報告
ベトナムの沿岸域で
1995 ～2007年に記録されたカニ類
Brachyuran species recorded from the coastal region of Vietnam in 1995–2007
和田恵次
Keiji Wada
はじめに
ベトナムの沿岸域に見られるカニ類相についての 知見は，これまで特定地域の記録種がまとめられた もの(Starobogatov, 1972; Hong & San, 1993; Kosuge et al., 1997: Nhuong, 2003; Chertoprud et al., 2012; Khac et al., 2012)や，特定種の記録を報じたもの(Serène, 1981; Davie & Xuan, 2003; Nagahashi et al., 2007; Ng et al., 2010; Shahdadi et al., 2017)があるものの，北部 から南部までを広域的にまとめた報告は，筆者の知 る限りまだない．筆者は，1995年から2007年にかけ て，ベトナム北部，中部，南部で沿岸性のカニ類の 調査をする機会があり，各地でカニ類を採集するこ とができた．本報告は，そのときに採集されたカニ 類の報告である．調査地（Table 1）は主にマング ローブ湿地と砂浜海岸であり，分類群はイワガニ上科 とスナガニ上科が中心で，いずれも大阪市立自然史 博物館 （OMNH），和歌山県立自然博物館 （WMNH）， 千葉県立中央博物館（CBM）に登録している．記 録された種ごとに，標本登録番号，標本個体数，採 集地（Table 1の番号），採集年月日を記した．また 日本に記録のある種については和名を学名に併記す るとともに，採集者が著者以外の場合は採集者名を 括弧内に表示した． キンセンガニ科Matutidae
Matuta planipes Fabricius, 1798 アミメキンセンガニ OMNH-Ar-11350: 1, Loc. 21, Jul. 27, 2000
コブシガニ科Leucosiidae
Philyra biprotubera Dai & Guan, 1986 OMNH-Ar-11282: 1, Loc. 2, Dec. 13, 1995 ガザミ科Portunidae
Thalamita crenata Rüppel, 1830 ミナミベニツケガニ OMNH-Ar-11351: 11, Loc. 16, Nov. 30, 2003
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Table 1. Localities of sampling sites in Vietnam.
No Locality Latitude & Longitude 1 Duong Quan, Haiphong 20°53′N, 106°42′E 2 Cam River Estuary, Haiphong 20°50′N, 106°46′E 3 Cam River Estuary, Haiphong 20°49′N, 106°48′E 4 Dinh Vu Is., Haiphong 20°48′N, 106°45′E 5 Ngoi Hai, Do Son 20°43′N, 106°48′E 6 Quynh Cu, Thai Thuy 20°38′N, 106°34′E 7 Xuan Thuy, Thai Thuy 20°35′N, 106°36′E 8 Con Ngan, Giao Thuy 20°16′N, 106°34′E 9 Giao Thuy 20°14′N, 106°30′E
10 Tinh Gia 19°28′N, 105°47′E
11 Tach Ha, Ha Tinh 18°25′N, 105°53′E 12 Ky Nam, Ha Tinh 17°59′N, 106°29′E 13 Tung Hai, Quang Tri 17°01′N, 107°03′E 14 Huong Phong, Thua Thien 16°34′N, 107°36′E 15 Co Du, Thua Thien 16°21′N, 107°58′E 16 Lang Co, Thua Thien 16°15′N, 108°06′E
17 Hoi An 15°53′N, 108°19′E
18 Hoi An 15°53′N, 108°20′E
19 Hoi An 15°53′N, 108°21′E
20 Can Thanh, Can Gio 10°25′N, 106°57′E 21 Long Thanh, Can Gio 10°24′N, 106°55′E
イワガニ科Grapsidae
Metopograpsus quadridentatus Stimpson, 1858
Ar-11254: 1, Loc. 2, Dec. 9, 1995; OMNH-Ar-11341: 2, Loc. 9, Jul. 18, 2000
Metopograpsus thukuhar (Owen, 1839) ハシリイワガ ニモドキ
WMNH-NU-000375: 1, Loc. 16, Nov. 30, 2003 ベンケイガニ科Sesarmidae
Chiromantes dehaani (H. Milne Edwards, 1853) クロ ベンケイガニ
Ar-11243: 1, Loc. 1, Dec. 1, 1995; OMNH-Ar-11246: 2, Loc. 2, Dec. 14, 1995
Episesarma lafondii (Hombron & Jacquinot, 1846) オ オベンケイガニ
WMNH-NU-000105: 1, Loc. 18, Dec. 15, 2007; WMNH-NU-000322: 1, Loc. 13, Nov. 29, 2003 Neosarmatium smithi (H. Milne Edwards, 1853) アシ ハラガニモドキ
WMNH-NU-000323: 1. Loc. 9, 2003
Parasesarma affine (De Haan, 1837) クシテガニ Ar-11211: 5, Loc. 2, Dec. 14, 1995; OMNH-Ar-11212: 3, Loc. 3, Dec. 13, 1995
Parasesarma bengalense (Davie, 2003) OMNH-Ar-11257: 2, Loc. 14, Dec. 2, 2003 Parasesarma eumolpe (De Haan, 1895)
OMNH-Ar-11347: 1, Loc. 7, Jul. 22, 2001 (col. by Do Van Nhuong)
Parasesarma peninsulare Shahdadi, Ng & Schubart, 2018
Ar-11213: 3, Loc. 2, Nov. 24, 1995; OMNH-Ar-11214: 1, Loc. 10, Dec. 3, 2003; OMNH-Ar-11215: 4, Loc. 14, Dec. 2, 2003; OMNH-Ar-11216: 1, Loc. 11, Dec. 3, 2003; OMNH-Ar-11260: 1, Loc. 2, Dec. 13, 1995; OMNH-Ar-11334: 1, Loc. 9, Jul. 21, 2000 Parasesarma semperi (Bürger, 1893) フタバカクガニ モドキ
WMNH-NU-000106: 1, Loc. 19, Dec. 16, 2007 Sarmatium germaini (A. Milne Edwards, 1869) ギ ザ テアシハラガニ
OMNH-Ar-11244: 2, Loc. 2, Dec. 14, 1995; WMNH-NU-000324: 1, Loc. 9, 2003
モクズガニ科Varunidae Helice latimera Parisi, 1918
OMNH-Ar-11210: 2, Loc. 1, Nov. 27, 1995
Hemigrapsus takanoi Asakura & Watanabe, 2005 タカ ノケフサイソガニ
OMNH-Ar-10778: 8, Loc. 16, Nov. 30, 2003 Metaplax crenulata (Gerstaecker, 1856) （図1A）
OMNH-Ar-11303: 2, Loc. 21, Jul. 25, 2000 Metaplax elegans De Man, 1888
Ar-11255: 1, Loc. 2, Dec. 14, 1995; OMNH-Ar-11360: 1, Loc. 7, Jul. 22, 2001 (col. by Do Van Nhuong); WMNH-NU-000228～000229: 2, Loc. 2, Dec. 15, 1995
Metaplax sheni Gordon, 1931
OMNH-Ar-11261: 1, Loc. 2, Dec. 14, 1995
Metaplax takahashii Sakai, 1939 テナガアシハラガニ Ar-11248: 1, Loc. 2, Dec. 14, 1995; OMNH-Ar-11249: 2, Loc. 2, Dec. 15, 1995; OMNH-Ar-11250: 1, Loc. 2, Nov. 23, 1995; OMNH-Ar-11251: 1, Loc. 2, Dec. 2, 1995; OMNH-Ar-11252: 3, Loc. 3, Dec. 13, 1995; WMNH-NU-000230: 1, Loc. 2, Dec. 7, 1995; WMNH-NU-000231: 1, Loc. 2, Dec. 11, 1995 Parapyxidognathus deianira (De Man, 1888) トゲア シヒライソガニモドキ
WMNH-NU-000325: 1, Loc. 14, Dec. 2, 2003 Varuna litterata (Fabricius, 1798) オオヒライソガニ
OMNH-Ar-11304: 3, Loc. 9, Jul. 19, 2000; WMNH-NU-000388: 1, Loc. 16, Nov. 30, 2003
ムツハアリアケガニ科Camptandriidae Baruna sinensis Tan & Huang, 1995
OMNH-Ar-11322: 1, Loc. 21, Jul. 27, 2000 Baruna trigranulum (Dai & Song, 1986)
Ar-11284: 3, Loc. 2, Dec. 14, 1995; OMNH-Ar-11285: 3: Loc. 2, Dec. 15, 1995; OMNH-Ar-11323: 1, Loc. 9, Jul. 20, 2000; WMNH-NU-000390～000392: 3, Loc. 2, Dec. 15, 1995
Camptandrium sexdentatum Stimpson, 1858 ムツハア リアケガニ
OMNH-Ar-11286: 3, Loc. 2, Dec. 8, 1995
Deiratonotus cristatus (De Man, 1895) アリアケモドキ OMNH-Ar-11310: 2, Loc. 1, Dec. 10, 1995
Ilyogynnis microcheirum (Tweedie, 1937) OMNH-Ar-11321: 1, Loc. 21, Jul. 27, 2000 Paracleistostoma depressum De Man, 1895
Ar-11287: 7, Loc. 2, Dec. 15, 1995; OMNH-Ar-11359: 1, Loc. 7, Jul. 22, 2001 (col. by Do Van Nhuong)
コメツキガニ科Dotillidae
Dotilla wichmanni De Man, 1892 （図1B）
Ar-11266: 6, Loc. 3, Nov. 25, 1995; OMNH-Ar-11324: 1, Loc. 9, Jul. 20, 2000; OMNH-Ar-11325: 8, Loc. 21, Jul. 26, 2000; OMNH-Ar-11349: 11, Loc. 22, Jul. 28, 2000; WMNH-NU-000170～000175: 6, Loc. 19, Dec. 1, 2003; WMNH-NU-000221～000227: 7, Loc. 5, Apr. 6, 1995; WMNH-NU-000321: 1, Loc. 5, Apr. 6, 1995
Ilyoplax formosensis Rathbun, 1921
OMNH-Ar-11241: 2, Loc. 8, Apr. 8, 1995; WMNH-NU-000083～000085: 3, Loc. 18, Dec. 15, 2007; WMNH-NU-000086～000092: 7, Loc. 18, Dec. 15, 2007; WMNH-NU-000093～000101: 9, Loc. 8, Dec. 16, 2007; WMNH-NU-000102: 1, Loc. 8, Dec. 16, 2007; WMNH-NU-000278～000286: 9, Loc. 6, Apr. 6, 1995; WMNH-NU-000342～000356: 15, Loc. 18, Dec. 1, 2003
Ilyoplax ningpoensis Shen, 1940 （図1C）
Ar-11293: 2, Loc. 2, Dec. 7, 1995; OMNH-Ar-11294: 12, Loc. 2, Dec. 8, 1995; OMNH-Ar-11295: 9, Loc. 2, Nov. 23, 1995; OMNH-Ar-11296: 2, Loc. 2, Dec. 15, 1995; WMNH-NU-000103～000104: 2, Loc. 18, Dec. 17, 2007; WMNH-NU-000287～000297: 11, Loc. 4, Apr. 6, 1995
Ilyoplax orientalis (De Man, 1888)
WMNH-NU-000389: 1, Loc. 21, Jul. 27, 2000 図1. A: Metaplax crenulata, B: Dotilla wichmanni, C: Ilyoplax ningpoensis, D: Ilyoplax serrata, E: Pseudogelasimus loii,
Ilyoplax punctata Tweedie, 1935
Ar-11318: 1, Loc. 21, Jul. 25, 2000; Ar-11319: 1, Loc. 20, Jul. 26, 2000; OMNH-Ar-11338: 10, Loc. 20, Jul. 25, 2000 (col. by Jun Kitaura); OMNH-Ar-11358: 11, Loc. 21, Jul. 27, 2000 (col. by Jun Kitaura)
Ilyoplax serrata Shen, 1931 （図1D）
Ar-11301: 6, Loc. 11, Dec. 3, 2003; OMNH-Ar-11239: 1, Loc. 5, Apr. 6, 1995; OMNH-Ar-11240: 1, Loc. 8, Apr. 8, 1995; OMNH-Ar-11344: 3, Loc. 9, July. 20, 2000; OMNH-Ar-11345: 3, Loc. 21, Jul. 27, 2000; OMNH-Ar-11356: 11, Loc. 21, Jul. 27, 2000 (col. by Jun Kitaura); OMNH-Ar-11357: 1, Loc. 9, Jul. 20, 2000 (col. by Jun Kitaura); WMNH-NU-000232～ 000253: 22, Loc. 2, Dec. 12, 1995; WMNH-NU-000313 ～000320: 8, Loc. 4, Apr. 5, 1995; WMNH-NU-000393 ～000397: 5, Loc. 4, Apr. 5, 1995
Ilyoplax tansuiensis Sakai, 1939
OMNH-Ar-11345: 1, Loc. 9, June 28, 2001 (col. by Do Van Nhuong)
Pseudogelasimus loii Serène, 1981 （図1E）
OMNH-Ar-7142: 1, Loc. 18, Dec. 1, 2003; WMNH-NU-000001: 1, Loc. 17, Dec. 16, 2007; WMNH-NU- 000002～000014: 13, Loc. 17, Dec. 16, 2007; WMNH- NU-000015～000025: 11, Loc. 17, Dec. 17, 2007 Scopimera bitympana Shen, 1930
OMNH-Ar-11365: 3, Loc. 9, Jul. 21, 2000; WMNH-NU-000193～000195: 3, Loc. 15, Nov. 30, 2003 Scopimera curtelsona Shen, 1936 （図1F）
WMNH-NU-000056～000060: 5, Loc. 18, Dec. 16, 2007; WMNH-NU-000061: 1, Loc. 18, Dec. 16, 2007; WMNH-NU-000062～000071: 10, Loc. 18, Dec. 17, 2007; WMNH-NU-000072～000079: 8, Loc. 18, Dec. 17, 2007; WMNH-NU-000080～000082: 3, Loc. 18, Dec. 16, 2007; WMNH-NU-000181～000189: 9, Loc. 18, Dec. 1, 2003; WMNH-NU-000190～000192: 3, Loc. 18, Dec. 1, 2003
Scopimera intermedia Balss, 1934
Ar-11264: 11, Loc. 2, Nov. 24, 1995; OMNH-Ar-11265: 1, Loc. 4, Apr. 5, 1995; WMNH-NU- 000026～000031: 6, Loc. 11, Dec. 3, 2003; WMNH-NU-000032: 1, Loc. 10, Dec. 3, 2003; WMNH-NU-
000033～000043: 11, Loc. 15, Nov. 30, 2003; WMNH- NU-000044～000055: 12, Loc. 19, Dec. 16, 2007 Tmethypocoelis ceratophora (Koelbel, 1897)
OMNH-Ar-11242: 4, Loc. 8, Apr. 8, 1995; OMNH-Ar- 11263: 8, Loc. 2, Nov. 24, 1995; OMNH-Ar-11292: 6, Loc. 2, Dec. 12, 1995; OMNH-Ar-11302: 4, Loc. 18, Dec. 1, 2003; WMNH-NU-000267～000277: 11, Loc. 2, Dec. 12, 1995; WMNH-NU-000327～000330: 4, Loc. 19, Dec. 1, 2003; WMNH-NU-000337～000341: 5, Loc. 19, Dec. 1, 2003; WMNH-NU-000371: 1, Loc. 19, Dec. 1, 2003
オサガニ科Macrophthalmidae
Ilyograpsus paludicola (Rathbun, 1909)
CBM-ZC-9307: 1, Loc. 9, May, 17, 2001 (col. by Do Van Nhuong)
Macrophthalmus abbreviatus Manning & Holthuis, 1981 オサガニ
Ar-11258: 1, Loc. 3, Dec. 13, 1995; OMNH-Ar-11259: 4, Loc. 3, Nov. 25, 1995; OMNH-Ar-11367: 4, Loc. 9, Jul. 20, 2000
Macrophthalmus convexus Stimpson, 1858 フタハオ サガニ
OMNH-Ar-11328: 3, Loc. 22, Jul. 28, 2000
Macrophthalmus definitus Adams & White, 1849 ヨコ スジオサガニ
WMNH-NU-000199: 1, Loc. 11, Dec. 3, 2003 Macrophthalmus erato De Man, 1888
Ar-11320: 1, Loc. 22, Jul. 28, 2000; OMNH-Ar-11330: 2, Loc. 9, Jul. 20, 2000; OMNH-Ar-11355: 1, Loc. 21, Jul. 27, 2000 (col. by Jun Kiatura)
Macrophthalmus laevimanus H. Milne Edwards, 1852 OMNH-Ar-11329: 3, Loc. 22, Jul. 28, 2000
Macrophthalmus pacificus Dana, 1851 タイヨウオサ ガニ （図1G）
Ar-11247: 2, Loc. 2, Dec. 14, 1995; OMNH-Ar-11297: 1, Loc. 3, Dec. 13, 1995; OMNH-Ar-11366: 1, Loc. 9, Jul. 19, 2000
Macrophthalmus sulcatus H. Milne Edwards, 1852 OMNH-Ar-11331: 3, Loc. 20, Jul. 26, 2000 Macrophthalmus tomentosus Eydoux & Souleyet, 1842
OMNH-Ar-11306: 1, Loc. 20, Jul. 26, 2000; WMNH-NU- 000398: 1, Loc. 2, Dec. 15, 1995
ミナミコメツキガニ科Mictyridae Mictyris brevidactylus Stimpson, 1858
Ar-11291: 2, Loc. 3, Dec. 13, 1995; OMNH-Ar-11342: 3, Loc. 9, Jul. 20, 2000
スナガニ科Ocypodidae
Austruca lactea (De Haan, 1835) ハクセンシオマネ キ
Ar-11270: 3, Loc. 2, Nov. 24, 1995; OMNH-Ar-11332: 2, Loc. 9, Jul. 19, 2000; OMNH-Ar-11333: 1, Loc. 9, Jul. 20, 2000; WMNH-NU-000357～000358: 2, Loc. 10, Dec. 3, 2003; WMNH-NU-000364～000368: 5, Loc. 14, Dec. 2, 2003
Austruca perplexa (H. Milne Edwards, 1837) オキナ ワハクセンシオマネキ
Ar-11326: 6, Loc. 22, Jul. 28, 2000; OMNH-Ar-11348: 1, Loc. 20, Jul. 26, 2000
Gelasimus borealis Crane, 1975 ホンコンシオマネキ Ar-11269: 1, Loc. 2, Dec. 13, 1995; OMNH-Ar-11354: 2, Loc. 9, Jul. 18, 2000
Gelasimus vocans (Linnaeus, 1758) ヒメシオマネキ WMNH-NU-000200～000205: 6, Loc. 19, Dec. 1, 2003; WMNH-NU-000326: 1, Loc. 19, Dec. 1, 2003 Ocypode ceratophthalma (Pallas, 1772) ツノメガニ Ar-11327: 1, Loc. 21, Jul. 26, 2000; OMNH-Ar-11343: 2, Loc. 20, Jul. 26, 2000
Ocypode sinesis Dai, Song & Yang, 1985 ナンヨウス ナガニ
WMNH-NU-000376～000380: 5, Loc. 12, Nov. 28, 2003; WMNH-NU-000381: 1, Loc. 19, Dec. 1, 2003; WMNH-NU-000382～000385: 4, Loc. 16, Nov. 30, 2003; WMNH-NU-000386～000387: 2, Loc. 15, Nov. 30, 2003
Paraleptuca splendida (Stimpson, 1858) （図1H） OMNH-Ar-11317: 7, Loc. 19, Dec. 16, 2007 Tubuca acuta (Stimpson, 1858) （図1I）
Ar-11298: 7, Loc. 1, Nov. 28, 1995; OMNH-Ar-11299: 11, Loc. 3, Dec. 13, 1995; OMNH-Ar- 11300: 2, Loc. 2, Dec. 14, 1995; WMNH-NU-000196
～000198: 3, Loc. 11, Dec. 3, 2003; WMNH-NU- 000359～000363: 5, Loc. 11, Dec. 3, 2003
Tubuca arcuata (De Haan, 1835) シオマネキ Ar-11267: 1, Loc. 1, Nov. 28, 1995; OMNH-Ar-11364: 2, Loc. 9, Jul. 19, 2000
Tubuca paradussumieri Bott, 1934 OMNH-Ar-11268: 1, Loc. 2, Dec. 3, 1995 メナシピンノ科Xenophthalmidae
Anomalifrons lightana Rathbun, 1931 OMNH-Ar-11283: 1, Loc. 2, Dec. 15, 1995
謝 辞
本 研 究 の 現 地 調 査 に 際 し て 協 力 を い た だ い た Phan Nguyen Hong教授，Do Van Nhuong教授．Phan Dinh Trong博士，Phan Due Thanh博士，小菅丈治博 士，北浦 純博士，並びに一部の標本の同定に協力 いただいた駒井智幸博士にお礼申し上げる． 本研究成果は次の研究助成事業による現地調査か ら得られたものである．伊藤科学振興財団研究助成 金，日本学術振興会科学研究費補助金，日本学術振 興会・拠点大学交流事業，日本学生支援機構・フォ ローアップ事業． 文 献
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