ῳ QQ Ῡ  QQ ῵ Q῵Q ̰ ῖ̮̮῜Ῐῌῐ῏῍̮Ί̮̮῜Ῑ῎Ὶ̮̮῕῔ῑῒΐῗ

^{1)1) 2)}

Shuˆhei Nomura¹⁾, Munetoshi Maruyama¹⁾and Shiho Arai²⁾: The Soil Staphylinoid Faunas of the Akasaka Imperial Gardens and

the Tokiwamatsu Imperial Villa, Tokyo, Japan

!"#$%& '()

* +,-./01"23/45 63 789:;<=>?@

A!BC D EF%BGHI45GJK%B? '()*

L?MBNO/45P& ;<QRSTAUV%BC T1976 !WXY"#Z[\?$] ! ^_(`*%12a?b& !5cB de fg 1976hC

ij'kA12(GH4(!WXY")*#Z/lm"n [\/op$]MBC #Z[\/qrMB%"& dNomuraet al., 2000 D+

,-.b&/sth Auvwx/0y1z;<{|} d~ 2001h #Z[\2

/lm"C

1.

345^{2,100 m}645^{1,380 m}3€c€77‚3cdƒ¹hC„

I51 ha …†‡+8/ˆ‰Š‹7Œf+8c5 #Z/lm"C

2002ῌ2004 9Pc5Ž⁶j M¹²( d:;‘’<h )*“”•–”—/lm"C –

˜™Pšu=M›5op>œ/c5†:ž/>bMBCTA01:Ÿ?@ 23TA 02:¡!¢A23 do 2002 11£²⁰:hTA03:¡A7¤23 TA04:¥¦…7¤

G3§0do2003 4£2:hTA05:¨B©CD3§0TA06:ª@ 

G3H§0doE ¹⁰£²⁴:hTA07:ADG3§0TA08:«¬…­

78doE 12£24:hTA09:®FGN H TA10:A23do 2004

2£²⁷:h TA11:Ÿ?@ 23 dTA01E(h TA12:¯A23 do E ⁶

£24:hC

°"GHI^{2 ha};<=> !"‡+8±²Ug+8c5

#Z/lm"dƒ2hC2003ῌ2004 9²"m5Ž2j4(c5)*“”•–”—/lm"C

1) vwx/0yH0³´+ µI¶J3ῌ23ῌ1

Department of Zoology, National Science Museum, Hyakunincho 3ῌ23ῌ1, Shinjuku, Tokyo, 169ῌ0073 Japan E-mail: [email protected]

2) ·¸¹KJt 4º3-22-13

Musashidai 3ῌ22ῌ13, Ranzan-machi, Saitama, 355ῌ0216 Japan

vw/¼$ (39), 2005 3£25:

Mem. Natn. Sci. Mus., Tokyo, (39), March 25, 2005

1.

162

TT01:῎ TT02: ῍῎ 2003³³¹ TT03:

῍῎ TT04:῎ 2004624

2.

ῌ !"# $%!"&'()*+),-./01 23 456789:;<=>[email protected].;10 mC10 m5DE&1#F6G1#FHIJ 1 mC1 m5KLMNLO&⁵PQRSG1 KLMNLOT5UL7VWX@YZ&[9\J]

^;_`a9\J]0)bcUL9d;efg1Y&hi)jklmSHno48pqr s&tG)Yuvw&xyG1 xy`^1vw⁷⁰ῌ80z{U|LkTH}~G1 5€

HV0)h`^1‚ƒ5Yu„vw&…†.‡Lˆ‰G1 Š^‹5k& ŒPŽ

b&‘’“”P•bHIe)–—. ˜5™HIe),-.> šƒ&/01

3.

˜^›^5€#F9`^1Yu„“”P•b῏œh&ž6G) ˜^‹5Ÿ‚ 

„HIe) ¡¢ ῌ¢£¤¥&/¦1iH 45³I5§ƒ&"e1

1) ‚ ¨§ƒ ©ª‚ ¨H῍ «¬‚ ¨J῍ ­‚ ¨H῍N® œh¯°5±²`&³*§´6 G) kœh5Ÿƒ6µ¶ƒ65·¸n‹¹Jy`^;‚ ¨§ƒ5?I9:;bº|

»2. $%!"H7o;€#F5žS

!"#6$%!"5Yu„“”P•b« 163

Hῌ1 !"#$%

&'(& )*+,-./ 0S N i120 ni3 4^{, H}ῌ56S(ni/N)log(ni/N 789:;<= >?@& AB²C4DE^bit =7 )*+,-.0F G H&IJKJ7L2CMAJῌ5 Hῌ/logS N8O)*+,-./#$%3/^HῌN

2) PQJaccardRSTU CCV WXYSimpsonU NSCU-.ZPQT3

C: 0[\PQ]7L2C²0PPQ3^:-.Z_`a(b7c:

deC '27C =7c: fCghiYjk(1989)lmnopqrs

῏ ῎ ῍ ῌ

tu7v%)*+w*x&y=z{|}~P0F7€3 H0‚& ƒ„…†‚

&‡.ˆ ‰ƒ„Š ‹Œ…Ž‚&‡.ˆ ‰‹ŒŠ &C Staphylinoidea z{|}~

Ptiliidae ‘}’o“”‘~

1. Dipentium japonicum(K. Sawada)”’•–—‘}’o“”‘~

‰ƒ„Š TA02 (11 exs.); TA04 (1 ex.); TA07 (9 exs.); TA10 (16 exs.); TA12 (2 exs.).

‰‹ŒŠ TT04 (1 ex.).

2. Acrotrichis thoracica(Waltl)z˜™š‘}’o“”‘~

‰ƒ„Š TA01 (11 exs.); TA02 (5 exs.); TA05 (4 exs.); TA06 (1 ex.); TA07 (13 exs.); TA09 (5 exs.); TA10 (31 exs.); TA11 (44 exs.); TA12 (20 exs.).

‰‹ŒŠ TT01 (1 ex.); TT02 (1 ex.); TT03 (6 exs.); TT04 (4 exs.).

3. Nossidium japonicumY. Sawada›œ*•žw‘}’o“”‘~

Nomura, Kishimoto & Watanabe (2000)& Ptiliidae, gen. et sp. indet.ˆ>0&I Ÿ† †& glG ¡¢£7¤¥ˆ8¦&I

‰ƒ„Š TA02 (1 ex.).

‰‹ŒŠ TT02 (1 ex.).

Leiodidae§¨o“”‘~

1. Dermatohomoeus terrenus(Hisamatsu)©•˜ª«§¨o“”‘~

‰ƒ„Š TA01 (3 exs.); TA02 (5 exs.); TA03 (2 exs.); TA04 (2 exs.); TA05 (3 exs.); TA06 (5 exs.);

TA07 (1 ex.); TA09 (2 exs.); TA10 (11 exs.); TA11 (8 exs.); TA12 (35 exs.).

‰‹ŒŠ TT01 (1 ex.); TT03 (2 exs.); TT04 (5 exs.).

Scydmaenidae ”¬‘~

1. Euconnussp. 1

‰ƒ„Š TA01 (1 ex.); TA02 (2 exs.); TA03 (2 exs.); TA04 (4 exs.); TA05 (2 exs.); TA06 (2 exs.);

TA07 (4 exs.); TA08 (4 exs.); TA09 (1 ex.); TA10 (6 exs.); TA11 (2 exs.); TA12 (6 exs.).

‰‹ŒŠ TT02 (8 exs.); TT04 (3 exs.).

2. Euconnussp. 2

‰ƒ„Š TA09 (2 exs.); TA11 (1 ex.).

3. Euconnussp. 3

>0Nomura, Kishimoto & Watanabe (2000)7­®:lG ¯t°±›§w WX²Y³´µ¶Y·¸¹º

164

! "#$%&

'()* TA06 (3 exs.).

'* TT01 (2 exs.); TT02 (2 exs.); TT03 (16 exs.); TT04 (18 exs.).

4. Cephennodessp.

'()* TA04 (1 ex.); TA07 (3 exs.); TA08 (1 ex.).

'* TT03 (1 ex.); TT04 (1 ex.).

Staphylinidae +,-./0

Omaliinae 123+,-./40

1. Olophrum arrowiScheerpeltz567123+,-./

()89:; <=>?

'()* TA03 (2 exs.); TA05 (1 ex.).

Proteininae +@A6+,-./40

2. Megarthrus convexusSharpBCD+@A6+,-./

'()* TA01 (4 exs.); TA02 (3 exs.); TA03 (2 exs.); TA04 (3 exs.); TA07 (1 ex.); TA08 (1 ex.);

TA09 (2 exs.); TA10 (4 exs.); TA11 (5 exs.); TA12 (2 exs.).

'* TT03 (5 exs.); TT04 (3 exs.).

Micropeplinae EA+,-./40

3. Micropeplus fulvus japonicusSharpBFGEA+,-./

'()* TA01 (2 exs.); TA02 (4 exs.); TA06 (1 ex.); TA09 (2 exs.); TA10 (2 exs.); TA12 (6 exs.).

'* TT03 (2 exs.); TT04 (7 exs.).

Pselaphinae 5HI-J/40

4. Morana discedensSharpC35HI-J/

'()* TA04 (3 exs.); TA09 (1 ex.); TA10 (1 ex.).

5. Plagiophorus fujiyamai(Kubota)KGLCMDC5HI-J/

'()* TA01 (24 exs.); TA02 (2 exs.); TA05 (3 exs.); TA11 (4 exs.).

Tachyporinae /HNO+,-./40

6. Mycetoporini, gen. & sp. undet. 1

'* TT02 (1 ex.); TT03 (1 ex.); TT04 (1 ex.).

7. Mycetoporini, gen. & sp. undet. 2 '()* TA02 (2 exs.); TA09 (1 ex.).

8. Sepedophilus germanus(Sharp)J.PQ3RST+,-./

'()* TA02 (2 exs.); TA03 (3 exs.); TA04 (5 exs.); TA08 (1 ex.); TA09 (2 exs.); TA10 (2 exs.);

TA11 (5 exs.); TA12 (4 exs.).

9. Sepedophilus pumilus(Sharp)+FUQ3RST+,-./

'()* TA01 (4 exs.); TA02 (29 exs.); TA03 (1 ex.); TA04 (9 exs.); TA05 (3 exs.); TA07 (2 exs.);

TA08 (1 ex.); TA10 (3 exs.); TA11 (23 exs.); TA12 (13 exs.).

()VWXYVWZ[\]^+,-./_ 165

10. Sepedophilussp. 1 TA10 (1 ex.).

11. Tachyporus celatusSharp

TA06 (1 ex.).

12. Tachinus mimulusSharp

TA01 (1 ex.); TA04 (1 ex.); TA08 (2 exs.); TA09 (4 exs.).

Aleocharinae 13. Aleocharasp.

! TT04 (1 ex.).

14. Athetasp. 1

TA03 (1 ex.); TA04 (1 ex.); TA07 (1 ex.); TA08 (6 exs.); TA09 (4 exs.); TA10 (1 ex.).

! TT03 (3 exs.); TT04 (4 exs.).

15. Athetasp. 2

TA02 (8 exs.); TA03 (5 exs.); TA04 (4 exs.); TA05 (34 exs.); TA06 (8 exs.); TA07 (20 exs.); TA08 (45 exs.); TA09 (19 exs.); TA10 (45 exs.); TA11 (5 exs.); TA12 (2 exs.).

! TT03 (15 exs.); TT04 (10 exs.).

16. Athetasp. 3

TA08 (8 exs.); TA09 (2 exs.); TA10 (15 exs.).

17. Athetasp. 4

TA02 (2 exs.); TA09 (2 exs.); TA11 (1 ex.); TA12 (2 exs.).

! TT02 (4 exs.).

18. Athetasp. 5

! TT02 (4 exs.); TT04 (2 exs.).

19. Athetasp. 6

! TT02 (3 exs.); TT03 (10 exs.).

20. Zyras optatus(Sharp)"# $% &'()*+ TA01 (2 exs.); TA05 (1 ex.).

21. Drusilla sparsa(Sharp)$,--./

! TT04 (1 ex.).

22. Oligotasp.

! TT02 (1 ex.); TT03 (1 ex.); TT04 (1 ex.).

23. Oxypoda sp.

TA02 (1 ex.); TA09 (1 ex.).

! TT02 (1 ex.).

Scaphidiinae 012

24. Scaphisoma rubrumReitter$.3012

TA06 (1 ex.).

25. Scaphisoma crassipesAchard$3012

TA01 (3 exs.); TA05 (5 exs.); TA10 (10 exs.).

! TT01 (4 ex.).

456789:;<8=>?@

166

1. 4

/ ! "#ῌ $%&'(

)*+,-./*01) 2 2 2 2

345/*01) 2 2 2

6789+:;/*01) 2 2 2

<+3=>?@01) 2 2 2 2

=AB+4C/ 2

Euconnussp. 1 2 2 2 2

Euconnussp. 2 2 2

Euconnussp. 3 Dῌῌ D

Cephennodessp. 2 2 2

E5FGH> D

I@B345 2 2 2 2

Megarthrussp. 2

IJK+4 2 2 2 2

Petaloscapussp. 2

@>E;L/ 2 2 2

Natypleurina, sp. 2

MN<1=*E;L/ 2 2

9KO@PB@E;L/ 2 2

Mycetoporini, sp. 1 2 2

Mycetoporini, sp. 2 2 2 2

/*=>01) 2 2 2

JQ8=>01) 2 2 2

Sepedophilussp. D

5R;7S 2 2

0T;@B4 2 2 2

Aleocharasp. 2

Athetasp. 1 2 2 2

Athetasp. 2 2 2 2 2

Athetasp. 3 D

Athetasp. 4 D D

Athetasp. 5 D

Athetasp. 6 D

)Q85E;1J 2 2

E6IIUV 2 2

Oligotasp. D

Oxypodasp. 2 2 2 2

Oxypodini, sp. 2

EUWC<01)/ 2 2 2

EXMWC<01)/ 2 2 2 2

=>5IJK 2 2 2

BYJIJK 2 2 2

E@PZ>P 2 2

E[Z>P\ 2

=ZE>P 2

)5>P 2 2

JKHO+4 2 2 2 2

5-.] 2

0E;^5 D

E=>7S 2 2

+,3).Z 2 2 2

5N4Y5 2 2 2

_ 34 25 73ῌ 27

ῌ` "#abc !defghijk ῌῌ` "# lm$%&'(nopqrstuv ubDwhij

xyd !xyz 167

Oxytelinae

26. Anotylus laticornis(Sharp)

TA04 (1 ex.).

TT01 (1 ex.); TT02 (2 exs.).

27. Anotylus lewisius(Sharp)

TT03 (2 exs.); TT04 (12 exs.) Steninae

28. Stenus cicindeloides(Schaller)

TA06 (1 ex.); TA12 (2 exs.).

29. Stenus melanarius verecundusSharp

TA05 (1 ex.).

Euaesthetinae

30. Edaphus carinicollisBernhauer !

TA01 (4 exs.); TA03 (3 exs.); TA04 (4 exs.); TA06 (11 exs.); TA07 (20 exs.); TA08 (1 ex.); TA09 (1 ex.); TA10 (21 exs.); TA12 (25 exs.).

TT01 (3 exs.); TT03 (5 exs.); TT04 (12 exs.).

Paederinae "#$

31. Astenus latifrons(Sharp)%"&

TT03 (1 ex.);

Staphylininae

32. Neobisnius pumilus(Sharp)'(

TA08 (1 ex.).

33. Philonthus gastralisSharp)*#

TA05 (2 exs.); TA11 (1 ex.).

TT04 (4 exs.).

34. Ocypus lewisiusSharp+

TA01 (1 ex.); TA03 (3 exs.); TA05 (1 ex.); TA06 (1 ex.); TA07 (1 ex.); TA10 (3 exs.).

,2. -./04123456789:;<=>?@

AB2C DE FGHIJ

KL 34 25 73 27

34 17 28 17

25 0.40/0.68 17 11

DE73 0.35/0.82 0.21/068 16

FGHIJ27 0.38/0.63 0.27/0.44 0.19/0.59 MNO PQKLR STO <=UVL(CC/NSC).

XYZ[\]^_`\abcd 168

ῌ ῍

4^34425442 !"#$%&'( ")*1+,-'./+Nomura, Kishimoto & Watanabe (2000)$0 1-'23456789:;<=>?+.!@A&B-'CD.EFGHIJ⁽²⁰⁰¹⁾$KL'M

*3 6NO POQR#>?STU U VWXYUZ[(

\ TA01 TA02 TA03 TA04 TA05 TA06 TA07 TA08 TA09 TA10 TA11 TA12

]^_`abc^de] 11 1 9 16 2 fghc^de] 11 5 4 1 13 5 31 4 20 ij=k_lmc^de] 1

n_fopqrde] 3 5 2 2 3 5 1 2 11 8 35

Euconnussp. 1 1 2 2 4 2 2 4 4 1 6 2 6

E.sp. 2 2 1

E.sp. 3 3

Cephennodessp. 1 3 1

shtu9p 2 1

vr:fgh 4 3 2 3 1 1 2 4 5 2

vwx_g 2 4 1 2 2 6

rpsmyc 3 1 1

kxzr{:rsmyc 24 2 3 4

Mycetoporini, gen. & sp. undet. 2 2 1

c^opde] 2 3 5 1 2 2 5 4 w|=opde] 4 29 1 9 3 2 1 3 23 13

Sepedophilussp. 1

h}mj~ 1

dmr:g 1 1 2 4

Athetasp. 1 1 1 1 6 4 1

A.sp. 2 8 5 4 34 8 20 45 19 45 5 2

A.sp. 3 8 2 15

A.sp. 4 2 2 1 2

]|=hsmew 2 1

sivv€ 1

Oxypodasp. 1 1

s€‚ƒnde] 1

s„…‚ƒnde] 3 5 0

ophvwx 1

:†wvwx 2 12

sr{‡p{ 1 2

]hp{ 1

wx9z_g 4 3 4 11 20 1 1 21 25

sopj~ 1

_`f]b‡ 2 1

hˆg†h 1 3 1 1 1 3

‰STU(N) 60 77 26 52 60 35 75 71 51 172 99 119 U(S) 12 14 11 15 12 11 11 11 16 16 11 12

Š‹VWX(Hῌ) 2.81 3.03 3.32 3.41 2.39 2.83 2.71 1.91 3.27 3.21 2.45 2.87

ŒVWX(Jῌ) 0.78 0.80 0.96 0.87 0.67 0.82 0.78 0.57 0.82 0.80 0.71 0.80

ŽVWX(HῌN) 169 234 86.3 178 144 99.1 203 140 167 553 243 341

6NO#6NŒ 169

1 4!"#$

%&'⁷³()*+,-(.'/0 12$ 345(.'/6789:$

34;<=>?@0

4!ABCD*²(CEF.*GH-²>I ²JCEF.

IKLM'NO-0@0P7 LM*QRST$U8>V0'7 %&CD' 2J0<>W0-5X0Y'=T Z789: [#5CD'X0 CD

\0#$712(.80]^_'%&`a-0Tb'c dT Z71289:

[#5CD$=@4\0

1289:[#7 ef.(.gh>ij,7 3(/EF.*GH(/D

*kHlm*³no>I (.p$12¹²!b>W0-^12.707 89:#^12.25(

#qT'7 6>89:#rsJ,'t,67 u@v$@0 ef.p$12#74.57 89 :#^50.0$H,4v'ws p/E^(Hῌ)$12#p^2.857 89:#^3.06(.

$xy-0T'7Y5v't,0Pu@v#$@0 Az/E(Jῌ)*wT12' p^0.78\67 89:#$^0.59{{X0 |/E^(HῌN)$7 ef.v't,6}~dT

4. 8€ ‚ƒ„!b…†‡ef.7 (.7 /EF.N

( TT01 TT02 TT03 TT04

ˆ‰Š‹ŒŽ‰ˆ 1

‘’“”Ž‰ˆ 1 1 6 4

•–—Š˜™Ž‰ˆ 1

šŠ’›œžˆ 1 2 5

Euconnussp. 1 8 3

E.sp. 3 2 2 16 18

Cephennodessp. 1 1

Ÿž ‘’“”‘¡¢Ž£ 5 3

Ÿ¤¥Š“‘¡¢Ž£ 2 7

Mycetoporini, gen. & sp. undet. 1 1 1 1

Aleocharasp. 1

Athetasp. 1 3 4

A.sp. 2 15 10

A.sp. 4 4

A.sp. 5 4 2

A.sp. 6 3 10

¦¢•ŸŸ§¨‘¡¢Ž£ 1

Oligotasp. 1 1 1

Oxypodasp. 1

¦£©ª«£¬šˆ 4

›œŽ”Ÿ¤¥‘¡¢Ž£ 1 2

 ­¤Ÿ¤¥‘¡¢Ž£ 2 12

¤¥®¯Š“‘¡¢Ž£ 3 5 12

¦££™°”‘¡¢Ž£ 1

Š‹’¡ˆŒ£±‘¡¢Ž£ 4

²ef.(N) 12 28 70 90

(.(S) 6 11 14 18

p/E(Hῌ) 2.36 3.07 3.20 3.59

Az/E(Jῌ) 0.66 0.64 0.52 0.55

|/E(HῌN) 28.3 85.9 224 323

´µ¶·¸¹¶º»¼ 170

²¹³ !"#$%&' ()*% + %,-./0 123245678. 9 :;<=>?@' ABC%D 9: E:[email protected] J KL'%H2+M@$ 9:%NIOPQRST

Summary

In the faunistic survey carried out from 2002 to 2003, 34 staphylinoid species in four families were collected from the Akasaka Imperial Gardens, Minato-ku, Tokyo, and 25 species in four families were recognized in the Tokiwamatsu Imperial Villa, Shibuya-ku, Tokyo. Soil staphylinoid faunas of Akasaka, Tokiwamatsu, Fukiage Garden of the Imperial Palace (Nomura, Kishimoto & Watanabe, 2000), and the Institute for Nature Study, National Science Museum (Nomura & Kishimoto, 2001) were compared to one another by Jaccard’s coe$cient of community (CC) and Nomura & Simpson’s coe$cient (NSC). The a$nity between the Imperial Palace and the Institute for Nature Study is low. The fauna of Akasaka is closely allied to that of the Imperial Palace with 80% of common species. Soil staphylinoid communities at twelve sites in Akasaka and four sites of Tokiwamatsu were quantitatively sampled, and analyzed using three kinds of diversity indices (H῍, J῍ andH῍N). The di#erence in average diversity index (Shannon function:H῍) was not significant, but the relative diversity index (Pielou’sJ῍) was slightly high in Akasaka.

ῌ ῏ ῎ ῍

Nomura, S., T. Kishimoto & Y. Watanabe, 2000. A faunistic study on the staphylinoid beetles (Insecta, Coleoptera) from the Garden of the Imperial Palace, Tokyo, Japan. Mem. natn. Sci. Mus., Tokyo, (36):

257ῌ286.

UVWXYZ[\]X^_`aXbcdeXf_ghXijklXm nXoVpqXrstu 1976 vwxNyAz{| }~€ Edaphologia, (14): 25ῌ44

‚ƒX„…†‡ 2001. gZˆ‰Š€‹ŒŽ‘’ }~“”•(1)–—˜™š<ˆ Ž‘’

›œ (33): 301ῌ312

[Nomura, S. & T. Kishimoto, 2001. The soil beetle fauna of the Institute for National Science Museum, Tokyo, Japan (1), Staphylinoidea.Misc. Rep. Inst. Nature Study, (33): 301ῌ312.]

žŸABCžŸ| }~ –—˜™š• 171