࡚࠸ࡿࡀ㸪࠸ࡎࢀࡢရ✀ࡶ͂┒ᒸ16ྕ̓ࡀ⫱ᡂ ࡉࢀࡓᚋࡢရ✀࡛㸪⫱✀ࡢ㐣⛬࡛͂┒ᒸ16ྕ̓ࡀ⫱

✀⣲ᮦ࡜ࡋ࡚฼⏝ࡉࢀ࡚࠸ࡿࡇ࡜ࡀ᫂ࡽ࠿࡟࡞ࡗ࡚

࠸ࡿ㸬

⪏⑓ᛶᰴࡢసฟἲ

࢖ࢳࢦࡢ⪏⑓ᛶရ✀ࡢ⫱ᡂ࡟࠶ࡓࡗ࡚ࡣ㸪௚స≀

࡜ྠᵝ࡟⑓Ẽ࡟ᙉ࠸ရ✀࣭⣔⤫➼ࡢ㑇ఏᏊࢆ஺㞧⫱

✀ἲ࡟ࡼࡗ࡚ᑟධࡍࡿࡢࡀ୍⯡ⓗ࡛࠶ࡿ㸬ࡋ࠿ࡋ㸪 ࡇࡢ⫱✀ἲ࡛ࡣ⪏⑓ᛶࡣ௜୚ࡉࢀࡓ࡜ࡋ࡚ࡶ஺㓄ぶ ࡢඃⰋ࡞ᯝᐇᙧ㉁➼ࢆኻ࠺ࡇ࡜ࡶ㠀ᖖ࡟ከ࠸㸬ࡑࡇ

࡛㸪ⴭ⪅ࡽࡣ͂┒ᒸ16ྕ̓ࡢඃⰋ࡞⏕⫱ࡸᯝᐇ≉ᛶ

ࢆኻࢃࡎ࡟㯮ᩬ⑓࡟⪏ᛶࢆࡶࡘᰴࢆ⫱ᡂࡍࡿࡇ࡜ࢆ

ࡡࡽ࠸࡜ࡋ㸪஺㞧⫱✀ἲࡼࡾࡶኚ␗ࡢᖜࡀᑠࡉ࠸࡜

ࡉࢀ࡚࠸ࡿయ⣽⬊ኚ␗㸦Somaclonal variation㸧฼⏝ࡢ

⫱✀ἲࢆ⏝࠸ࡿࡇ࡜࡜ࡋࡓ㸬

ᮏ◊✲࡛ࡣ͂┒ᒸ16ྕ̓ࡢⱼ㡬ᇵ㣴࡟ࡼࡗ࡚෌ศ

໬ಶయ㸦࢝ࣜࢡ᳜ࣟࣥ≀㸧ࢆసฟࡋ㸪ࡑࢀࡽࡢಶయ

࡟͂┒ᒸ16ྕ̓ࡢ㯮ᩬ⑓⨯⑓ᰴ࠿ࡽศ㞳ࡋࡓ㯮ᩬ

⑓⳦㸦ศ㞳⳦ᰴ㸸OH-5㸪⬊Ꮚ⃰ᗘ㸸5×105ಶml㸧ࢆ

᥋✀ࡋ⪏⑓ᛶಶయࢆ㑅ᢤࡋࡓ㸬

⪏⑓ᛶᰴࡢㅖ≉ᛶ

㑅ᢤ࡟ࡣ2237ಶయࡢ࢝ࣜࢡ᳜ࣟࣥ≀ࢆ⏝࠸ࡓࡀ㸪ࡑ

ࢀࡽࡢ࠺ࡕࡢ3ಶయࡢࡳ㯮ᩬ⑓⳦᥋✀ᚋࡶ඲ࡃⓎ⑓

ࡋ࡞࠿ࡗࡓ㸬ࡑࡇ࡛㸪ࡇࢀࡽࡢ3ಶయ࡟͂M16-AR1̓㸪

(3)

͂M16-AR2̓㸪͂M16-AR3̓ࡢ⣔⤫ྡࢆ௜ࡋᮏⴥ5 㹼6ᯛࡢᡂⱑࡲ࡛⏕⫱ࡉࡏ㸪෌ᗘᮾ໭࣭໭ᾏ㐨ࡢྛ

ᆅᇦ࡛᱂ᇵࡉࢀ࡚࠸ࡿ͂┒ᒸ16ྕ̓ࡢ⨯⑓ᰴ࠿ࡽศ 㞳ࡋࡓ㯮ᩬ⑓⳦ࢆ᥋✀ࡋࡓࡀ㸪࠸ࡎࢀࡢ⣔⤫࡟ࡶⓎ

⑓ࡣ඲ࡃㄆࡵࡽࢀ࡞࠿ࡗࡓ㸦⾲1㸧㸬

ࡉࡽ࡟㸪ࡇࢀࡽ3⣔⤫ࡀ࡝ࡢ⛬ᗘ͂┒ᒸ16ྕ̓ࡢ ᙧែࡸᯝᐇ≉ᛶ➼ࢆഛ࠼࡚࠸ࡿ࠿ࢆㄪᰝࡍࡿࡓࡵ㸪

ྛ⣔⤫࡜ぶရ✀ࡢ͂┒ᒸ16ྕ̓ࢆẚ㍑᳨ウࡋࡓ㸬ࡑ ࡢ⤖ᯝ㸪͂M16-AR̓ࡢ3⣔⤫ࡢⲡጼࡣ͂┒ᒸ16ྕ̓

࡜ྠᵝ❧ᛶ࡛㸪ᑠⴥࡶ඾ᆺⓗ࡞ᮼ≧ᙧ࡛࠶ࡗࡓ㸬ࡲ

ࡓ㸪㏻ᖖࡢ࢖ࢳࢦࡢ」ⴥࡀ3ᯛ࡛࠶ࡿࡢ࡟ᑐࡋ͂┒

ᒸ16ྕ̓࡟ࡣ4㹼5ᯛࡢ」ⴥࡢⓎ⏕ࡀ㧗㢖ᗘ࡛ㄆࡵ

ࡽࢀࡿࡀ㸪͂M16-AR̓ࡢྛ⣔⤫ࡶࡇࡢ≉ᛶࢆഛ࠼࡚

࠸ࡓ㸦෗┿2㸧㸬㛤ⰼᮇ㸪཰✭ᮇ㸪ᯝᙧ㸪ᯝ⓶Ⰽ㸪ᯝ

⫗Ⰽ➼࡟ࡘ࠸࡚ࡶ͂┒ᒸ16ྕ̓࡜ࡢ㛫࡟㐪࠸ࡣࡳࡽ

ࢀ࡞࠿ࡗࡓ㸦TakahashiTakai and Matsumoto1992㸧㸬

⪏⑓ᛶᰴࡢ㑇ఏᏊᆺ࠾ࡼࡧ⪏⑓ᛶ࡟ಀࢃࡿ㑇ఏ

͂M16-AR̓⣔⤫ࡣᐇ⏝ရ✀࡜ࡋ࡚ࡢࡳ࡞ࡽࡎ⫱

✀⣲ᮦ࡜ࡋ࡚฼⏝ࡉࢀࡿྍ⬟ᛶࡀ㧗࠸࡜⪃࠼ࡽࢀࡓ㸬 ࡋ࠿ࡋ㸪ᮏ⣔⤫࡟ࡘ࠸࡚ࡣ஺㞧⫱✀ἲ࡛ᚓࡽࢀࡓ᳜

≀య࡜␗࡞ࡾ㸪ኚ␗࡛ᚓࡽࢀࡓ᭷⏝ᙧ㉁ࡀ㑇ఏⓗ࡞

ࡶࡢ࠿࡝࠺࠿᫂ࡽ࠿࡟ࡋ࡚࠾ࡃᚲせࡀ࠶ࡿ㸬ࡑࡇ࡛㸪

͂M16-AR̓ 3⣔⤫ࡢ㑇ఏゎᯒࢆ⾜ࡗࡓ㸬ゎᯒ⤖ᯝ ࡣ3⣔⤫࡜ࡶ࡯ࡰྠᵝ࡛࠶ࡗࡓࡢ࡛㸪ᮏሗ࿌࡛ࡣ

͂M16-AR2̓ࡢ⤖ᯝ࡟ࡘ࠸࡚♧ࡋࡓ㸦⾲2㸧㸬ࡇࢀࡽ

ࡢゎᯒ⤖ᯝ࠿ࡽࡶ᫂ࡽ࠿࡞ࡼ࠺࡟3⣔⤫࡜ࡶ࢖ࢳࢦ 㯮ᩬ⑓࡟㛵ࡋ࡚⨯⑓ᛶ࡛࠶ࡿ͂┒ᒸ16ྕ̓ࡢ㑇ఏᏊ ᆺ㸦ඃᛶ࡬ࢸࣟ㸧ࡀ⪏⑓ᛶရ✀ࡢ㑇ఏᏊᆺ㸦ຎᛶ࣍

ࣔ㸧࡟ኚ␗ࡋ࡚࠸ࡿ࡜⪃࠼ࡽࢀࡓ㸬ࡲࡓ㸪͂M16-AR̓

⣔⤫ࡢ⪏⑓ᛶࡣᚋ௦࡟㑇ఏࡍࡿࡇ࡜ࡶ᫂ࡽ࠿࡜࡞ࡗ ࡓ㸬

⪏⑓ᛶ⣔⤫ࡢ᱂ᇵ㐺ᛂᛶ

͂M16-AR̓ࡢ ⣔⤫ࡣᙧែⓗ≉ᛶ㸪ᯝᐇ≉ᛶ㸪 㑇ఏⓗ≉ᛶ࠿ࡽࡳ࡚͂┒ᒸ16ྕ̓࡟௦ࢃࡿᐇ⏝ရ✀

࡜ࡋ࡚ࡢྍ⬟ᛶࡀ㧗࠸ࡇ࡜࠿ࡽ㸪͂┒ᒸ16ྕ̓ࡀ⏝

࠸ࡽࢀ࡚࠸ࡿ㟢ᆅ᱂ᇵ࠾ࡼࡧᐮ෭ᆅᆺ༙ಁᡂ᱂ᇵ 㸦௨ୗ㸪ప ࢝ࢵࢺ᱂ᇵ㸧࡟࠾ࡅࡿ᱂ᇵ㐺ᛂᛶ࡟ࡘ

࠸᳨࡚ウࡋࡓ㸬ࡑࡢ⤖ᯝ㸪୧సᆺ࡟࠾࠸࡚࢖ࢳࢦ㯮

ᩬ⑓ࡢⓎ⏕ࡣࡳࡽࢀࡎ㸪⏕⫱㸪཰㔞㸪ᯝᐇရ㉁➼࡟

࠾࠸࡚ࡶ͂┒ᒸ16ྕ̓࡜ࡢ㛫࡟᭷ពᕪࡣㄆࡵࡽࢀ࡞

࠿ࡗࡓ㸦㧗ᶫ㸪㧗஭ཬࡧᯇᮏ㸪1993㸧㸬

෗┿2 ͂M16-AR̓⣔⤫࡟ㄆࡵࡽࢀࡓ4ᯛࡢ」ⴥ㸬

෗┿3 㟢ᆅ᱂ᇵ࡟࠾ࡅࡿ͂࢔࢟ࢱ࣮࣋ࣜ̓㸬

(4)

ࡑࡇ࡛㸪ࡇࢀࡽ3⣔⤫ࡢ࠺ࡕࡸࡸᯝᐇ཰㔞ࡀከ࠿

ࡗࡓ͂M16-AR2̓ࢆ͂࢔࢟ࢱ࣮࣋ࣜ̓㸦෗┿3㸧࡜࿨

ྡࡋ࡚1990ᖺ9᭶࡟㎰ᯘỈ⏘┬࡟✀ⱑⓏ㘓ࡢ⏦ㄳࢆ

ࡋ㸪1992ᖺ9᭶࡟Ⓩ㘓㸦➨3224ྕ㸧ࡀㄆࡵࡽࢀࡓ 㸦㧗ᶫ㸪1994㸧㸬

࢖ࢳࢦⴎ㯤⑓⪏⑓ᛶရ✀ࡢ⫱ᡂ

ࢃࡀᅜࡢ࢖ࢳࢦⴎ㯤⑓⪏⑓ᛶရ✀⫱ᡂࡢ⌧≧

࢖ࢳࢦⴎ㯤⑓㸦෗┿4㸧ࡣᅵተ⑓ᐖ࡛⸆๣➼࡟ࡼ

ࡿ㜵㝖ࡀࡁࢃࡵ࡚ᅔ㞴࡞⑓ᐖࡢ୍ࡘ࡛࠶ࡿࡀ㸪1990 ᖺ௦ึࡵࡲ࡛ࡣᮏ⑓⪏ᛶࡢရ✀⫱ᡂࡣዉⰋ┴㎰ᴗヨ

㦂ሙ㸦⌧ዉⰋ┴㎰ᴗ⥲ྜࢭࣥࢱ࣮㸧ࡢෆ⸨㸦1991㸧

࡟ࡼࡗ࡚ಁᡂ᱂ᇵ⏝ࡢရ✀࡛⾜ࢃࢀ࡚࠸ࡿ࡟㐣ࡂ࡞

࠿ࡗࡓ㸬ᮏ⑓ࡣᬮᆅࡢࡳ࡞ࡽࡎᮾ໭ࡸ໭ᾏ㐨࡞࡝ࡢ ᐮᆅ࣭ᐮ෭ᆅ࠾࠸࡚ࡶࡑࡢⓎ⏕ࡀ☜ㄆࡉࢀ⿕ᐖࡀ ᚎࠎ࡟ᣑ኱ࡋࡘࡘ࠶ࡗࡓࡀ㸪ࡇࢀࡽࡢᆅᇦ࡟㐺ࡍࡿ

࢖ࢳࢦⴎ㯤⑓⪏⑓ᛶရ✀ࡢ⫱ᡂࡣ඲ࡃᡭࡘ࠿ࡎࡢ≧

ែ࡛࠶ࡗࡓ㸬ࡑࡇ࡛㸪ⴭ⪅ࡽࡣᐮ෭ᆅࡢ㟢ᆅ᱂ᇵࡸ

ప ࢝ࢵࢺ᱂ᇵ࡟㐺ࡍࡿⴎ㯤⑓⪏⑓ᛶရ✀ࡢ⫱ᡂ࡟

ྲྀࡾ⤌ࡴࡇ࡜࡜ࡋࡓ㸬

⪏⑓ᛶᰴࡢసฟἲ

⪏⑓ᛶရ✀ࡢ⫱ᡂ࡟࠶ࡓࡗ࡚㸪ࡲࡎ⫱✀⣲ᮦ࡟ࡘ

࠸᳨࡚ウࡋࡓ㸬ࡑࡢ⤖ᯝ㸪ࢃࡀᅜ࡛⫱ᡂࡉࢀࡓᐮ෭ ᆅྥࡁရ✀ࡢ୰୍࡛Ꮨᡂࡾᛶࡢ࣮͂࣋ࣝࣝࢪࣗ̓ࡸ

ᅄᏘᡂࡾᛶࡢ࢚͂ࣂ࣮࣮࣋ࣜ̓ࡀⓎ⑓ࡋ࡟ࡃ࠸ࡇ࡜

ࡀ᫂ࡽ࠿࡟࡞ࡗࡓ㸦㧗ᶫ㸪ྂᒇ㸪ᯇᮏཬࡧ㧗஭㸪1997 TakahashiFuruyaMatsumotoKanda and Yoshida㸪

1998㸧㸬ࡑࡇ࡛㸪⫱✀ぶ࡜ࡋ࡚ⴎ㯤⑓࡟ᑐࡋ࡚ࡢ⪏ᛶ

ࡣ࡞࠸ࡀᯝᐇࡀ◳ࡃ᪥ᣢࡕᛶ࡟ඃࢀ㟢ᆅ᱂ᇵ➼࡟㐺 ࡋ࡚࠸ࡿ͂Pajaro̓ࢆẕᮏ࡟㸪ⴎ㯤⑓࡟ẚ㍑ⓗᙉࡃ ᐮ෭ᆅࡢ㟢ᆅ࠾ࡼࡧప ࢝ࢵࢺ᱂ᇵ➼࡟ᗈࡃ฼⏝ࡉ

ࢀ࡚࠸ࡿ࣮͂࣋ࣝࣝࢪࣗ̓ࢆ∗ᮏ࡜ࡋ࡚⏝࠸ࡓ㸬ࡇ

ࢀࡽ2ရ✀ࡢ஺㞧࡟ࡼࡗ࡚ᚓࡽࢀࡓᐇ⏕ࢆ⏝࠸࡚

1997ᖺ࡜1998ᖺࡢ2ᅇ࡟ศࡅ࡚࢖ࢳࢦⴎ㯤⑓⳦㸦ศ 㞳⳦ᰴ㸸91-40㸪⬊Ꮚ⃰ᗘ㸸1×106ಶml㸧ࢆ᥋✀ࡋ⪏

⑓ᛶᰴࡢ㑅ᢤࢆ⾜ࡗࡓ㸬㑅ᢤᚋ㸪⪏⑓ᛶᰴࡢቑṪࢆ

⾜࠸1999ᖺ㹼2001ᖺ࡟⛅⏣┴‮ἑᕷ⛅ࣀᐑࡢ࢖ࢳ ࢦⴎ㯤⑓Ⓨ⏕ᅡሙ࠾ࡼࡧ⛅⏣┴❧኱Ꮫ▷ᮇ኱Ꮫ㒊㝃 ᒓ㎰ሙ㸦⌧⛅⏣┴❧኱Ꮫ⏕≀㈨※⛉Ꮫ㒊㝃ᒓࣇ࢕࣮

ࣝࢻᩍ⫱◊✲ࢭࣥࢱ࣮㸧࡟࠾࠸࡚⏕⫱≉ᛶ࠾ࡼࡧⓎ

⑓ࡢ᭷↓ࢆㄪᰝࡋࡓ㸬

⪏⑓ᛶ⣔⤫ࡢㅖ≉ᛶ࠾ࡼࡧ᱂ᇵ㐺ᛂᛶ

㑅ᢤᰴ࡟ࡣ1006ಶయࡢᐇ⏕ࢆ⏝࠸ࡓࡀ㸪ࡑࡢ୰࡛

⑓ᚩࡀ඲ࡃㄆࡵࡽࢀࡎᡂⱑ࡟࡞ࡗ࡚ࡶ⏕⫱ࡀ㡰ㄪ࡞

3 ಶ య ࡟ ࡑ ࢀ ࡒ ࢀ ͂WB-A15̓㸪͂WB-B22̓ 㸪

͂WB-B33̓ࡢ⣔⤫ྡࢆ௜ࡋ㸪ᅡሙ࡟࠾ࡅࡿⴎ㯤⑓

⪏ᛶࡸ᱂ᇵ㐺ᛂᛶ➼࡟ࡘ࠸࡚ㄪᰝࡋࡓ㸬ࡑࡢ⤖ᯝ㸪

͂WB̓ࡢ3⣔⤫ࡣᅡሙ࡟࠾࠸࡚ࡶ1999㹼2001ᖺࡢ 3࢝ᖺ㛫඲ࡃⓎ⑓ࡀㄆࡵࡽࢀ࡞࠿ࡗࡓ㸦⾲3㸧㸬ࡲࡓ㸪 㟢ᆅ᱂ᇵ࡛ࡢ⏕⫱㸪཰㔞㸪ᯝᐇရ㉁➼࡟࠾࠸࡚ࡶ᪤

Ꮡရ✀ࡢ͂Pajaro̓㸪࣮͂࣋ࣝࣝࢪࣗ̓㸪͂┒ᒸ 16 ྕ̓࡜ྠ➼࠿ࡑࢀ௨ୖࡢ್ࢆ♧ࡋࡓ㸦Takahashi YoshidaKandaFuruya and Matsumoto2003㸧㸬

෗┿4 ࢖ࢳࢦⴎ㯤⑓ࡢึᮇࡢ⑓ᚩ㸬

ⱝ࠸」ⴥࡀ㯤໬ࡋ୙ṇᙧ࡟࡞ࡾ

▮༳㸪ࡑࡢᚋᰴࡀᯤṚࡍࡿ

ⴭ⪅ࡽࡢ◊✲࡛ࡣⴎ㯤⑓࡟㛵ࡍࡿ㑇ఏⓗ≉ᛶ࡟ࡘ

࠸ ࡚ ㄪ ᰝ ࡍ ࡿ ࡲ ࡛ ࡟ ⮳ ࡽ ࡞ ࠿ ࡗ ࡓ ࡀ 㸪Mori㸪

(5)

Kitamura and Kuroda㸦2005㸧ࡢ◊✲࡟ࡼࢀࡤ࢖ࢳࢦ ရ✀ࡢⴎ㯤⑓࡟ᑐࡍࡿ⪏ᛶ࡟ࡣ㉁ⓗ㑇ఏᏊࡲࡓࡣ㔞

ⓗ㑇ఏᏊࡢ࠸ࡎࢀ࠿ࡀ㛵୚ࡋ࡚࠸ࡿ࡜ࡉࢀ࡚࠸ࡿ㸬

࢖ࢳࢦࡢ᱂ᇵ✀ࡣከࡃࡢస≀࡜␗࡞ࡾ8ಸయ࡛࠶ࡿ

ࡇ࡜࠿ࡽ㸪㑇ఏᵝᘧࡢヲ⣽࡟ࡘ࠸࡚ࡣࡲࡔ୙᫂࡞Ⅼ

ࡶከ࠸㸬ⴭ⪅ࡽࡀ⫱ᡂࡋࡓ͂WB̓ࡢ 3 ⣔⤫ࡢ⪏⑓

ᛶࡀ㉁ⓗ㑇ఏᏊࡲࡓࡣ㔞ⓗ㑇ఏᏊࡢ࠸ࡎࢀ࡟ࡼࡿࡶ

ࡢ࠿࡟ࡘ࠸࡚ࡣࡲࡔ᫂ࡽ࠿࡛࡞࠸ࡀ㸪௒ᚋ㸪3 ⣔⤫

ࡀᐇ⏝ရ✀࡜ࡋ࡚ࡢ฼⏝࡟࡜࡝ࡲࡽࡎⴎ㯤⑓⪏ᛶࡢ

⫱✀⣲ᮦ࡜ࡋ࡚฼⏝ࡉࢀࡿྍ⬟ᛶࡶ㧗࠸ࡇ࡜࠿ࡽ㸪 㑇ఏᵝᘧ࡟ࡘ࠸࡚᫂ࡽ࠿࡟ࡋ࡚࠾ࡃࡇ࡜ࡀ㔜せ࡛࠶

ࡿ࡜ᛮࢃࢀࡿ㸬

෗┿ 6 ᯝᐇࡢගἑࡀ

≉࡟ඃࢀ࡚࠸ࡿ͂ࢽ

ࣗࡣࡿࡳ̓ࡢᯝᐇ㸬 ⴭ⪅ࡽࡀ⫱ᡂࡋࡓ3⣔⤫ࡢ࠺ࡕ㸪≉࡟͂WB-B33̓

ࡀⴎ㯤⑓⪏ᛶ࡛⛅⏣┴ෆࡢ⏘ᆅ࡟࠾ࡅࡿヨ㦂᱂ᇵ࡟

࠾࠸࡚ࡶ཰㔞ࡀᏳᐃࡋ࡚㧗࠿ࡗࡓࡢ࡛⫱ᡂ᏶஢ᚋ ࡢ2006ᖺ1᭶࡟✀ⱑⓏ㘓ࡢ⏦ㄳࢆ⾜ࡗࡓ㸬ࡑࡢᚋ㸪

᭩㢮ᑂᰝ࡜⌧ᆅᑂᰝ➼ࢆ⤒࡚2009ᖺ3᭶࡟͂ࡇࡲࡕ

࣮࣋ࣜ̓㸦෗┿5㸧࡜ࡋ࡚࿨ྡⓏ㘓㸦➨17763ྕ㸧ࡉ

ࢀࡓ㸦㧗ᶫ㸪2013㸧㸬͂ࡇࡲࡕ࣮࣋ࣜ̓ࡢ᱂ᇵ㠃✚

ࡣ࠶ࡲࡾከࡃ࡞࠸ࡶࡢࡢ㸪⌧ᅾ㸪㮵ゅᕷࢆ୰ᚰ࡟᱂ ᇵࡉࢀ࡚࠸ࡿ㸬࡯࠿ࡢ2⣔⤫㸦͂WB-A15̓͂WB- B22̓㸧࡟ࡘ࠸࡚ࡣⴎ㯤⑓⪏ᛶࡢⅬ࡛͂ࡇࡲࡕ࣋ࣜ

࣮̓࡜኱ࡁ࡞㐪࠸ࡀㄆࡵࡽࢀ࡞࠿ࡗࡓࡢ࡛✀ⱑⓏ㘓

ࢆ⾜ࢃ࡞࠿ࡗࡓࡀ㸪‮ἑᕷ࢖ࢳࢦ⏕⏘㞟ฟⲴ⤌ྜࡀ

͂WB- A15̓ࢆ͂ࡣࡿࡳ̓㸪͂WB-B22̓ࢆ͂ࢽ࣮ࣗ

ࡣࡿࡳ̓࡜ࡋ࡚ᮾி㸪ᶓ὾㸪௝ྎ㸪ᶓᡭ➼ࡢᕷሙ࡟

ฟⲴࡋࡓ࡜ࡇࢁ୍⯡ⓗ࡞ᯝᐇရ㉁㸦⢾ᗘ㸪㓟ᗘ㸧࡟

ຍ࠼࡚ᯝᐇࡢගἑ㸦෗┿6㸧ࡀඃࢀ࡚࠸ࡿ࡜ࡢホ౯

ࢆᚓࡓࡇ࡜࠿ࡽ㸪⌧ᅾ㸪ࡇࢀࡽࡢ2⣔⤫ࡶ᱂ᇵရ✀

࡜ࡋ࡚┴ෆࡢ࢖ࢳࢦ⏘ᆅ࡛฼⏝ࡉࢀ࡚࠸ࡿ㸬

⪏⑓ᛶ⫱✀ࡀᐮ෭ᆅࡢ࢖ࢳࢦ⏕⏘࡟ᯝࡓࡋࡓᙺ๭

ⴭ⪅ࡽࡢ◊✲ࢢ࣮ࣝࣉࡣ㸪1986ᖺ࡟࢖ࢳࢦࡢ⪏⑓

ᛶရ✀⫱ᡂࡢ◊✲ࢆ㛤ጞࡋ࡚௨᮶2012ᖺࡲ࡛ࡢ27 ᖺ㛫࡟✀ⱑⓏ㘓ရ✀2ရ✀㸪ᮍⓏ㘓ရ✀2ရ✀㸪ィ 4ရ✀ࡢᐇ⏝ရ✀ࢆ⫱ᡂࡋࡓ㸬࠸ࡎࢀࡢ⫱ᡂရ✀ࡶ

⛅⏣┴ࢆ୰ᚰ࡜ࡋࡓᐮ෭ᆅ࡟࠾ࡅࡿ࢖ࢳࢦ᱂ᇵ㎰ᴗ

࡟ᑐࡋ࡚㸪ᐇ⏝㠃࡛ᐤ୚ࡋ࡚࠸ࡿࡇ࡜ࡣ㛫㐪࠸࡞࠸㸬 ᐇ㝿࡟㸪‮ἑᕷ⛅ࣀᐑᆅ༊➼࡟ࡳࡿࡼ࠺࡟㸪ᙜヱရ

✀ࡢ㛤Ⓨᬑཬ࣭ᐃ╔࡟ࡼࡾ࢖ࢳࢦ᱂ᇵࡢ୍኱⏘ᆅᙧ ᡂࢆᐇ⌧ࡉࡏ࡚࠸ࡿࡢ࡛࠶ࡿ㸬

ᮏ◊✲ࢆ㛤ጞࡋࡓᙜึࡣ㸪኱Ꮫࡢᙺ๭࡜ࡋ࡚⌧ᅾ

࡯࡝ᆅᇦ㈉⊩ࡀᙉࡃྉࡤࢀ࡚࠸࡞࠿ࡗࡓࡇ࡜ࡶ࠶ࡗ

࡚࠿㸪◊✲ෆᐜࡢࡳ࡞ࡽࡎ◊✲ᴗ⦼ᩘࡀ㔜せどࡉࢀ

ࡿ኱Ꮫ࡟࠾࠸࡚ᐇ⏝ⓗ࡞⫱✀ࢆ◊✲ࢸ࣮࣐࡜ࡋ࡚ྲྀ

ࡾୖࡆࡿࡢࡣ኱Ꮫࡢ◊✲࡜ࡋ࡚㥆ᰁࡲ࡞࠸ࡢ࡛ࡣ࡞

࠸࠿㸪࡜࠸࠺ኌࡶᑡ࡞࠿ࡽࡎ࠶ࡗࡓࡇ࡜ࡣ☜࠿࡛࠶

ࡿ㸬ࡋ࠿ࡋ࡞ࡀࡽ㸪ⴭ⪅ࡽࡢ◊✲ࢢ࣮ࣝࣉࡣබ❧኱

Ꮫ㸦⌧ᅾ㸪බ❧኱Ꮫἲே㸧࡛◊✲࡟ᦠࢃࡿ⪅࡜ࡋ࡚㸪 ᆅᇦࡀᢪ࠼࡚࠸ࡿ῝้࡞ၥ㢟ࢆࡑࡢࡲࡲᨺ⨨ࡋ࡚࠾

ࡃࡇ࡜ࡣዲࡲࡋࡃ࡞࠸࡜ࡢᛮ࠸࠿ࡽᮏ◊✲࡟ྲྀࡾ⤌

ࢇ࡛ࡁࡓ㸬኱Ꮫࡢ◊✲࡜ࡋ࡚㸪ඛ➃ⓗ࡛኱ࡁ࡞ᡂᯝ ࡀᮇᚅ࡛ࡁࡿ◊✲ࡀᚲせ୙ྍḞ࡛࠶ࡿࡇ࡜ࡣㄡࡋࡶ

␲࠺వᆅࡣ࡞࠸ࡀࠊᮏ◊✲ࡢࡼ࠺࡟ከࡃࡢ᫬㛫ࢆせ ࡍࡿࡀᆅᇦ㔜どࡢᆅ㐨࡞◊✲ࡶᚲせ࡛࡞࠸࠿࡜⪃࠼

࡚࠸ࡿ㸬

ᩥ ⊩

Maekawa㸪N 㸪YamamotoMNishimura S㸬㸪 Kohmoto㸪K㸬 KuwadaM㸬and WatanabeY㸬 㸦1984㸧㸬Studies on host-specific AF-toxins produced by Alternaria alternata strawberry pathotype causing Alternaria black spot of strawberry㸬㸦1㸧Production of host-specific toxins and their biological activities 㸬 Annals of Phytopathological Society of Japan50 600-609㸬

෗┿5 㟢ᆅ᱂ᇵ࡟࠾ࡅࡿ

͂ࡇࡲࡕ࣮࣋ࣜ̓㸬

୕⃝▱ኸ㸪すᕝ┪ኈ㸪ᰰ᳃⨾ዴ㸦2012㸧㸬ࠕ࢖ࢳࢦရ

(6)

✀ࠕ࢚ࢵࢳ࢚ࢫ-138ࠖ࡟࠾ࡅࡿ㯮ᩬ⑓Ⓨ⏕࡜ࡑ ࡢ⑓ཎ⳦ࡢᙧែࠖࠗ໭᪥ᮏ⑓ᐖ⹸◊✲఍ሗ࠘63 92-96㸬

Mori㸪T㸬㸪KitamuraH㸬and Kuroda K㸬㸦2005㸧㸬Varietal differences in Fusarium wilt-resistance in strawberry cultivars and the segregation of this trait in F1 hybrids 㸬 Journal of Japanese Society Horticultural Science74(1) 㸪57-59㸬

ෆ⸨ ₩㸦1991㸧㸬ࠕ࢖ࢳࢦⴎ㯤⑓᢬ᢠᛶရ✀ࡢ⫱ᡂࠖ

ࠗ1991 ᖺᗘ࢖ࢳࢦࢭ࣑ࢼ࣮⣖せ࡜ࡑࡢ௚ ࠘ 106-118㸬

Namiki㸪F㸬㸪YamamotoM㸬㸪NishimuraS㸬㸪Nakatsuka㸪 S㸬Goto㸪T㸬KohmotoK㸬and Otani H㸬㸦1986a㸧㸬 Studies on host-specific AF-toxins produced by Alternaria alternata strawberry pathotype causing Alternaria black spot of strawberry㸬㸦4㸧Protective effect of AF-toxinϩ on AF-toxinϨinduced toxic action and fungal infection 㸬 Annals of Phytopathological Society of Japan52 428-436㸬 Namiki㸪F㸬㸪 Okamoto㸪H㸬Katou㸪 K㸬㸪 Yamamoto㸪

M㸬Nishimura㸪S㸬㸪Nakatsuka㸪S㸬Goto㸪T㸬 Kohmoto㸪K㸬㸪Otani㸪H㸬 and Novacky㸪A㸬 㸦1986b㸧㸬Studies on host-specific AF-toxins produced by Alternaria alternata strawberry pathotype causing Alternaria black spot of strawberry㸬㸦5㸧Effect of toxinson membrane potential of susceptible plants as assessed by electrophysiological method 㸬 Annals of Phytopathological Society of Japan52610-619㸬 㧗ᶫ᫓ᐿ㸦1994㸧ࠕ࢔࢟ࢱ࣮࣋ࣜࠖ㸬ఀᮾ ṇ㸦┘ಟ㸧

ࠗⶰ⳯ࡢ᪂ရ✀12࠘㸦pp㸬72㸧㸬ㄔᩥᇽ᪂ග♫㸬 㧗ᶫ᫓ᐿ㸦2001㸧ࠕ͂┒ᒸ16ྕ̓̿✀ⱑⓏ㘓ࡢክࢆ⤯

ࡓࢀࡓྡရ✀̿ࠖࠗ᪥ᮏ࢖ࢳࢦࢭ࣑ࢼ࣮⣖せ 2001 ࠘No.10 119-123㸬

㧗ᶫ᫓ᐿ㸦2006㸧㸬ࠕ⛅⏣┴࡟࠾ࡅࡿ࢖ࢳࢦ᱂ᇵࡢ⌧

≧࡜ᒎᮃࠖࠗ⛅⏣┴❧኱Ꮫ▷ᮇ኱Ꮫ㒊⣖せ࠘ 7 29-35㸬

㧗ᶫ᫓ᐿ㸦2013㸧ࠕࡇࡲࡕ࣮࣋ࣜࠖ㸬ఀᮾ ṇ㸦┘ಟ㸧

ࠗⶰ⳯ࡢ᪂ရ✀18࠘㸦pp㸬65㸧㸬ㄔᩥᇽ᪂ග♫㸬 Takahashi㸪H㸬㸪 Furuya㸪H㸬 MatsumotoT㸬㸪

KandaH㸬and YoshidaY㸬㸦1998㸧㸬Reactions of

F1 strawberry seedling crossed with cultivars differing in susceptibility to Fusarium wilt㸬 Supplement to Journal of Japanese Society Horticultural Science67(1) 㸪209㸬

㧗ᶫ᫓ᐿ㸪ྂᒇᘅග㸪ᯇᮏ ໅㸪㧗஭㝯ḟ㸦1997㸧㸬 ࠕࣛࣥࢼ࣮ᰴ࠾ࡼࡧ⮬Ṫ㸦S1㸧ᐇ⏕ࢆ⏝࠸ࡓ࢖

ࢳࢦရ✀㛫ࡢ࢖ࢳࢦⴎ㯤⑓᢬ᢠᛶࡢẚ㍑ࠖࠗᅬ ⱁᏛ఍㞧ㄅ࠘66 ู෉㸯 270-271㸬

㧗ᶫ᫓ᐿ㸪㧗஭㝯ḟ㸪ᯇᮏ ໅㸦1988㸧㸬ࠕ࢖ࢳࢦࡢ 㯮ᩬ⑓᢬ᢠᛶᰴࡢ⫱ᡂ࡟㛵ࡍࡿ◊✲㸦➨1ሗ㸧 ⱼ㡬⤌⧊ࡢᇵ㣴࡟ࡼࡗ࡚సฟࡋࡓࢡ࣮ࣟࣥಶ య฼⏝࡟ࡼࡿ㯮ᩬ⑓᢬ᢠᛶᰴࡢ㑅ᢤࠖࠗᅬⱁᏛ

఍ᮾ໭ᨭ㒊᫛࿴63ᖺᗘ◊✲Ⓨ⾲せ᪨࠘55-56㸬 Takahashi HTakaiTand MatsumotoT㸦1992㸧㸬

Resistant plant to Alternaria alternata strawberry pathotype selected from calliclones of strawberry cultivar Morioka-16 and their characteristics.

Jounal of Japanese Society Horticultural Science 61㸦2㸧323-329

㧗ᶫ᫓ᐿ㸪㧗஭㝯ḟ㸪ᯇᮏ ໅㸦1993㸧㸬ࠕᐮ෭ᆅࡢ

༙ಁᡂ᱂ᇵ࠾ࡼࡧ㟢ᆅ᱂ᇵ࡟࠾ࡅࡿ࢖ࢳࢦ㯮

ᩬ⑓᢬ᢠᛶ⣔⤫M16-AR1㸪2࠾ࡼࡧ3ࡢ㐺ᛂᛶࠖ

ࠗᅬⱁᏛ఍㞧ㄅ࠘62㸦1㸧129-134㸬

Takahashi㸪H㸬㸪YoshidaY㸬㸪Kanda H㸬㸪Furuya H㸬and Matsumoto T㸬㸦2003㸧㸬Breeding of Fusarium wilt-resistant strawberry cultivar suitable for field culture in Northern Japan 㸬 Acta Horticulture 626 113-118㸬

㧗஭㝯ḟ㸪᪋ᒣ⣖⏨㸪ⰼᒸ ಖ㸪బ⸨ᛅᘯ㸦1979㸧㸬 ࠕ࢖ࢳࢦ┒ᒸ 16 ྕࡢ≉ᛶ࡟ࡘ࠸࡚ࠖࠗ᫛࿴ 54 ᖺᗘᅬⱁᏛ఍⛅Ꮨ኱఍◊✲Ⓨ⾲せ᪨࠘146-147㸬 Ώ㎶ᗣṇ㸪ᱵᕝ Ꮫ㸦1977㸧㸬ࠕAlternariaᒓ⳦࡟ࡼࡿ

࢖ࢳࢦࡢ᪂⑓ᐖ࡟ࡘ࠸࡚ࠖࠗ᪥ᮏ᳜≀⑓⌮Ꮫ఍

ሗ࠘43 82㸬

Ώ㎶ᗣṇ㸪ᱵᕝ Ꮫ㸪すᮧṇᬰ㸦1978㸧㸬ࠕ࢖ࢳࢦ㯮

ᩬ⑓ࡢ⑓ཎ⳦࡟ࡘ࠸࡚ࠖࠗ᪥ᮏ᳜≀⑓⌮Ꮫ఍ሗ࠘

44 363㸬

ᖹᡂ25ᖺ11᭶30᪥ ཷ௜

ᖹᡂ25ᖺ12᭶11᪥ ཷ⌮

(7)

Breeding Strawberry Cultivars Resistant to Alternaria Black Spot and Fusarium Wilt Disease Suitable for Northern Japan

Harumi Takahashi1, Yasunori Yoshida1, Hiroomi Kanda1, Hiromitsu Furuya2, Tsutomu Matsumoto3 and Takatsugu Takai4

1 Department of Agribusiness, Faculty of Bioresource Sciences, Akita Prefectural University

2 Department of Biological Production, Faculty of Bioresource Sciences, Akita Prefectural University

3,4Akita Prefectural College of Agriculture

Alternaria black spot and Fusarium wilt are problematic diseases affecting strawberry production in Akita Prefecture in northern Japan. In this study, we aimed to address these diseases from the perspective of breeding. The strawberry cultivar “Morioka-16” , which is commonly cultivated in northern Japan, bears large fruits that are of excellent quality. However, this cultivar is particularly susceptible to Alternaria black spot. We attempted to develop an improved “Morioka-16” line that is resistant to the disease using somaclonal variations induced during tissue culture. Using this method, we succeeded in generating three resistant lines, “M16-AR 1” , “M16-AR 2” and “M16-AR 3” . The “M16-AR 2”

line developed in this study was registered as a new cultivar named “Akita Berry” and was introduced to strawberry-producing districts in Akita Prefecture. Further, to develop a new cultivar resistant to Fusarium wilt, we crossed the “Pajaro” and “Belle Rouge” cultivars. Three resistant lines, namely “WB-A15” , “WB-B22” and “WB-B33” were selected from the resulting F1 seedlings. Among these, “WB-B33” which retained a high crop yield, was registered as a new cultivar named “Komachi Berry” and introduced to strawberry-producing districts in Akita Prefecture.

Although “WB-A15” and “WB-B22” were not registered as cultivars, they were given the names “Harumi” and “New Harumi” respectively, and provided to various growers and cultivated throughout Akita Prefecture.

Keywords: Morioka-16㸪Alternaria black spot disease㸪somaclonal variation㸪Fusarium wilt disease㸪soil-borne disease㸪disease resistance

Correspondence to: Harumi Takahashi, 418 Ohmaki, Murayama, Yamagata 995-0203, Japan. E-mail: [email protected]

参照

関連したドキュメント

熱力学計算によれば、この地下水中において安定なのは FeSe 2 (cr)で、Se 濃度はこの固相の 溶解度である 10 -9 ~10 -8 mol dm

In the present study, we will again use integral transforms to study the Black-Scholes-Merton PDE, specifically Laplace and Mellin transforms, which are the natural transforms for

Therefore, with the weak form of the positive mass theorem, the strict inequality of Theorem 2 is satisfied by locally conformally flat manifolds and by manifolds of dimensions 3, 4

Furthermore, the upper semicontinuity of the global attractor for a singularly perturbed phase-field model is proved in [12] (see also [11] for a logarithmic nonlinearity) for two

“Breuil-M´ezard conjecture and modularity lifting for potentially semistable deformations after

[9, 28, 38] established a Hodge- type decomposition of variable exponent Lebesgue spaces of Clifford-valued func- tions with applications to the Stokes equations, the

Section 3 is first devoted to the study of a-priori bounds for positive solutions to problem (D) and then to prove our main theorem by using Leray Schauder degree arguments.. To show

A real matrix with nonnegative entries and having a largest eigenvalue of multiplicity just one has a strictly positive left eigenvector (see the Appendix) if and only if there