㟷᳃┴䛻䛚䛡䜛䜹䜶䝹㢮䛾ᐤ⏕⹸┦

௧࿴ඖ (2019) ᖺᗘ

ᘯ๓኱Ꮫ኱Ꮫ㝔ᩍ⫱Ꮫ◊✲⛉ಟኈㄽᩥ

㟷᳃┴䛻䛚䛡䜛䜹䜶䝹㢮䛾ᐤ⏕⹸┦

ᘯ๓኱Ꮫ኱Ꮫ㝔ᩍ⫱Ꮫ◊✲⛉Ꮫᰯᩍ⫱ᑓᨷᩍ⛉ᐇ㊶䝁䞊䝇⌮⛉ᩍ⫱㡿ᇦ 18GP303

▼㒓ᒸ ༓ḷ㡢

2020 ᖺ 1 ᭶ 22 ᪥

┠㻌 㻌 㻌 㻌 㻌 㻌 ḟ

䠍䠊 䛿䛨䜑䛻㻌 㻌 --- 1

䠎䠊 ◊✲䛾⫼ᬒ --- 2

䠏䠊 ᮦᩱ䛸᪉ἲ --- 3

䠐䠊 ⤖ᯝ 4-1 ほᐹ䛥䜜䛯ᐤ⏕⹸䛾䝸䝇䝖 --- 4

4-2 ྛ✀䛾グ㍕䛸ฟ⌧≧ἣ --- 6

4-2 䜹䜶䝹ᐤ⏕⹸㢮䛾ᐤ⏕≧ἣ㻌 --- 31

䠑䠊 ⪃ᐹ 5-1 㟷᳃┴䛾䜹䜶䝹ᐤ⏕⹸㢮䛾ᐤ⏕≧ἣ䛸ศᕸ䛾≉ᚩ --- 32

5-2 㻌 㟷᳃┴⏘䜹䜶䝹ᐤ⏕⹸䛾ᐟ୺䛸ᐤ⏕㒊఩ --- 33

䠒䠊 ᦬せ --- 34

䠓䠊 ㅰ㎡ --- 34

䠔䠊 ᘬ⏝ᩥ⊩ --- 35 ῧ௜⾲ ⾲䠍ࠥ⾲ 4

ῧ௜ᅗ㻌 㻌 㻌 㻌 㻌 ᅗ䠍ࠥᅗ 28

䛿䛨䜑䛻

␗✀䛾⏕≀䛜୍⥴䛻⏕ά䛩䜛⌧㇟䜢ඹ⏕䛸䛔䛖 (ᕑబ 2013) . 䛣䛾䛖䛱, ⏕⫱䜔⦾Ṫ䛻 ᚲせ䛺ᰤ㣴䜢ᐟ୺䛻౫Ꮡ䛩䜛㛵ಀ䜢ᐤ⏕䛸䛔䛖 (ᾆ㒊 2003) . ື≀㻌 (ᚋ⏕ື≀) 䛿 35 㛛 䛛䜙䛺䜛䛜, 䛭䛾୰䛾 5 㛛䛿ᵓᡂ✀䛾䛩䜉䛶䛜ᐤ⏕ᛶ✀䛷༨䜑䜙䜜䛶䛔䜛. 䛭䜜௨እ䛾 4 㛛 䜒ከ䛟䛾ᐤ⏕✀䜢ྵ䜏, 䜎䛯, ୍㒊䛰䛜ᐤ⏕✀䜢ྵ䜐㛛䜒 10 㛛䛻䛾䜌䜛䛣䛸䛛䜙 (㛗㇂ᕝ

2016) , ᐤ⏕䛿ື≀⏺䛻ᗈ䛟䜏䜙䜜䜛⌧㇟䛷䛒䜛䛣䛸䛜䜟䛛䜛. ከ䛟䛾ᐤ⏕⹸䛿䜂䛸䛴, 䜎䛯

䛿䛯䛟䛥䜣䛾୰㛫ᐟ୺䜢ᣢ䛱, ᤕ㣗䜢䛿䛨䜑䛸䛩䜛⏕≀ྠኈ䛾䛛䛛䜟䜚䛾୰䛷⏕άྐ䜢᏶⤖

䛩䜛. 䛭䛾ព࿡䛷, ᐤ⏕⹸䛾ୡ⏺䛿⏕ែ⣔䛾⦰ᅗ䛸䜒䛔䛘䜛 (ᾆ㒊 2016) .

䜹䜶䝹㢮 (↓ᑿ୧⏕㢮) 䛿, ᗂ⏕᫬௦䛿“䛚䛯䜎䛨䜓䛟䛧”䛸䛧䛶Ỉ୰䛷⏕ᜥ䛧, ኚែᚋ䛿 ⲡᆅ䜔᳃ᯘ䛺䛹䛾㝣ୖ䛷⏕ά䛩䜛 (ᕷᕝ 1970) . 䜎䛯, ᗂ⏕䛿㞧㣗ᛶ䛰䛜, ᡂయ䛻䛺䜛䛸

⫗㣗ᛶ䛜ቑ䛧䛶᪻⹸䛺䛹䛾䛥䜎䛦䜎䛺ᑠື≀䜢ᤕ㣗䛧, 䛥䜙䛻, 䜹䜶䝹㢮⮬㌟䜒䝦䝡㢮䜔㫽 㢮, ့ங㢮䛺䛹䛾ከ䛟䛾㧗ḟᤕ㣗⪅䛾㣵䛸䛺䜛 (ᕷᕝ 1970) . Ỉ⏺䛸㝣ୖ䛻䜎䛯䛜䛳䛶ᬽ䜙 䛧, పḟ䛛䜙㧗ḟ䜈䛾㣗≀㐃㙐䛾୰㛫䛻఩⨨䛩䜛䜹䜶䝹㢮䛾⏕ែᏛⓗ䛺≉ᚩ䛛䜙, యෆ䛻 䛿㣗≀㐃㙐䜢฼⏝䛧䛶⏕ά䜢㏦䜛ᐤ⏕⹸䛜㇏ᐩ䛻ぢ䜙䜜䜛䛸᥎ 䛥䜜䜛. ஦ᐇ, 䜹䜶䝹㢮䛻 䛿ᐤ⏕⹸䛜ከ䛟, “ᐤ⏕䛥䜜䛶䛔䛺䛔䜹䜶䝹䜢᥈䛩䛣䛸䛾᪉䛜↓⌮” (ᕷᕝ 1970) 䛸ゝ䜟䜜䜛 䜋䛹䛷䛒䜛. ᪥ᮏ䛻䛚䛡䜛䜹䜶䝹㢮䛾ᐤ⏕⹸䛿 1900 ᖺ௦䛾ึ䜑䛛䜙◊✲䛜ጞ䜎䜚, ᒣཱྀᕥ

௰䜔ෆ⏣᫂ᙪ䜢୰ᚰ䛻䛧䛯ሗ࿌䛜䛒䜛 (Uchida and Ooi 2018; Uchida et al. 2019 ཧ↷) . 䛣 䜜䜎䛷, ᪥ᮏ䛾୧⏕㢮䛛䜙䛿 4 㛛 88 ✀䛾ෆ㒊ᐤ⏕⹸䛜▱䜙䜜䛶䛔䜛 (Uchida and Ooi 2018;

Uchida et al. 2019) .

䜹䜶䝹㢮䛿⛣ືᛶ䛜ప䛔䛯䜑ᆅ⌮ⓗ䛺ศ໬䛜㉳䛣䜚䜔䛩䛟, ᪥ᮏ䛾ᅾ᮶䛾䜹䜶䝹㢮 (39

✀) 䛾ᅛ᭷✀䛾๭ྜ䛿 70.3%䛻ཬ䜆 (ᯇ஭ 2016) . ᪥ᮏ䛻䛿እ᮶✀ 3 ✀䜢ྵ䜑 42 ✀䛾䜹 䜶䝹㢮䛜ศᕸ䛩䜛 (ᯇ஭ 2016) . 䛣䛾䛖䛱, ᮏᕞ໭➃䛻఩⨨䛩䜛㟷᳃┴䛻䛿 4 ⛉ 10 ✀䛾䜹 䜶䝹㢮䛜ศᕸ䛩䜛 (ྥᒣ 2003) . እ᮶䛾 1 ✀ (䜴䝅䜺䜶䝹) 䜢㝖䛟ᅾ᮶䛾 9 ✀䛾䜹䜶䝹㢮䛾 䛖䛱, ໭ᾏ㐨䛻䜒ศᕸ䛩䜛䝙䝩䞁䜰䝬䜺䜶䝹䜢㝖䛟 8 ✀䛿䛩䜉䛶㟷᳃┴䛜⮬↛ศᕸ䛾໭㝈䛸 䛺䛳䛶䛔䜛 (ྥᒣ 2003) . ከ䛟䛾䜹䜶䝹䛾ศᕸ䛾໭㝈䛷䛒䜛㟷᳃┴䛷䛿, ᐤ⏕⹸┦䛻䜒ᆅ

⌮ⓗ䛺≉ᚩ䛜䜏䜙䜜䜛ྍ⬟ᛶ䛜䛒䜛. 䛧䛛䛧, 䛣䜜䜎䛷㟷᳃┴䛾䜹䜶䝹㢮䛛䜙ᐤ⏕⹸䛜✀䝺 䝧䝹䛷ሗ࿌䛥䜜䛯஦౛䛿, ྾⹸㢮 2 ✀ (Uchida and Itagaki 1975; Uchida and Itagaki 1980) , ༢⏕㢮 1 ✀ (Uchida et al. 1988) , ⥺⹸㢮 1 ✀ (Oyamada et al. 1998) ࡢィ 4 ✀䛻䛸䛹䜎䛳䛶 䛚䜚, ᐤ⏕⹸┦䛾඲ㇺ䛿᫂䜙䛛䛻䛺䛳䛶䛔䛺䛔.

㻌 ▼㒓ᒸ (2018) 䛿ᘯ๓኱Ꮫ䛾༞ᴗ◊✲䛷, 2015 ᖺ䛛䜙 2017 ᖺ䛻䛛䛡䛶㟷᳃┴䛾 9 ᕷ⏫

ᮧ䛷᥇㞟䛧䛯䜹䜶䝹㢮 10 ✀ 94 ಶయ䛛䜙, ⧄ẟ⹸㛛, ᡥᙧື≀㛛༢⏕⥘, ྾⹸⥘, ᮲⹸⥘,

㖋㢌ື≀㛛, ⥺ᙧື≀㛛, ⎔ᙧື≀㛛䝠䝹⥘䛾 5 㛛 7 ศ㢮⩌䛻䜎䛯䛜䜛ᐤ⏕⹸䜢ぢ䛔䛰䛧

䛯. 䛧䛛䛧, 䛣䛾◊✲䛿ୖ఩ศ㢮⩌䛷䛾ྠᐃ䛻䛸䛹䜎䛳䛶䛚䜚, ከᵝᛶ䛾ᐇయ䛿䛔䜎䛰᫂䜙

䛛䛻䛷䛝䛺䛛䛳䛯. 䛭䛣䛷, ᮏ◊✲䛿, 䛣䜜䜎䛷᥇㞟䛧䛯㟷᳃┴⏘䛾䜹䜶䝹㢮䛾ᐤ⏕⹸䛾ᶆ

ᮏ䛸, 㟷᳃┴䜢୰ᚰ䛸䛧䛯໭ᮾ໭䛛䜙᪂䛯䛻᥇㞟䛧䛯䜹䜶䝹㢮䛾ᐤ⏕⹸䜢ศ㢮Ꮫⓗ䛻⢭ᰝ 䛩䜛䛣䛸䛻䜘䛳䛶, 㟷᳃┴䛾䜹䜶䝹㢮䛻ぢ䜙䜜䜛ᐤ⏕⹸┦䛾ᐇయ䜢᫂䜙䛛䛻䛩䜛䛣䛸䜢┠ⓗ䛸 䛧䛯.

2. ◊✲䛾⫼ᬒ

2-1㻌 㟷᳃┴䛻䛚䛡䜛䜹䜶䝹┦䛾≉ᚩ

㻌 㟷᳃┴䛻䛿௨ୗ䛾 4 ⛉ 10 ✀䛾䜹䜶䝹㢮䛜ศᕸ䛩䜛 (ྥᒣ 2003) (䜹䜶䝹㢮䛾Ꮫྡ䛿᪥

ᮏ∐⹸୧᳇㢮Ꮫ఍䝩䞊䝮䝨䞊䝆 (http://zoo. zool. kyoto-u. ac. jp/herp/wamei. html, 2019 ᖺ 1 ᭶ 19 ᪥᭱⤊☜ㄆ) 䛻䜘䜛) :

䝠䜻䜺䜶䝹⛉㻌 䜰䝈䝬䝠䜻䜺䜶䝹 Bufo japonicus formosus Boulenger, 1883 䜰䝬䜺䜶䝹⛉㻌 䝙䝩䞁䜰䝬䜺䜶䝹 Hyla japonica Günther, 1859

䜰䜹䜺䜶䝹⛉㻌 䝍䝂䜺䜶䝹 Rana tagoi Okada, 1928, 䝲䝬䜰䜹䜺䜶䝹 Rana ornativentris Werner, 1903, 䝖䝜䝃䝬䜺䜶䝹 Pelophylax nigromaculatus (Hallowell, 1861) , 䝒䝏䜺䜶 䝹 Glandirana rugosa (Temminck et Schlegel, 1838) , 䜴䝅䜺䜶䝹 Lithobates

catesbeianus (Shaw, 1802)

䜰䜸䜺䜶䝹⛉㻌 䝰䝸䜰䜸䜺䜶䝹 Rhacophorus arboreus (Okada et Kawano, 1924) , 䝅䝳䝺䞊䝀 䝹䜰䜸䜺䜶䝹 Rhacophorus schlegelii (Günther, 1858) , 䜹䝆䜹䜺䜶䝹 Buergeria buergeri (Temminck et Schlegel, 1838)

䛣䛾䛖䛱, ᶟኴ, ᮅ㩭, 䝻䝅䜰ἢᓊᕞ, ୰ᅜ໭㒊䛻䜒ศᕸ䛩䜛䝙䝩䞁䜰䝬䜺䜶䝹, ᮅ㩭, ୰ ᅜ, 䝻䝅䜰䛾୍㒊䛻䜒ศᕸ䛩䜛䝖䝜䝃䝬䜺䜶䝹, ୡ⏺ྛᆅ䛻⛣ධ䛥䜜䛶䛔䜛໭⡿ཎ⏘䛾䜴䝅 䜺䜶䝹௨እ䛾 7 ✀䛿᪥ᮏ䛾ᅛ᭷✀䞉ᅛ᭷ள✀䛷䛒䜛 (ᯇ஭ 2016) . ᪥ᮏᅾ᮶䛾 9 ✀䛾䛖䛱䠈

໭ᾏ㐨䛻䜒ぢ䜙䜜䜛䝙䝩䞁䜰䝬䜺䜶䝹䜢㝖䛟 8 ✀䛿㟷᳃┴䛜ᅜෆ䛾⮬↛ศᕸ䛾໭㝈䛻䛺䛳䛶 䛔䜛 (ྥᒣ 2003) . 䜰䝈䝬䝠䜻䜺䜶䝹䛸䝖䝜䝃䝬䜺䜶䝹䛿⌧ᅾ, ໭ᾏ㐨䛻䜒ศᕸ䛧䛶䛔䜛䛜, 䛣䜜䛿ᮏᕞ䛛䜙䛾ேⅭⓗ⛣ධ䛻䜘䜛 (ᯇ஭ 2016) . 䛣䛾䜋䛛, 㟷᳃┴䛷䛿䝙䝩䞁䜰䜹䜺䜶䝹 Rana japonica Boulenger, 1879 䛾グ㘓䛜䛒䜛䜒䛾䛾, ␲ၥど䛩䜛◊✲⪅䜒ከ䛟, ྥᒣ (2003) 䛿㟷᳃┴䛾䝣䜯䜴䝘䛛䜙㝖እ䛧䛶䛔䜛.

⛅⏣┴䛸ᒾᡭ┴䛻䛚䛡䜛䜹䜶䝹┦䛿㟷᳃┴䛷䛾≧ἣ䛸䜘䛟ఝ䛶䛔䜛. 㟷᳃┴⏘䛾䜹䜶䝹 㢮䛻ຍ䛘䛶, ⛅⏣┴䛸ᒾᡭ┴䛷䛿䝙䝩䞁䜰䜹䜺䜶䝹䛜, ᒾᡭ┴䛷䛿䠈䛥䜙䛻䠈䝖䜴䜻䝵䜴䝎䝹 䝬䜺䜶䝹 Pelophylax porosus porosus (Cope, 1868) 䛜ศᕸ䛩䜛 (ኴ⏣䜋䛛 2010) .

2-2㻌 㟷᳃┴䛻䛚䛡䜛ᐤ⏕⹸䛾◊✲ྐ

䜹䜶䝹㢮䛾ᐤ⏕⹸䛻㛵䛩䜛◊✲䛾ከ䛟䛿, 㛵すཬ䜃㛵ᮾᆅ᪉䜔༡すㅖᓥ䛷䛺䛥䜜, ໭᪥

ᮏ䛛䜙䛾グ㘓䛿ᴟ䜑䛶ᑡ䛺䛔. 㟷᳃┴䛛䜙䛾䜹䜶䝹ᐤ⏕⹸䛾グ㘓䛷, ✀䜎䛷≉ᐃ䛥䜜䛶䛔

䜛䜒䛾䛿, Uchida and Itagaki (1980) 䛻䜘䜛“㟷᳃┴⏘” (᪥௜䛸ሙᡤ䛾ヲ⣽䛿୙᫂) 䛾䝖䝜䝃

䝬䜺䜶䝹䛛䜙ぢฟ䛥䜜䛯྾⹸㢮 Omphalometridae ⛉䛾 Opisthioglyphe japonicus Uchida and

Itagaki, 1980, ෆ⏣ (1981) 䛜㟷᳃┴⏘䛾䝖䝜䝃䝬䜺䜶䝹䛛䜙グ㘓䛧䛯྾⹸㢮

Haematoloechidae ⛉䛾 Haematoloechus japonicus (Yamaguti, 1936) , Uchida et al. (1988) 䛜

“㟷᳃┴⏘” (᪥௜䛸ሙᡤ䛿୙᫂) 䛾䜰䝈䝬䝠䜻䜺䜶䝹䛛䜙グ㘓䛧䛯༢⏕㢮 Polystoma ozakii, 䛚䜘䜃, Oyamada et al. (1998) 䛜㟷᳃┴⏘䛾䝖䝜䝃䝬䜺䜶䝹䛸䜴䝅䜺䜶䝹䛛䜙グ㘓䛧䛯⥺⹸

㢮 Gnathostoma nipponicum Yamaguti, 1941 䛾 4 ✀䛾䜏䛷䛒䜛.

3. ᮦᩱ䛸᪉ἲ

㻌 䜹䜶䝹㢮䛾ᐤ⏕⹸䜢ㄪ䜉䜛䛯䜑䛻, 2015 ᖺ 6 ᭶䡚2018 ᖺ 12 ᭶䛻䛛䛡䛶, 㟷᳃┴䛸⛅⏣┴, ᒾᡭ┴䛛䜙᥇㞟䛥䜜䛯ᅾ᮶䛾䜹䜶䝹㢮 9 ✀ 243 ಶయ䜢ㄪ䜉䛯 (⾲ 1) . 䜹䜶䝹㢮䛾✀㢮䛿, 䝠䜻䜺䜶䝹⛉䛻ᒓ䛩䜛䜰䝈䝬䝠䜻䜺䜶䝹, 䜰䝬䜺䜶䝹⛉䛻ᒓ䛩䜛䝙䝩䞁䜰䝬䜺䜶䝹, 䜰䜹䜺䜶 䝹⛉䛻ᒓ䛩䜛䝍䝂䜺䜶䝹, 䝲䝬䜰䜹䜺䜶䝹, 䝖䝜䝃䝬䜺䜶䝹, 䝒䝏䜺䜶䝹, 䜴䝅䜺䜶䝹, 䜰䜸 䜺䜶䝹⛉䛻ᒓ䛩䜛䝰䝸䜰䜸䜺䜶䝹, 䝅䝳䝺䞊䝀䝹䜰䜸䜺䜶䝹, 䜹䝆䜹䜺䜶䝹䛷, 㟷᳃┴䛷䛣 䜜䜎䛷グ㘓䛥䜜䛶䛔䜛䜹䜶䝹㢮䛾䛩䜉䛶 (ྥᒣ 2003) 䜢ྵ䜣䛷䛔䜛. 䛣䜜䜙䛾䜹䜶䝹㢮䛿,

⮬ศ䛷᥇㞟䛧䛯䜒䛾䛸༠ຊ⪅䛛䜙ᥦ౪䜢ཷ䛡䛯䜒䛾䜢ྵ䜣䛷䛚䜚, ᥦ౪䜢ཷ䛡䛯䜒䛾䛾୰䛻 䛿, 䜹䜶䝹䛾ಶయ඲య䜢ᥦ౪䛥䜜䛯䜒䛾䛾௚䛻, ᦬ฟ䛧䛯ᾘ໬⟶䛰䛡䛾ᥦ౪䜢ཷ䛡䛯䜒䛾䛜 17 ಶయྵ䜎䜜䛶䛔䜛.

ᐤ⏕⹸䛾᳨ᰝ᫬䛻䛿, 䜎䛪, 䜹䜶䝹㢮䜢䝆䜶䝏䝹䜶䞊䝔䝹䜢ධ䜜䛯ᐦ㛢ᐜჾ䛾୰䛷㯞㓉 䛧, ື䛛䛺䛟䛺䛳䛯ᚋ䛻ྲྀ䜚ฟ䛧, య㛗䜢䝕䝆䝍䝹䝜䜼䝇䛷 ᐃ䛧䛯. 䛣䛾䛸䛝, య㛗䛿࿀➃䛛 䜙⥲᤼ฟ⭍䜎䛷䛾┤⥺㊥㞳䛸䛧䛯. 䛭䛾ᚋ, య⾲䛾ᐤ⏕⹸䜢ㄪ䜉, ゎ๗─䛾ୖ䛷ゎ๗䜢⾜

䛳䛯. ⫵, ᚰ⮚, 㣗㐨䛛䜙⥲᤼ฟ⭍䛻⮳䜛ᾘ໬⟶, ⭈⮚ཬ䜃⭤⬔, ⏕Ṫ⭢䜢ྲྀ䜚ฟ䛧, 䛭䜜 䛮䜜ᐇయ㢧ᚤ㙾ୗ䛷ษ䜚㛤䛔䛶, ᐤ⏕⹸䜢᳨ᰝ䛧, ぢ䛴䛛䛳䛯ሙྜ䛿඲䛶ᅇ཰䛧䛶ィᩘ䛧 䛯. ෆ⮚䜢ྲྀ䜚ฟ䛧䛯ᚋ䛾య⭍䛷䜒ᐤ⏕⹸䜢ㄪ䜉䛯. ྲྀ䜚ฟ䛧䛯ᐤ⏕⹸䛿, ୍㒊䜢ᅽᖹᶆᮏ

䛸䛧, ṧ䜚䛿 70%䛒䜛䛔䛿 100%䜶䝍䝜䞊䝹䛷ಖᏑ䛧䛯. ௒ᅇᢅ䛳䛯ᐤ⏕⹸䛿“⽸⹸”䛻䛒䛯䜛

ᚋ⏕ື≀䛻㝈ᐃ䛧䛯. ཎ⏕ື≀䛿グ㘓䛩䜛䛻䛸䛹䜑ᅇ཰䛿⾜䜟䛺䛛䛳䛯.

ᐤ⏕⹸䛾ほᐹ䛾䛯䜑䛻, ௨ୗ䛾ᡭ㡰䛷, 㕲䝦䝬䝖䜻䝅䝸䞁䛷ᰁⰍ䛧䛯ᅽᖹᶆᮏ䜢సᡂ䛧䛯.

䜎䛪, ⦪ 2mm, ᶓ 20mm 䛾⣬∦ 2 ᮏ䜢䝇䝷䜲䝗䜺䝷䝇䛾ୖ䛻ᶆᮏ䛾䝃䜲䝈䜘䜚䜔䜔ᗈ䛔㛫㝸

䛷⨨䛝, 䛭䛾㛫䛻ᑡ㔞䛾Ỉ䜢⁲ୗ䛧ᐤ⏕⹸䜢⨨䛝, 䜹䝞䞊䜺䝷䝇䜢㍕䛫䛯. ᐇయ㢧ᚤ㙾䛷ᶆ ᮏ䛾≧ែ䜢☜ㄆ䛧䛺䛜䜙, ᶆᮏ䛜ᆒ➼䛻ᅽᖹ䛥䜜䜛䜘䛖䛻ᡭⱁ⣒䜢䜹䝞䞊䜺䝷䝇䛤䛸䝇䝷䜲䝗 䜺䝷䝇䛻ᕳ䛝௜䛡䛯. ᅽᖹ䛧䛯ᶆᮏ䛿䛭䛾䜎䜎 70%䜶䝍䝜䞊䝹䛻ᾐ䛧䛶ᅛᐃ䛧䛯. ᶆᮏ䛾ᰁ Ⰽ䛻䛒䛯䛳䛶䛿, ᅽᖹᶆᮏ䛾⣒䜢ษ䛳䛶䜹䝞䞊䜺䝷䝇䜢ྲྀ䜚㝖䛝, ⹸య䛰䛡䜢ᙇ䜚௜䛡䛯≧

ែ䛷䝇䝷䜲䝗䜺䝷䝇䜢Ỉ䛷Ὑ䛔 2. 5%㕲䝭䝵䜴䝞䞁Ỉ⁐ᾮ䛾୰䛻 12 ᫬㛫ධ䜜䛯. 䛣䜜䜢⵨␃Ỉ 䛷୎ᑀ䛻Ὑ䛳䛯ᚋ䛻, ಸ㔞ᕼ㔘䛾䝦䝬䝖䜻䝅䝸䞁⁐ᾮ䛻 24 ᫬㛫䡚48 ᫬㛫ධ䜜䛶ᰁⰍ䛧, ෌

䜃 2. 5%㕲䝭䝵䜴䝞䞁Ỉ⁐ᾮ䛻ධ䜜, ᘚⰍ䜢⾜䛳䛯. 24 ᫬㛫䡚1 㐌㛫䛷ᘚⰍ䛜᏶஢䛧䛯䜙, 30

ศ䡚60 ศ㛫ὶỈὙί䜢⾜䛳䛯䛾䛱, 70%, 80%, 90%, 95%, 100%, 100%䜶䝍䝜䞊䝹䛾㡰䛻䛭

䜜䛮䜜 15 ศ௨ୖᾐ䛧⬺Ỉ䛧䛯. ⬺Ỉ䜢᏶஢䛧䛯ᶆᮏ䛿䝺䝰䝌䞊䝹䛻 30 ศ௨ୖᾐ䛧䛶㏱ᚭ䛧,

䝞䝹䝃䝮䛒䜛䛔䛿䝨䝹䝬䜴䞁䝖䛷ໟᇙ䛧䛶䜹䝞䞊䜺䝷䝇䜢䛛䛡䛯. 䛣䜜䜢 50Υ䛻タᐃ䛧䛯䝠䞊 䝍䞊䛻ᨺ⨨䛧䛶ໟᇙ๣䜢ᅛ䜑䛯.

ຍ䛘䛶, ௒ᅇ᥇㞟䛧䛯ᐤ⏕⹸䛾୍㒊䛿㑇ఏᏊゎᯒ䛾䛯䜑䛻 100%䜶䝍䝜䞊䝹䛷ᅛᐃ䛧䛶 ෭෾ಖᏑ䛧䛯. 䛣䜜䜙䛾ᶆᮏ䛿᪫ᕝ་⛉኱Ꮫᐤ⏕⹸Ꮫㅮᗙ䛾୰ᑿ⛱ඛ⏕䛸⁠㈡┴❧኱Ꮫ

⎔ቃ⛉Ꮫ㒊䛾ᾆ㒊⨾బᏊඛ⏕䛾◊✲ᐊ䛷 DNA 䛾ሷᇶ㓄ิ䛾ゎᯒ䛜⾜䜟䜜䛯. 㑇ఏᏊ䝬 䞊䜹䞊䛸䛧䛶䛿, 䝭䝖䝁䞁䝗䝸䜰 DNA 䝏䝖䜽䝻䞊䝮䜸䜻䝅䝍䞊䝊䝃䝤䝴䝙䝑䝖 1 (cox1) 䛾 648BP 㡿ᇦ䛸᰾ 28S rDNA 䛾ሷᇶ㓄ิ䜢⏝䛔䛯.

ᙧែ䛾ㄝ᫂䛾䛯䜑䛾ᅗ䛷⏝䛔䛯␎グྕ䛿௨ୗ䛾䛸䛚䜚䛷䛒䜛.

a, anus (⫠㛛) ; bc, buccal cavity (ཱྀᄞ) ; br, bursal ray (⫘) ; bu, bursa (஺᥋ᄞ) ; cg, cement glands (䝉䝯䞁䝖⭢) ; ci, cephalic vesicle (㢌⬊) ; cp, caudal sucker (ᚋ྾┙) ; e, egg (༸) ; es, esophagus (㣗㐨) ; ex, excretory pore (᤼ἥᏍ) ; ey, eye (║) ; gs, genital spine (⏕

ṪᲲ) ; gu, gubernaculum (๪஺᥋่) ; in, intestine (⭠) ; l, lemniscus (ᆶᲨ) ; lh, large hook (኱㖋) ; m, mouth (ཱྀ) ; n, neck (㢕㒊) ; nr, nerve ring (⚄⤒⎔) ; o, ovary (༸ᕢ) ; ob, oesophageal bulb (㣗㐨⌫) ; od, oral diverticlum (྾ᄞ) ; op, opisthaptor (ᢕᥱჾ) ; os, oral sucker (ཱྀ྾┙) ; p,

(ဗ㢌) ; pp, prepharynx (๓ဗ㢌) ; pr,

(࿀) ; pre, proboscis receptacle (࿀㠧) ; s,

(྾┙) ; sap, Saefftigen’s pouch (䝉䝣䝔䜱䝀䞁Ặᄞ) ; sh, small hook (ᑠ㖋) ; su, spicle (஺᥋่) ; t, testis (⢭ᕢ) ; tr, trunc (⬗) ; u, uterus (Ꮚ ᐑ) ; v, vitellarium (༸㯤⭢) ; va, vagina (⭼) ; ve, vestibule (๓ᗞ) ; vs, ventral sucker (⭡

྾┙) ;㻌 vu, vulva (㝜㛛) .

4. ⤖ᯝ

4-1 ほᐹ䛥䜜䛯ᐤ⏕⹸䛾䝸䝇䝖

௒ᅇ䛾◊✲䛻䜘䛳䛶, 㟷᳃┴䜢୰ᚰ䛸䛧䛯໭ᮾ໭䛻ศᕸ䛩䜛ᅾ᮶䛾䜹䜶䝹㢮 9 ✀㢮䛛䜙, ᡥᙧື≀㛛 14 ✀, 㖋㢌ື≀㛛 2 ✀, ⥺ᙧື≀㛛 6 ✀, ⎔ᙧື≀㛛 2 ✀䛾ィ 24 ✀䛾ᐤ⏕

⹸䛜☜ㄆ䛥䜜䛯. ௨ୗ䛻, ฟ⌧✀䛾䝸䝇䝖䜢♧䛩. 㧗ḟศ㢮య⣔䛿 Uchida and Ooi (2018)

䛚䜘䜃 Uchida et al. (2019) 䛻䛧䛯䛜䛳䛯. ࿴ྡ䛾䛒䜛ศ㢮⩌䛻㝈䛳䛶࿴ྡ䜢௜䛧䛯.

Phylum Platyhelminthes㻌 ᡥᙧື≀㛛㻌 Class Monogenea㻌 ༢⏕⥘

Order Polyopisthocotylea㻌 Family Polystomatidae㻌

Diplorchis ranae Ozaki, 1931㻌㻌

Polystoma ozakii㻌 Price, 1939

Class Trematoda㻌 ྾⹸⥘

Order Plagiorchiida㻌 Family Hemiuridae

Halipegus japonicus Yamaguti, 1936 Family Paramphistomidae㻌 ཮ཱྀ྾⹸⛉

Diplodiscus japonicas Yamaguti, 1936 Diplodiscus sp.

Family Gorgoderidae㻌 䝂䝹䝂䝕䝷⛉

Gorgodera sp.

Gorgoderina tanigawaensis Uchida et Itagaki, 1974 Family Brachycoeliidae

Mesocoelium brevicaecum Goto et Ozaki, 1929 Mesocoelium sp.

Brachycoeliidae sp.

Family Haematoloechidae㻌

Haematoloechus japonicus (Yamaguti, 1936) Family Omphalometridae

Opisthioglyphe japonicus Uchida, Itagaki et Inoue, 1980 Family Pleurogenidae

Pleurolobatus lobatus (Ozaki, 1926) Cestoda㻌 ᮲⹸⥘

Cestoda sp.

Phylum Acanthocephala㻌 㖋㢌ື≀㛛 Class Palaeacanthocephala㻌 ྂ㖋㢌⹸⥘

Order Echinorhynchida㻌 䝁䜴䝖䜴䝏䝳䜴┠

Family Echinorhynchidae㻌 䝖䝀䝝䝅䝁䜴䝖䜴䝏䝳䜴⛉

Pseudoacanthocephalus toshimai Nakao, 2016㻌㻌 䜶䝌䜹䜶䝹㖋㢌⹸

Acanthocephala sp.

Phylum Nematoda㻌 ⥺ᙧື≀㛛 Class Chromadorea

Order Rhabditida Family Cosmocercidae

Cosmocerca japonica Yamaguti, 1938 Cosmocercoides pulcher Wilkie, 1930 Family Rhabdiasidae㻌 ᜷⥺⹸⛉

Rhabdias sp. 1

Rhabdias sp. 2 Family Trichostrongylidae

Oswaldocruzia sp.

Nematoda sp.

Phylum Annelida㻌 ⎔ᙧື≀㛛 Class Clitellata ⎔ᖏ⥘

Order Rhynchobdellida

Family Glossiphoniidae 䜾䝻䝅䝣䜷䝙⛉

Glossiphoniidae sp. 1 Glossiphoniidae sp. 2

4-2. ྛ✀䛾グ㍕䛸ฟ⌧≧ἣ

௨ୗ䛻, ᮏ◊✲䛷☜ㄆ䛥䜜䛯ᐤ⏕⹸䛻䛴䛔䛶, ✀䛤䛸䛻, ศ㢮Ꮫⓗ఩⨨, ᶆᮏ᝟ሗ (᪥

௜, ⏘ᆅ, ᐟ୺, ᐟ୺䛾㞤㞝, ᐤ⏕㒊఩, ᥇㞟⪅) , ᐟ୺䛸ᐤ⏕㒊఩䛚䜘䜃ᐤ⏕ᩘ, ᙧែ

䛾グ㍕, ศ㢮Ꮫⓗ⪃ᐹ䜢グ䛧䛯. ᐤ⏕ᩘ䛿, Bush et al. (1997) 䛾 intensity, 䛴䜎䜚, ⿕ᐤ⏕

ᐟ୺䛷䛾ᐤ⏕⹸䛾ಶయᩘ䛸䛧䛯.

Phylum Platyhelminthes㻌 ᡥᙧື≀㛛 Class Monogenea㻌 ༢⏕⥘

Order Polyopisthocotylea Family Polystomatidae

Diplorchis ranae Ozaki, 1931 ᅗ 1

Diplorchis ranae Ozaki, 1931a, p. 181-184, textfig. p. 182.

Diplorchis ranae Ozaki. Ozaki, 1935a, p. 196-213, fig. 5-17, pls. 1-3. Ozaki, 1935b, p. 16-17.

Yamaguti, 1936, p. 552. figs. 1-2.

ほᐹಶయ

2 ಶయ, 2018 ᖺ 7 ᭶ 18 ᪥, ⛅⏣┴䛻䛛䜋ᕷ, 䝒䝏䜺䜶䝹 2 ಶయ (㞝 1 㞤 1) , ⭤⬔, ụ

⏣⣫ኈ᥇㞟. 1 ಶయ, 2019 ᖺ 5 ᭶ 25 ᪥, 㟷᳃┴῝ᾆ⏫༑஧†ඵᬒ䛾ụὶฟἙᕝ, 䝒䝏䜺 䜶䝹 1 ಶయ (㞝 1) , ⭤⬔, ᑠ㔝᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䝒䝏䜺䜶䝹䛾⭤⬔䛻䜏䜙䜜, ᐤ⏕ᩘ䛿 1-2 䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿⦪䛻㛗䛟, ඛ➃䛛䜙ᢕᥱჾ䛾ᮎ➃䜎䛷䛾㛗䛥䛿 2. 7-3. 6 mm, ᖜ䛿 0. 8 mm (ᅗ 1A) .

⹸ཱྀ䛿య䛾ඛ➃䛻㛤䛟. ᢕᥱჾ䛿㛗䛥 350 μm, ᖜ䛿⹸య䛸䜋䜌ྠᵝ䛷 0. 8 mm 䛸ᶓ㛗䛷, 6 ಶ䛾྾┙䛜୪䜣䛷䛔䜛. ᭱ᚋ᪉䛾୰ኸᐤ䜚䛻఩⨨䛩䜛 2 ྾┙䛾㛫䛻, ᇶ㒊䛛䜙ᮎ➃䜎䛷䛾

┤⥺䛾㛗䛥䛜 0. 26-0. 29 mm 䛾 1 ᑐ䛾኱㖋䜢䜒䛴. 䛭䜜䛮䜜䛾኱㖋䛿ᖜᗈ䛔ᇶ㒊䛻㝯㉳䜢 䜒䛱, ᇶ㒊ഃ䛻❑䜏䛒䜛䛔䛿 V Ꮠᆺ䜔 U Ꮠᆺ䛾ษ䜜㎸䜏䜢ᣢ䛴 (ᅗ 1B) . ኱㖋䛾ᇶ㒊䛾ษ 䜜㎸䜏䛿, ኱㖋䛾㛗䛥䛾 5 ศ䛾 1 䛛䜙 3 ศ䛾䠍⛬ᗘ䜎䛷䛒䜛. ኱㖋䛿, ᚋ᪉䛻ྥ䛛䛳䛶ᖜ䛜

⊃䜎䜚, ഃ᪉䛻᭤䛜䜚, ᮎ➃䛻㛗䛥 53-58 μm 䛾㗦䛔∎䜢ᣢ䛳䛶䛔䜛. ဗ㢌䛿➽⫗㉁䛷, ဗ㢌 䛾䜔䜔ᚋ᪉䛻⏕ṪᏍ䛜㛤䛟. ⏕ṪᏍ௜㏆䛻䛿⏕ṪᲲ䛿㆑ู䛷䛝䛺䛛䛳䛯. ⭼Ꮝ䛿㆑ู䛷䛝 䛺䛛䛳䛯. ༸ᕢ䛿 100-150 μm 䛷, ๓➃䛛䜙 5 ศ䛾 2 ⛬ᗘ䛾๓᪉㒊䛾ഃ᪉䛻఩⨨䛩䜛. ⢭ᕢ 䛿㆑ู䛷䛝䛺䛛䛳䛯. ༸䛿⣳㗽ᙧ䛷㛗䛥 100 μm 䛷, 58-131 ಶ䛜య⭍䛾⭠⟶䜘䜚ෆഃ䜘䜚䛻 ᐦ㞟䛧䛶䛔䛯. ⭠⟶䛿ဗ㢌䛾䛩䛠ᚋ᪉䛛䜙ᕥྑ䛻ศ䛛䜜యഃ䛻ἢ䛳䛶, ᯞศ䛛䜜䛫䛪䛻⣽

䛛䛟ᒅ᭤䛧䛺䛜䜙ᚋ᪉䛻ఙ䜃, ᅛ╔ჾ䛾๓᪉㒊⫼㠃䛷⧅䛜䜛. ᾘ໬⟶䛻䛿≉ᐃ䛾䜎䛸䜎䜚䛿 䜏䜙䜜䛪, ┣ᄞ䛿䛺䛔.

≉グ஦㡯

᪥ᮏ䛷䛿, Diplorchis ᒓ䛸䛧䛶 Diplorchis ranae 䛾 1 ✀䛜グ㘓䛥䜜䛶䛔䜛䛾䜏䛷䛒䜛 (Ozaki

1931a; ᪂⏣, 2018) . ௒ᅇほᐹ䛧䛯ᶆᮏ䛾኱㖋䛾ᇶ㒊䛛䜙ᮎ➃䜎䛷䛾┤⥺䛾㛗䛥䛿 0. 26-0.

29 μm 䛷, Ozaki (1935a) 䛻䜘䜛 0. 28-0. 33 mm 䛸኱䛝䛺㐪䛔䛿䛺䛛䛳䛯. 䜎䛯, ௒ᅇほᐹ䛧䛯 ᶆᮏ䛾∎䛾㛗䛥 53-58 μm 䜒 Ozaki (1935a) 䛷䛾 50-58 μm 䛸䜋䜌ྠᵝ䛷䛒䛳䛯. 䛥䜙䛻, ⭠

⟶䛾ᙧែ䜒 Ozaki (1935a) 䛾グ㏙䛸㐪䛔䛜䛺䛛䛳䛯. 䛣䜜䜙䛾ほᐹ⤖ᯝ䛻ᇶ䛵䛔䛶, ௒ᅇほ ᐹ䛧䛯ᶆᮏ䜢 D. ranae 䛸ྠᐃ䛧䛯.

୍᪉䛷, ௒ᅇほᐹ䛧䛯ಶయ䛾య㛗䛿 2. 7-3. 6 mm 䛷, Ozaki (1935a) 䛾 4. 25-6. 5 mm 䛻 ẚ䜉䜛䛸⣙༙ศ⛬ᗘᑠᆺ䛷䛒䛳䛯. Ozaki (1935a) 䛻䜘䜛䛸, ኱㖋䛾ษ䜜㎸䜏䛿ಶయኚ␗䛜 㠀ᖖ䛻኱䛝䛟, 䜋䛸䜣䛹䛝䜜䛣䜎䛺䛔䜒䛾䛛䜙䜋䜌༙ศ⛬ᗘ䜎䛷 V Ꮠ≧䛻ษ䜜㎸䜐ಶయ䜔, Ἴᡴ䛳䛯䜚୙つ๎䛻ษ䜜㎸䜐ಶయ䛜グ㏙䛥䜜䛶䛔䜛. ௒ᅇほᐹ䛧䛯 2 ಶయ䛾ษ䜜㎸䜏䛿䛹

䛱䜙䜒 Ozaki (1935a) 䛜グ㍕䛧䛶䛔䜛ಶయኚ␗䛾⠊ᅖ䛻཰䜎䛳䛶䛔䛯.

ศᕸ䛸ᐟ୺

᪂⏣ (2018) 䜔 Uchida and Ooi (2018) 䛻䜘䜛䛸, D. ranae 䛾䛣䜜䜎䛷䛾ᐟ୺䛿䛩䜉䛶䝒䝏 䜺䜶䝹䛷, ᮏᕞ, ᅄᅜ, ஑ᕞ䛛䜙グ㘓䛥䜜䛶䛔䜛. 䛣䜜䜎䛷ᮾி䜘䜚䜒໭᪉䛛䜙䛿グ㘓䛥䜜䛶 䛔䛺䛔䛯䜑, ௒ᅇ䛾㟷᳃┴䛛䜙䛾グ㘓䛿 D. ranae 䛾᭱໭䛾グ㘓䛸䛺䜛. ᐟ୺䛿䛣䜜䜎䛷ྠ

ᵝ䝒䝏䜺䜶䝹䛷䛒䛳䛯.

Polystoma ozakii Price, 1939

ᅗ 2-3

Polystoma integerrimum (

. ᑿᓮ, 1931b, p. 23. Ozaki, 1935a, 194-195, figs.

1-3.

Polystoma ozakii Price, 1939, p. 82.

Polystoma ezoensis Uchida, Machida, Uchida et Itagaki, 1988, 9-11, figs. 5-6.

ほᐹಶయ

4 ಶయ, 2017 ᖺ 4 ᭶ 14 ᪥, ᘯ๓ᕷᆏඖ, 䝲䝬䜰䜹䜺䜶䝹 2 ಶయ (㞝 1 㞤 1) , ⭤⬔, ▼㒓 ᒸ༓ḷ㡢᥇㞟. 1 ಶయ, 2017 ᖺ 5 ᭶ 31 ᪥, ᘯ๓ᕷᒾᮌᒣᘺ⏕Ꮚ䛹䜒䛾ᗈሙ, 䝲䝬䜰䜹䜺䜶 䝹 1 ಶయ (㞝 1) , ⭤⬔, ụ⏣⣫ኈ᥇㞟. 2 ಶయ, 2018 ᖺ 4 ᭶ 13 ᪥, ᘯ๓ᕷᆏඖ, 䝲䝬䜰䜹 䜺䜶䝹 1 ಶయ (ND1) , ⭤⬔, ▼㒓ᒸ༓ḷ㡢䞉኱㧗᫂ྐ᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䝲䝬䜰䜹䜺䜶䝹䛾⭤⬔䛻䜏䜙䜜, ᐤ⏕ᩘ䛿, 1-4 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿⦪䛻㛗䛟, ඛ➃䛛䜙ᢕᥱჾᮎ➃䜎䛷䛾㛗䛥䛿 7. 1-9. 5 mm (ᅗ 2A) . ཱྀ䛿⹸య䛾ඛ➃ 䛻㛤䛟. ᢕᥱჾ䛿 6 ಶ䛾྾┙䜢䜒䛴. ᭱ᚋ᪉䛾 2 ྾┙䛾㛫䛻, ᫂░䛺ษ䜜㎸䜏䛾䛒䜛 2 ཫ ᆺ䛷ᇶ㒊䛻㝯㉳䛜䛒䜛 1 ᑐ䛾኱㖋䜢䜒䛴 (ᅗ 2B) . ኱㖋䛿๓᪉㒊䛜 V Ꮠᆺ䛒䜛䛔䛿 U Ꮠᆺ 䛻኱䛝䛟ษ䜜㎸䜏, ᚋ᪉䛻䛛䛡䛶ᖜ䛜ᚎ䚻䛻⊃䜎䜛䛸䛸䜒䛻ഃ᪉䛻᭤䛜䜚, ඛ➃䛻㗦䛔∎䜢 䜒䛴. ኱㖋䛾ᇶ㒊䛾ษ䜜㎸䜏䛿, ඲㛗䛾 3 ศ䛾䠍䛛䜙༙ศ⛬ᗘ䜎䛷䛷䛒䜛. ኱㖋䛾㛫䛻, ୍ ᑐ䛾ᑠ㖋䜢ᣢ䛴ಶయ (ᅗ 3A) 䜒䛒䛳䛯. ᑠ㖋䛿Წ≧䛷ඛ➃䛜඲㛗䛾༙ศ䛻䛒䛯䜛఩⨨䛷 ศ䛛䜜䛶䛚䜚, ඛ➃ഃ䛻ᘏ㛗䛧䛯㒊ศ䛿, ᇶ㒊ᐤ䜚䛾఩⨨䛷ศ䛛䜜䛯㒊ศ䛻ẚ䜉䛶ⴭ䛧䛟㛗 䛔. 㝜ⱼ䛾ඛ➃䛻㛗䛥 20-25 μm 䛷‴᭤䛧䛯 9 ᮏ䛾⏕ṪᲲ (ᅗ 2C) 䜢ᣢ䛴. ⭼Ꮝ䛿➽⫗㉁

䛷, య๓➃䛛䜙య㛗䛾 1/5 䛾ᕥྑ୧ഃ䛻㛤䛟. Ꮚᐑ䛿༸ᕢ䜘䜚య䛾ඛ➃䜘䜚䛾ሙᡤ䛻䛒䜛.

⢭ᕢ䛿㆑ู䛷䛝䛺䛛䛳䛯. ほᐹ䛧䛯ᶆᮏ䛷䛿, ༸䛿䛣䜜䜢ᣢ䛴ಶయ䛷䜒 4-6 ಶ䛸ᑡ䛺䛟, ಶ య䛻䜘䛳䛶䛿༸䛜඲䛟ぢ䜙䜜䛺䛔ሙྜ䜒䛒䛳䛯. ⭠䛿⹸య඲య䛻䜟䛯䛳䛶୍ᵝ䛻」㞧䛺ᯞศ 䛛䜜䛜䜏䜙䜜, 䜎䛯, ┣ᄞ䜒ከᩘ䜏䜙䜜䛯. ⭠⟶䛾ᯞศ䛛䜜䛿䜎䛸䜎䛳䛯༢఩䜢స䜛䛣䛸䛿䛺 䛛䛳䛯.

ኚ␗

኱㖋䛾ᇶ㒊䛾ษ䜜㎸䜏䛾⛬ᗘ䛿ಶయ䛻䜘䛳䛶኱䛝䛟␗䛺䛳䛶䛔䛯 (ᅗ 3) . 䛧䛛䛧, ᭱䛾 ὸ䛔ሙྜ䛷䜒 5 ศ䛾 1 ⛬ᗘ, ᭱䜒ษ䜜㎸䜣䛷䛔䛶䜒༙ศ䜢㉺䛘䜛䛣䛸䛿䛺䛛䛳䛯. ኱㖋䛿㏻

ᖖᮎ➃䛻∎䜢ᣢ䛳䛶䛔䛯䛜, ほᐹᶆᮏ䛾 1 ಶయ䛿∦ഃ䛾኱㖋䛷∎䜢Ḟ䛔䛶䛔䛯. 䜎䛯, ほ ᐹᶆᮏ඲ 6 ಶయ䛾䛖䛱 1 ಶయ䛷䛿, ኱㖋䛸ᚋ྾╔ჾ䛾ᚋ⦕䛾㛫䛻 2 ᑐ䛾ᑠ㖋䜢ᣢ䛳䛶䛔䛯.

⏕ṪᲲ䛿, ほᐹᶆᮏ 7 ಶయ୰ 3 ಶయ䛷 9 ᮏ䛒䜛䛣䛸䛜☜ㄆ䛥䜜䛯. ୍᪉䛷, ほᐹᶆᮏ䛾䛖䛱 4 ಶయ䛷䛿୍㒊䛜୙᫂░䛷䛒䜚, ⏕ṪᲲ䛾ᮏᩘ䜢㆑ู䛩䜛䛣䛸䛜䛷䛝䛺䛛䛳䛯.

≉グ஦㡯

䝫䝸䝇䝖䝬┠䛿⹸య䛾ඛ➃䛻䛿྾┙䛜䛺䛟, ཱྀ䛜㛤䛔䛶䛔䜛 (Price 1939) . ௒ᅇほᐹ䛧䛯

ᶆᮏ䛷䜒, ඛ➃䛻྾┙≧䛾ჾᐁ䛿䜏䜙䜜䛪, ཱྀ䛜ぢ䜙䜜䛯䛯䜑䝫䝸䝇䝖䝬┠䛸ุ᩿䛧䛯. 䜎䛯,

䝫䝸䝇䝖䝬ள⛉䛿, ᢕᥱჾ䛻 6 䛴䛾྾┙䛜䛒䜛 (Price 1939) . ௒ᅇほᐹ䛧䛯ᶆᮏ䛾඲䛶䛷,

௜╔┙䛻 6 䛴䛾྾┙䛜䜏䜙䜜䛯䛯䜑, 䝫䝸䝇䝖䝬ள⛉䛸ุ᩿䛧䛯. 䛥䜙䛻, Polystoma ᒓ䛿 1)

」ᩘ䛾⭼䜢䜒䛴, 2) Ꮚᐑ䛿༸ᕢ䛾๓䛻䛒䜛, 3) 」ᩘ䛾⢭ᕢ䜢ᣢ䛴, 䛒䜛䛔䛿 1 䛴䛾⢭ᕢ䜢 ᣢ䛴, 4) Ꮚᐑ䛿┦ᑐⓗ䛻㛗䛟, ከ䛟䛾༸䜢ᣢ䛱, 」ᩘ䛾⢭ᕢ䜢ᣢ䛴䛸䛔䛖≉ᚩ䛜䛒䜛 (Price

1939) . ௒ᅇほᐹ䛧䛯ᶆᮏ䛿 2 䛴䛾⭼䛜䛒䜚, Ꮚᐑ䛿༸ᕢ䜘䜚ඛ➃ᐤ䜚䛾㒊఩䛻䛒䜚, 4-6 ಶ

䛾༸䛜䛒䛳䛯䛣䛸䛛䜙, ☜ㄆ䛷䛝䛶䛔䛺䛔⢭ᕢ䜢㝖䛟䛸䛔䛪䜜䜒 Price (1939) 䛾ᐃ⩏䛻ヱᙜ䛧 䛯䛯䜑, Polystoma ᒓ䛸ุ᩿䛧䛯.

᪥ᮏ䛷䜹䜶䝹㢮䜢⤊ᐟ୺䛸䛧䛶ሗ࿌䛥䜜䛶䛔䜛༢⏕㢮䛿䛣䜜䜎䛷, Diplorchis ranae, Polystoma integerrimum, Polystoma ozakii, Polystoma racophori 䛾 4 ✀㢮䛜▱䜙䜜䛶䛔䜛 (Yamaguti 1936; Price 1939; Uchida et al. 1988; Goldberg and Bursey 2002) . 䛣䛾䛖䛱 3 ✀䛜 Polystoma ᒓ䛻ᒓ䛧䛶䛔䜛. ௒ᅇほᐹ䛧䛯ᶆᮏ䜢䛣䜜䜎䛷▱䜙䜜䛶䛔䜛 Polystoma ᒓ䛾 3 ✀䛸 ẚ㍑䛩䜛. 䜎䛪, ᮏ✀䛾య㛗䛿 7. 1-9. 5 mm 䛷, P. racophori 䜘䜚䜒᫂䜙䛛䛻኱ᆺ䛷, 䛛䛴 P.

integerrimum 䜘䜚᫂䜙䛛䛻ᑠᆺ䛷䛒䜛. 䛭䜜䛻ᑐ䛧, P. ozakii 䛾య㛗䛾⠊ᅖ䛸኱䛝䛟㔜」䛧䛶 䛔䜛. ᮏ✀䛾⭠⟶䛾ᵓ㐀䛿, 」㞧䛻ศᒱ䛧, 䜎䛸䜎䛳䛯༢఩䛿ぢ䛔䛰䛫䛺䛔Ⅼ䛷 P. ozakii 䛸

୍⮴䛩䜛. 䛣䛾Ⅼ䛿⭠⟶䛾ศᒱ䛜 3 ⴥ䛻ศ䛛䜜䜛 P. integerrimum 䛸䛿኱䛝䛟␗䛺䛳䛶䛔䜛.

ᮏ✀䛾኱㖋䛿, ኱㖋䛾඲㛗䛾 3 ศ䛾 1 䛛䜙༙ศ⛬ᗘ䜎䛷ษ䜜㎸䜐. 䛣䛾Ⅼ䛿, ༙ศ⛬ᗘ䛾 ษ䜜㎸䜏䜢ᣢ䛴 P. ozakii 䛾グ㏙䛸䜘䛟ఝ䛶䛔䜛. ௨ୖ䛾䜘䛖䛻, య㛗䜔⭠ᐁ䛾ᙧែ, ኱㖋䛾 ᙧ≧䛻䛴䛔䛶䛾≉ᚩ䛜୍⮴䛩䜛䛣䛸䛛䜙, ௒ᅇ䛾ほᐹᶆᮏ䜢 P. ozakii 䛸ྠᐃ䛧䛯.

୍᪉䛷, P. ozakii 䜢ྵ䜐䛣䜜䜎䛷᪥ᮏ䛛䜙グ㘓䛥䜜䛶䛔䜛 Polystoma ᒓ 3 ✀䛾⏕ṪᲲ䛿䛔 䛪䜜䜒 8 ᮏ䛷䛒䜛䛾䛻ᑐ䛧䛶, ᮏ✀䛷䛿 9 ᮏ䛾⏕ṪᲲ䛜ぢ䜙䜜䛯. ௒ᅇ䛾ほᐹ䛿ಶయᩘ䛜 ᑡ䛺䛔䛯䜑䛻⏕ṪᲲ䛾ᩘ䛾ኚ␗䛻䛴䛔䛶䛿୙᫂䛷䛒䜛.

ศᕸ䛸ᐟ୺

௒ᅇ䛾◊✲䛷, ᮏ✀䛿㟷᳃┴ᘯ๓ᕷ䛾 2 ᆅⅬ䛾䝲䝬䜰䜹䜺䜶䝹䛛䜙☜ㄆ䛥䜜䛯. Uchida et al. (1988) , Uchida and Ooi (1988) , ᪂⏣ (2018) 䛻䜘䜛䛸, P. ozakii 䛿䛣䜜䜎䛷, ໭ᾏ㐨, 㟷᳃┴, ᐑᇛ┴, ᮾி㒔, ⚄ዉᕝ┴, ᗈᓥ┴䛛䜙グ㘓䛥䜜䛶䛔䜛.

Class Trematoda㻌 ྾⹸⥘

Order Plagiorchiida Family Hemiuridae

Halipegus japonicus Yamaguti, 1936 ᅗ 4

Halipegus japonicus Yamaguti, 1936, p. 561-563, figs. 16-17.

ほᐹಶయ

1 ಶయ, 2018 ᖺ 7 ᭶ 15 ᪥, 㟷᳃┴ᘯ๓ᕷᆏඖ, 䝒䝏䜺䜶䝹 1 ಶయ (㞝 1) , య⭍, ▼㒓 ᒸ༓ḷ㡢䞉῝⃝ᖾᜨ᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䝒䝏䜺䜶䝹 1 ಶయ䛾య⭍䛛䜙䜏䜙䜜, ᐤ⏕ᩘ䛿 4 䛷䛒䛳䛯. 䛯䛰䛧, ゎ๗᫬䛾᭱ᚋ䛻ぢ䛴 䛡䛯䛯䜑, ᐇ㝿䛾ᐤ⏕㒊఩䛿య⭍䛷䛿䛺䛔ྍ⬟ᛶ䛜䛒䜛.

ᙧែ䛾グ㍕

య䛿⦪䛻㛗䛟, ඛ➃䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 6. 8 mm, ⹸య䛾୰ኸ䛾ᖜ䛿 2. 1mm 䛷, ཱྀ྾

┙䛾ඛ➃㒊ศ䛾ᖜ䛿 0. 58 mm, ᭱኱ᖜ䛿୰ኸ䜘䜚䜔䜔ᚋ᪉䛷 2. 3 mm (ᅗ 4) ⹸య䛾ඛ➃ 䛻⦪᪉ྥ䛻䜔䜔㛗䛔ཱྀ྾┙䛜䛒䜚, 㛗䛥 0. 72 mm, ᖜ 0. 65 mm. ⭡྾┙䛿⦪᪉ྥ䛻䜟䛪䛛 䛻㛗䛔෇ᙧ䛷㛗䛥 1. 1 mm ᖜ 1. 0 mm 䛷, 䜋䜌⹸య䛾୰ኸ䛻఩⨨䛩䜛. ဗ㢌䜔㣗㐨, ⭠䛿

㆑ู䛷䛝䛺䛛䛳䛯. 㝜ⱼᄞ䛿䜏䜙䜜䛺䛛䛳䛯. Ꮚᐑ䛿ᖜ 0. 20-0. 25 mm 䛾⟶≧䛷, ᒅ᭤䛧䛺 䛜䜙⹸య䛾ඛ➃䛸ᚋ➃䜢㝖䛟య⭍䜢ᗈ䛟༨䜑䜛. ⢭ᕢ䛿 2 ಶ䛷, ⭡྾┙䛾ᚋ᪉䛻ᶓ䛻୪䜣䛷

఩⨨䛩䜛. 䛹䛱䜙䛾⢭ᕢ䜒䜋䜌⌫ᙧ䛻㏆䛔ᴃ෇ᙧ䛰䛜, ⭡྾┙䛻㏆䛔᪉䛜䜔䜔኱䛝䛟, 㛗䛥 1. 0 mm ᖜ 0. 86 mm, 䜒䛖୍᪉䛿㛗䛥 0. 8 mm ᖜ 0. 72 mm. ༸ᕢ䛿 1 ಶ䛷, 㛗䛥 0. 35 mm,

ᖜ 0. 36 mm 䛾෇ᙧ䛷, ⹸య䛾ᚋ➃㏆䛟䛻఩⨨䛩䜛. ༸㯤⭢䛿༸ᕢ䛾ᚋ᪉䛻㞟୰䛧䛶䛚䜚,

」ᩘ䛾ⴥ≧䛾⤌⧊䛜ᕥྑ䛻䛔䛟䛴䛛䜎䛸䜎䛳䛶ᕥྑ䛻㓄⨨䛩䜛. ༸㯤⭢䛾ⴥ≧䛾ᵓ㐀䛿䛭 䜜䛮䜜㛗䛥䛜 0. 23-0. 36 mm, ᖜ䛜 0. 13-0. 23 mm 䛷䛒䜛.

≉グ஦㡯

ᮏ✀䛿, ⭡྾┙䛾఩⨨䜔኱䛝䛥, ⢭ᕢ䛾఩⨨, ༸ᕢ䛾఩⨨䜔኱䛝䛥, ༸㯤⭢䛾ᵓ㐀䛜 Yamaguti (1936) 䛜グ㍕䛧䛯 Halipegus japonicus 䛸୍⮴䛧䛯䛯䜑, ᮏ✀䜢 H. japonicas 䛸ྠ

ᐃ䛧䛯.

ศᕸ䛸ᐟ୺

ᮏ✀䛿໭ᾏ㐨䛛䜙୰㒊ᆅ᪉䜎䛷䛷, 4 ✀䛾䜹䜶䝹㢮 (䜰䝈䝬䝠䜻䜺䜶䝹, 䜴䝅䜺䜶䝹, 䝖䝜 䝃䝬䜺䜶䝹, 䝘䝂䝲䝎䝹䝬䜺䜶䝹) 䛛䜙グ㘓䛥䜜䛶䛔䜛 (Uchida and Ooi 2018) . ௒ᅇぢ䛴 䛛䛳䛯䝒䝏䜺䜶䝹䛿᪂ᐟ୺䛸䛺䜛.

Family Paramphistomidae

Diplodiscus japonicus 㻌㻌 Yamaguti, 1936 ᅗ 5

Diplodiscus amphichrus japonicus Yamaguti, 1936, p. 559-561, figs. 13-15.

Diplodiscus japonicus Yamaguti. ᕷᕝ, 1970, p. 189-190, ᅗ 108.

Diplodiscus japonicas Yamaguti. Li, 1937. Besprozvannykh et al. , 2018, p. 65-66,

tables. 2-3 figs. 1.

Diplodiscus subclavatus (Pallas, 1760) . 㷌஭ , 1923, p. 155-159, ᅗ 1-4.

ほᐹಶయ

3 ಶయ, 2017 ᖺ 4 ᭶ 24 ᪥, 㟷᳃┴ᘯ๓ᕷ๓ᆏ, 䜰䝈䝬䝠䜻䜺䜶䝹 1 ಶయ (㞤 1) , ᑠ⭠, ▼ 㒓ᒸ༓ḷ㡢᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䜰䝈䝬䝠䜻䜺䜶䝹䛾ᑠ⭠䛷䜏䜙䜜䛯. ᐤ⏕ᩘ䛿 5 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿⣽㛗䛔෇㗹ᙧ䛷, ඛ➃䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 2. 1 mm, ⹸య䛾୰ኸ䛷ᖜ䛿 0. 58 mm.

ཱྀ྾┙䛿⹸య䛾ඛ➃䛻఩⨨䛧, 㛗䛥 242 μm, ᖜ 212 μm 䛷, ྾ᄞ䜢䜒䛴. ᚋ྾┙䛿⹸య䛾 ᮎ➃䛻䛒䜚, ⴭ䛧䛟኱䛝䛟㛗䛥 400 μm, ᖜ 636 μm 䛷, ᗏ㠃୰ኸ䛻➽⫗㉁䛾㝯㉳䜢䜒䛴. ྾ᄞ 䛻⥆䛟㣗㐨䛿, 㛗䛥 252 μm, ᖜ 25 μm. 㣗㐨䛾ᚋ᪉䛻, 㛗䛥 111 μm, ᖜ 30 μm 䛷ᕥྑ୍ᑐ 䛾➽⫗㉁䛾ቨ䛻䛛䛣䜎䜜䛯ဗ㢌䛜䛒䜛. ⭠⟶䛿⹸య䛾ඛ➃䛛䜙 4 ศ䛾 1 䛾఩⨨䛷ᕥྑ䛻஧

ศᒱ䛧, ᚋ྾┙㏆䛟䜎䛷ఙ䜃, ஫䛔䛻᥋⥆䛫䛪䛻┣≧䛻䛺䛳䛶⤊䜟䜛. ༸㯤⭢䛿䛭䜜䛮䜜

㛗ᚄ 101 μm, ▷ᚄ 60. 6 μm 䛾༸෇ᙧ䛷, ဗ㢌䛾ഃ᪉䛛䜙ᚋ྾┙䛾๓䜎䛷䛾ᕥྑ䛻䠈⢭ᕢ

䜔༸ᕢ䛜ศᕸ䛩䜛㡿ᇦ䜢㑊䛡䜛䜘䛖䛻⣙ 20 ಶ䛪䛴ศᕸ䛩䜛. ⢭ᕢ䛿 1 ಶ䛷, 㛗ᚄ 101μm,

▷ᚄ 60. 6 μm 䛾༸ᙧ䛷, ⹸య䛾୰ኸ䜘䜚䜔䜔ᚋ᪉䛾ṇ୰⥺ୖ䛷, ᾘ໬⟶䛾ෆഃ䛻఩⨨䛩䜛.

༸ᕢ䛿 1 ಶ䛷, 㛗ᚄ 141 μm, ▷ᚄ 80. 8 μm 䛷, ᚋ➃䛛䜙 3 ศ䛾 1 䛻఩⨨䛩䜛.

≉グ஦㡯

Uchida and Ooi (2018) 䛻䜘䜛䛸, ᪥ᮏ䛾 Diplodiscus ᒓ䛿 Diplodiscus japonicus 䛾 1 ✀䛾 䜏䛷䛒䜛.㻌 ᮏ✀䛿, Yamaguti (1936) 䛾 Diplodiscus japonicus 䛸ẚ㍑䛩䜛䛸, య㛗䜔㣗㐨䛾 㛗䛥, ༸㯤⭢䛾ಶᩘ䜔㓄⨨, ⢭ᕢ䛾㓄⨨䜔኱䛝䛥䛿▩┪䛧䛺䛔. 䜘䛳䛶, ᮏ✀䜢 Diplodiscus japonicus 䛸ྠᐃ䛧䛯.

Diplodiscus japonicus 䛿 Yamaguti (1936) 䛜 Diplodiscus amphichrus 䛾୍ள✀䛸䛧䛶グ㍕

䛧䛯✀㢮䛷䛒䜛. 䛭䛾ᚋ, Li (1937) 䛿ᮏ✀䜢 D. amphichrus 䛛䜙⊂❧䛥䛫, D. japonicas 䛸䜏 䛺䛧䛯. 䛣䛾ᢅ䛔䛿, Pande (1937) , ྜྷ⏣䞉ᒣୗ (1965) , ෆ⏣ (1975) 㛗㇂ᕝ (2003) 䛺䛹 䛻ᨭᣢ䛥䜜䛶䛚䜚, ௒ᅇ䜒䛣䛾య⣔䛻㡰䛨䛯.

୍᪉, Singh (1954) 䛿, D. amphichrus var. magunus 䛸 D. mehrai 䜢 D. amphichrus 䛾䝅䝜 䝙䝮䛸䛧䛯. 䜎䛯, Mukherijee (1966) 䛿, D. japonicus 䜢 D. amphichrus 䛾䝅䝜䝙䝮䛸䜏䛺䛧䛯.

Fischthal and Thomas (1968) 䛿, D. amphichrus 䜢 D. magunus 䛾䝅䝜䝙䝮䛸䜏䛺䛧䛯. 䛧䛯䛜 䛳䛶, Fischthal and Thomas (1968) 䛾䜒䛸䛷䛿, D. japonicus 䜔 D. mehrai 䛿 D. magunus 䛾䝅 䝜䝙䝮䛸䛺䜛. Besprozvannykh et al. (2018) 䛿, ᴟᮾ䝻䝅䜰䛾ᡂయ䛸䝉䝹䜹䝸䜰䛾ᶆᮏ䜢౑䛳 䛶, Diplodiscus 䛾ᙧែ䜔⣔⤫䜢㆟ㄽ䛧, D. japonicus 䜢, D. mehrai 䜔 D. amphichrus 䛸䛿␗䛺 䜛⊂❧✀䛸䜏䛺䛧䛯. 䛣䛣䛷䛿, ྜྷ⏣䞉ᒣୗ (1965) 䜔 ෆ⏣ (1975) 㛗㇂ᕝ (2003) , Besprozvannykh et al. (2018) 䛾య⣔䛻䛧䛯䛜䛳䛶 D. japonicus 䜢⊂❧✀䛸䛧䛶ᢅ䛳䛯.

ศᕸ䛸ᐟ୺

ᮏ✀䛿᭷ᑿ㢮䛸↓ᑿ㢮䛻䜎䛯䛜䜛 14 ✀㢮䛾୧⏕㢮䛛䜙グ㘓䛥䜜䛶䛔䜛 (Uchida and Ooi

2018) . ⏘ᆅ䜒ᮏᕞ䛸஑ᕞ䛾ᗈ䛔⠊ᅖ䛻䛚䜘䜆䛜, 䛣䜜䜎䛷䛾グ㘓䛾᭱໭䛿᪂₲┴䛺䛾䛷,

௒ᅇ䛾㟷᳃┴ᘯ๓ᕷ䛷䛾Ⓨぢ䛿, ໭㝈䛾᭦᪂䛸䛺䜛.

Diplodiscus sp.

ᅗ 6 ほᐹಶయ

1 ಶయ, 2017 ᖺ 8 ᭶ 1 ᪥, 㟷᳃┴ᘯ๓ᕷᆏඖ, 䝲䝬䜰䜹䜺䜶䝹 1 ಶయ (㞤 1) , ᑠ⭠, 㮵 ಛ㢧඾᥇㞟. 2 ಶయ, 2019 ᖺ 3 ᭶ 26 ᪥, 㟷᳃┴ᘯ๓ᕷᆏඖ, 䝲䝬䜰䜹䜺䜶䝹 1 ಶయ (㞝 1) , ᑠ⭠, ▼㒓ᒸ᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䝲䝬䜰䜹䜺䜶䝹䛾኱⭠䛷䜏䜙䜜䛯.ᐤ⏕ᩘ䛿 1-2 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿ᚋ᪉䛻䜅䛟䜙䜏䜢ᣢ䛴䛜䛚䛚䜐䛽෇㗹ᙧ䛷, ඛ➃䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 4. 57 mm,

⹸య䛾୰ኸ䛷ᖜ䛿 1. 65 mm, ᮎ➃䛛䜙 4 ศ䛾 1 䛷᭱኱ᖜ 1. 87 mm 䛻㐩䛩䜛 (ᅗ 6) . ཱྀ྾

┙䛿⹸య䛾ඛ➃䛻఩⨨䛧, ྾ᄞ䜢䜒䛴. ᚋ྾┙䛿⹸య䛾ᮎ➃䛻䛒䜚, ⴭ䛧䛟኱䛝䛟㛗䛥

0. 3mm, ᖜ 1. 3 mm 䛷, ᗏ㠃୰ኸ䛻㝯㉳䜢ᣢ䛴. ྾ᄞ䛻⥆䛟㣗㐨䛿, 㛗䛥 300 μm. 㣗㐨䛾

ᚋ᪉䛻, 㛗䛥 111 μm, ᖜ 30 μm 䛷ᕥྑ୍ᑐ䛾➽⫗㉁䛾ቨ䛻䛛䛣䜎䜜䛯㣗㐨⌫䛜䛒䜛. ⭠⟶

䛿⹸య䛾ඛ➃䛛䜙 5 ศ䛾 1 䛾఩⨨䛷ᕥྑ䛻஧ศᒱ䛧, ᚋ྾┙㏆䛟䜎䛷ఙ䜃, ┣≧䛻䛺䛳䛶

⤊䜟䜛. ༸㯤⭢䛿༸෇ᙧ䛷, ๓᪉㒊䛻ศᕸ䛩䜛䜒䛾䛜ᑠ䛥䛟, ᚋ᪉㒊䛷䛿༸ᕢ䛸ྠ⛬ᗘ䜎䛷

኱ᆺ䛻䛺䜚, 㣗㐨⌫䛾ഃ᪉䛛䜙ᚋ྾┙䛾㏆䛟䜎䛷ศᕸ䛩䜛. ⢭ᕢ䛿 1 ಶ䛷, ┤ᚄ 625 μm 䛾

⌫ᙧ䛷, ⹸య䛾๓ᚋ䛾୰ኸ䜔䜔ᕥ䜘䜚䛷, 2 ศᒱ䛧䛯⭠⟶䛾㛫䛻఩⨨䛩䜛. ༸ᕢ䛿 1 ಶ䛷,

㛗ᚄ 375 μm, ▷ᚄ 275 μm 䛾༸ᙧ䛷, ⢭ᕢ䛸䛿䛿䛺䜜䛶ᚋ➃䛛䜙 5 ศ䛾 2 䛻఩⨨䛩䜛.

≉グ஦㡯

ᮏ✀䛿๓㏙䛾 D. japonicus 䛻ẚ䜉䛶, య㛗䛜䛚䜘䛭 2 ಸ䛸኱ᙧ䛷䛒䜛Ⅼ䛜␗䛺䛳䛶䛔䜛. 䜎 䛯, D. japonicus 䛿⢭ᕢ䛜୰ኸ䜘䜚䜔䜔ᚋ᪉䛻఩⨨䛩䜛䛾䛻ᑐ䛧, ᮏ✀䛷䛿⹸య䛾୰ኸ䛻䛒 䜛Ⅼ䜒␗䛺䛳䛶䛔䜛. 䜎䛯, D. japonicus 䛷䛿, 㣗㐨䛾㛗䛥䛜 0. 33 mm 䛷, య㛗 (1. 5-2. 5 mm) 䛻ᑐ䛧䛶 22 䝟䞊䝉䞁䝖䛛䜙 13%䜢༨䜑䜛䛾䛻ᑐ䛧䛶, ᮏ✀䛷䛿, 㣗㐨䛾㛗䛥䛜య㛗䛾

6%䛸య㛗䛻ẚ䛧䛶ⴭ䛧䛟▷䛔. 䛥䜙䛻, D. japonicus 䛷䛿, ༸㯤⭢䛾㛗ᚄ䛜⣙ 100 μm 䛷䛒䜛

䛾䛻ᑐ䛧, ᮏ✀䛾༸㯤⭢䛿⹸య๓᪉䛷ᑠ䛥䛟, ᚋ᪉䛷኱䛝䛔䛸䛔䛖ኚ␗䛜䛒䜚, D. japonicus

䜘䜚䜒኱ᆺ䛾䜒䛾䜢ྵ䜣䛷䛔䛯. 䛣䜜䜙ᙧែ䛾㐪䛔䜢㔜ど䛧䛶, D. 䡆 _{aponicus 䛸䛿ู✀䛸䜏䛺䛧,}

Diprodiscus sp. 䛸䛧䛯.

Family Gorgoderidae㻌 䝂䝹䝂䝕䝷⛉

Gorgodera sp.

ᅗ 7 ほᐹಶయ

1 ಶయ, 2018 ᖺ 7 ᭶ 18 ᪥, ⛅⏣┴⏤฼ᮏⲮᕷ, 䝖䝜䝃䝬䜺䜶䝹 1 ಶయ (㞝 1) , ⭤⬔, ụ

⏣⣫ኈ᥇㞟.

ᐟ୺ཬ䜃ᐤ⏕㒊఩

䝖䝜䝃䝬䜺䜶䝹 1 ಶయ䛾⭤⬔䛛䜙䜏䜙䜜, ᐤ⏕ᩘ䛿 2 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿⦪䛻㛗䛟, ᚋ➃䛿㕌䛟ᑤ䜚, ඛ➃䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 6. 2 mm, ⹸య䛾୰ኸ䛾ᖜ䛿

1. 5 mm 䛷, ᭱኱ᖜ䛿ᮎ➃䛛䜙 3 ศ䛾 1 䛾఩⨨䛷 1. 8 mm 䛻㐩䛩䜛 (ᅗ 7) . ཱྀ྾┙䛿⹸య

䛾ඛ➃䛻఩⨨䛧, ┤ᚄ 0. 6 mm. ⭡྾┙䛿┤ᚄ 0. 75-0. 80 mm 䛾෇ᙧ䛷, ⹸య䛾ඛ➃䛛䜙 3 ศ䛾 1 䛻఩⨨䛩䜛. ๓ဗ㢌䜒➽⫗㉁䛺ဗ㢌䜒䜒䛯䛺䛔. 㣗㐨䛿▷䛟, ⭠䛿⹸య䛾ඛ➃䛛䜙 6 ศ䛾 1 䛾఩⨨䛷ᕥྑ䛻஧ศᒱ䛧, 䛭䜜䛮䜜䛿ᮎ➃㏆䛟䜎䛷ఙ䜃䛶᥋⥆䛫䛪䛻⤊䜟䜛. Ꮚᐑ 䛿⟶≧䛷, ᕥྑ䛻኱䛝䛟ᒅ᭤䛧䛺䛜䜙ᘏ䜃, ⭡྾┙䜘䜚ᚋ᪉䛾య⭍䜢ᗈ䛟༨䜑䜛 (ᅗ 7B, C) .

༸㯤⭢䛿⹸య䛾୰ኸ䛻఩⨨䛧, ᕥྑ䛻 4 䛴䛪䛴䛾ⴥ≧䛻䛺䛳䛶㓄⨨䛩䜛. ༸ᕢ䛿 1 ಶ䛷,

༸ᆺ䜢䛧䛶䛚䜚, ༸㯤⭢䛾ᚋ᪉䛻఩⨨䛩䜛. ༸ᕢ䛾㛗ᚄ䛿 0. 63 mm, ▷ᚄ䛿 0. 38 mm 䛷䛒 䜛. 9 ಶ䛾⢭ᕢ䛜, ⹸య䛾ඛ➃䛛䜙 5 ศ䛾 4 䜋䛹䛾ᚋ᪉䛾, ⭠⟶䛸⭠⟶䛾ෆഃ䛻఩⨨䛩䜛. 䛭 䜜䛮䜜䛾⢭ᕢ䛿㛗ᚄ 0. 28-0. 35 mm ▷ᚄ 0. 15-0. 20 mm 䛾኱䛝䛥䛷, ༸ᕢഃ䛻 5 ಶ, ཯ᑐ

ഃ䛻 4 ಶ䛜⦪䛻 2 ิ䛻୪䜆. ᤼ἥᏍ䛿⹸య䛾ᮎ➃䛻㛤䛟.

≉グ஦㡯

௒ᅇほᐹ䛧䛯ᶆᮏ䛿, ⭠⟶䛜య䛾ᮎ➃㏆䛟䜎䛷ఙ䜃, ୧⏕㢮䜢⤊ᐟ୺䛸䛧䛶䛔䛯䛣䛸䛛䜙,

Bray (2008) 䛛䜙 Gorgoderoidea ୖ⛉䛷䛒䜛䛣䛸䛜䜟䛛䜛. 䜎䛯, ⢭ᕢ䛜⭠䛸⭠䛾ෆഃ䛻఩⨨

䛧, ୧⏕㢮䛾⭤⬔䛻ᐤ⏕䛩䜛Ⅼ䛛䜙, Bray and Blair (2008) 䛻ᚑ䛳䛶 Gorgoderidae 䛸䛧䛯. 䛥 䜙䛻, 9 䛴䛾⢭ᕢ䜢䜒䛴䛸䛔䛖Ⅼ䛛䜙 Campbell (2008) 䛻ᚑ䛳䛶 Gorgodera ᒓ䛸ྠᐃ䛧䛯.

᪥ᮏ䛾୧⏕㢮䛛䜙▱䜙䜜䛶䛔䜛 Gorgodera ᒓ䛷, ✀䜎䛷☜ᐃ䛥䜜䛶䛔䜛䛾䛿 Gorgodera japonica Yamaguti (1936) 䛾䜏䛷䛒䜛 (Uchida and Ooi 2018) . ௒ᅇ䛾ほᐹᶆᮏ䛿, ྾┙䛾

఩⨨, Ꮚᐑ䛾఩⨨, ⢭ᕢ䛾ಶᩘ, ⭠䛾ᙧែ䛿 Gorgodera japonica 䛸▩┪䛧䛺䛔. 䛧䛛䛧, G.

japonica 䛿ཱྀ྾┙䛻ẚ䜉䛶⭡྾┙䛜㢧ⴭ䛻኱䛝䛔䛣䛸䛷▱䜙䜜䛶䛚䜚, 㡑ᅜ⏘䛾 G.

japonica 䛾グ㍕ (Kim et al. 1995) 䛷䛿, ཱྀ྾┙䛸⭡྾┙䛾኱䛝䛥䛾ẚ⋡䛿 1:2 䛸⭡྾┙䛜

኱䛝䛔. 䛭䜜䛻ᑐ䛧䛶, ௒ᅇ䛾ほᐹᶆᮏ䛷䛿, ⭡྾┙䛿ཱྀ྾┙䛾⣙ 1. 2 ಸ䛸, 䜟䛪䛛䛻኱䛝 䛔䛰䛡䛷䛒䜛Ⅼ䛷 G. japonica 䛸䛿䛳䛝䜚䛸␗䛺䛳䛶䛔䜛.௒ᅇ䛾ほᐹᶆᮏ䛿䠈G. japonica 䛸䛿

ู✀䛷᪥ᮏ䛷䛿グ㘓䛾↓䛔✀㢮䛷䛒䜛ྍ⬟ᛶ䛜㧗䛔䛯䜑䠈䛣䛣䛷䛿 Gorgodera sp. 䛸䛧䛯.

Gorgoderina tanigawaensis Uchida et Itagaki, 1974 ᅗ 8

Gorgoderina tanigawaensis Uchida et Itagaki, 1974, p. 66-233, fig. 1-4.

ほᐹಶయ

2 ಶయ, 2017 ᖺ 5 ᭶ 8 ᪥, 㟷᳃┴ᘯ๓ᕷⓒἑ, 䜰䝈䝬䝠䜻䜺䜶䝹 1 ಶయ (㞤 1) , ⭤⬔, ▼ 㒓ᒸ༓ḷ㡢᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䜰䝈䝬䝠䜻䜺䜶䝹 1 ಶయ䛾⭤⬔䛛䜙䜏䜙䜜, ᐤ⏕ᩘ䛿 26 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿⫼⭡䛻ᡥᖹ䛺⣳㗽ᙧ䛷, ๓ᚋ䛷᫂░䛻༊ู䛥䜜䛺䛔 (ᅗ 8) . ᚋ➃䛿㕌䛟ᑤ䜚, ඛ➃ 䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 9. 6 mm, ཱྀ྾┙䛾ᖜ䛿 0. 7 mm, ⹸య୰ኸ䛾ᖜ䛿 2. 1 mm 䛷, ᭱኱

ᖜ䛿⹸య䛾ᮎ➃䛛䜙 3 ศ䛾 1 䛾ᚋ᪉䛷 2. 3 mm 䛻㐩䛩䜛. ཱྀ྾┙䛿⹸య䛾ඛ➃䛻఩⨨䛧,

┤ᚄ 0. 69 mm 䛾෇ᙧ䛷䛒䜛. ⭡྾┙䛿ཱྀ྾┙䜘䜚䜒䜔䜔኱䛝䛟┤ᚄ 0. 82 mm 䛾෇ᙧ䛷, ⹸

య䛾ඛ➃䛛䜙 4 ศ䛾 1 䛾๓᪉㒊䛻఩⨨䛩䜛. ๓ဗ㢌䜒➽⫗㉁䛾ဗ㢌䜒ᣢ䛯䛺䛔. 㣗㐨䛿䛔 䛳䛯䜣‴᭤䛧䛶䛛䜙, ཱྀ྾┙䛾㏆䛟䛷⭠⟶䛸䛺䛳䛶ᕥྑ䛻ศ䛛䜜, ⹸య䛾ᮎ➃㏆䛟䜎䛷ᘏ䜃 䜛䛜┣⟶䛷⤊䜟䜛. Ꮚᐑ䛿⟶≧䛷, ⭡྾┙䛾ᚋ᪉䛷ᕥྑ䛻ᒅ᭤䛧䛶ഃ᪉䛻ఙ䜃, ṇ୰⥺䛾

㏆䛟䛻఩⨨䛩䜛⢭ᕢ䛸䜋䛸䜣䛹㔜䛺䜛䛣䛸䛺䛟యቨ䛻ἢ䛳䛶ᚋ➃䜎䛷ᘏ䜃, ⹸యᚋ➃䛷」ᩘ

䛾䝹䞊䝥䜢ᙧᡂ䛩䜛. ༸㯤⭢䛿⹸య䛾ඛ➃䛛䜙 5 ศ䛾 2 䛾๓᪉㒊䛻఩⨨䛧, 䛭䜜䛮䜜ᴃ෇

ᙧ䛷ᕥྑ䛻 1 䛴䛪䛴㓄⨨䛩䜛. ༸ᕢ䛿 1 ಶ䛷, 㛗ᚄ 0. 62 mm ▷ᚄ 0. 27 mm 䛾ᚋ᪉䛜䜟䛪 䛛䛻䜅䛟䜙䜣䛰༸ᆺ䜢䛧䛶䛚䜚, ༸㯤⭢䛾ᚋ᪉䛷, ⭠⟶䛾ෆഃ䛷ṇ୰⥺䜘䜚䜒ᕥᐤ䜚䛻㓄⨨

䛩䜛. ⢭ᕢ䛿 2 ಶ䛷, 㛗ᚄ 1. 34-1. 75 mm ▷ᚄ 0. 31-0. 34 mm, ⹸య䛾୰ኸ䜘䜚䜔䜔ᚋ᪉䛾,

⭠䛸⭠䛾㛫䛻఩⨨䛩䜛. ⢭ᕢ䛾ᙧ≧䛿⦪䛻㛗䛔୙ᐃᙧ䛷, ⴥ≧䛻䛺䜛䜋䛹῝䛔ษ䜜㎸䜏䜢 5-7 ಶᣢ䛴. ๓᪉㒊䛾⢭ᕢ䛿ṇ୰⥺䜘䜚䜒ᕥᐤ䜚䛻, ᚋ᪉䛾⢭ᕢ䛿ྑᐤ䜚䛻఩⨨䛩䜛.

㑇ఏᏊ᝟ሗ

ᘯ๓ᕷⓒἑ䛾ಶయ䛻䛴䛔䛶, 䝭䝖䝁䞁䝗䝸䜰 DNA28S 䝃䝤䝴䝙䝑䝖䛾ሷᇶ㓄ิ䜢౑䛳䛯⣔⤫

ゎᯒ䛛䜙ᚓ䜙䜜䛯⤖ᯝ䛷䛿, Phyllodistomum kanae 䛚䜘䜃 P. patellare 䛸䛾㏆䛔䜒䛾䛾, 䛭䜜 䜙䛸䛿ู䛺⊂❧䛧䛯఩⨨䛻䛒䜛䛣䛸䛜♧၀䛥䜜䛯 (ᾆ㒊 ᮍⓎ⾲) .

≉グ஦㡯

ほᐹᶆᮏ䛿⢭ᕢ䛜⭠䛸⭠䛾ෆഃ䛻఩⨨䛧, ୧⏕㢮䛾⭤⬔䛻ᐤ⏕䛩䜛Ⅼ䛛䜙, Bray and

Blair (2008) 䛻䛧䛯䛜䛖䛸 Gorgoderidae ⛉䛻ྵ䜎䜜䜛. 䜎䛯, ほᐹᶆᮏ䛿⢭ᕢ䛜 2 䛴䛷⹸య

䛾ᚋ➃䛜⣽㛗䛔䛸䛔䛖ᙧែ䛛䜙 Campbell (2008) 䛻䛧䛯䛜䛳䛶 Gorgoderina ᒓ䛻䛺䜛. ௒ᅇ

ᢅ䛳䛯ᶆᮏ䛿, ᮏ✀䛾య㛗䛜 9. 6 mm 䛸኱䛝䛔Ⅼ䜔, ဗ㢌䜢ᣢ䛯䛺䛔䛣䛸, ⹸య䛜๓ᚋ䛷᫂

░䛻༊ู䛥䜜䛺䛔䛣䛸, ⢭ᕢ䛜 2 ಶ䛷῝䛔ษ䜜㎸䜏䜢䜒䛴䛣䛸, 䛥䜙䛻Ꮚᐑ䛾ᵓ㐀䛾䛔䛪䜜䜒 䛜▩┪䛧䛺䛔䛣䛸䛛䜙 Gorgoderina tanigawaensis 䛸ྠᐃ䛧䛯.

ศᕸ䛸ᐟ୺

ᮏ✀䛿⩌㤿┴䛾㇂ᕝᓅ䛾䜰䝈䝬䝠䜻䜺䜶䝹䛛䜙グ㍕䛥䜜䛯✀㢮䛷䛒䜛 (Uchida et Itagaki

1974) . ௒ᅇ䛾㟷᳃┴ᘯ๓ᕷ䛛䜙䛾Ⓨぢ䛿, ཎグ㍕௨᮶ึ䜑䛶䛾グ㘓䛷䛒䜛.

Family Brachycoeliidae

Mesocoelium brevicaecum Goto et Ozaki, 1929 ᅗ 9

Mesocoelium brevicaecum Goto et Ozaki, 1929a, p. 213-214, fig. 1.

Mesocoelium brevicaecum Goto et Ozaki. ㉺ᬛ, 1930, 388-403, figs. 1-8.

Mesocoelium elongatum Goto et Ozaki, 1929a, p. 214-215, fig. 2.

M. lanceatum Goto et Ozaki, 1929a, p. 216-217, fig. 3.

M. pearsei Goto et Ozaki, 1929b, p. 76-77, fig. 3.

M. japonicum Goto et Ozaki, 1929b, p. 77-78 fig. 4.

M. ovatum Goto et Ozaki, 1929b, p. 78-79 fig. 5

ほᐹಶయ

1 ಶయ, 2017 ᖺ 5 ᭶ 8 ᪥, 㟷᳃┴ᘯ๓ᕷⓒἑ, 䜰䝈䝬䝠䜻䜺䜶䝹 1 ಶయ (㞤 1) , ᑠ⭠, ▼ 㒓ᒸ༓ḷ㡢᥇㞟. 3 ಶయ, 2017 ᖺ 8 ᭶ 6 ᪥, 㟷᳃┴῝ᾆ⏫༑஧†, 䜰䝈䝬䝠䜻䜺䜶䝹 1 ಶయ (㞤 1) , ᑠ⭠, ụ⏣⣫ኈ᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䜰䝈䝬䝠䜻䜺䜶䝹䛾ᑠ⭠䛛䜙䜏䜙䜜䛯.ᐤ⏕ᩘ䛿 1-3 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿⦪䛻㛗䛟, 㛗ᴃ෇ᙧ䛷య⾲䛻䛿ඛ➃䛛䜙 3 ศ䛾 1 䜎䛷᫂░䛺Ჲ䜢ᣢ䛴. Ჲ䛿య䛾୰

ኸ㒊䜎䛷ศᕸ䛩䜛䛜ᚋ➃䛻䛿䛺䛔 (ᅗ 9) . ⹸య䛾ඛ➃䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 2. 1 mm, ⹸ య䛾ᖜ䛿୰ኸ㒊䛷 0. 65 mm 䛷, ඛ➃㒊䛸ᚋ➃㒊䜢㝖䛟䛸ኚ໬䛿ᑠ䛥䛔. ཱྀ྾┙䛿⹸య䛾ඛ

➃㏆䛟䛻఩⨨䛧, ┤ᚄ 0. 27 mm 䛾෇ᙧ䛷䛒䜛. ⭡྾┙䛿ཱྀ྾┙䜘䜚䜒ᑠ䛥䛟┤ᚄ 0. 17 mm 䛾

෇ᙧ䛷, ⹸య䛾ඛ➃䛛䜙 3 ศ䛾 1 䛾๓᪉㒊䛻఩⨨䛩䜛. ๓ဗ㢌䜢䜒䛯䛺䛔. ཱྀ྾┙䛻⥆䛟ဗ 㢌䛿➽⫗㉁䛷, 㛗䛥 80 μm, ᖜ 30 μm. 㣗㐨䛿▷䛟, ⭠䛿ཱྀ྾┙䛛䜙⭡྾┙䛾㊥㞳䛻ᑐ䛧๓ ᪉ 3 ศ䛾 1 䛾఩⨨䛷ᕥྑ䛻஧ศᒱ䛧, 䛭䜜䛮䜜⭡྾┙䜘䜚䜒๓᪉䛷▷䛟⤊䜟䜛. ༸㯤⭢䛿ᴃ

෇ᆺ䛷, ဗ㢌䛾ᚋ᪉䛛䜙⭡྾┙䜘䜚๓᪉㒊䛷, ᕥྑ䛻⣙ 30 ಶ䜋䛹䛜ᐦ㞟䛧䛶㓄⨨䛩䜛. ༸

ᕢ䛿 1 ಶ䛷, ┤ᚄ 0. 17 mm 䛾⌫ᙧ䜢䛧䛶䛚䜚, ⭡྾┙䛾ᕥ┿ᶓ䛻఩⨨䛩䜛. ⢭ᕢ䛿 2 䛴䛒

䜚, ⭡྾┙䛾୰ኸ䜘䜚๓᪉䛻, ᕥഃ䜢๓䛻䛧䛶ᩳ䜑䛻㓄⨨䛩䜛. ⢭ᕢ䛾኱䛝䛥䛿ᕥ䛾༸ᕢഃ

䛜䜔䜔኱䛝䛟㛗ᚄ 0. 18 μm ▷ᚄ 150 μm, 䜒䛖୍᪉䛜㛗ᚄ 0. 15 mm ▷ᚄ 0. 11 mm 䛷, 䛹䛱 䜙䜒༸ᆺ䛷䛒䜛. ᤼ἥᏍ䛜⹸య䛾ᮎ➃䛻 I Ꮠ≧䛻㛤䛟. ⏕ṪᏍ䛾఩⨨䛿୙᫂䛷䛒䜛䠊 㑇ఏᏊ᝟ሗ

ᘯ๓ᕷ䛾䜰䝈䝬䝠䜻䜺䜶䝹䛛䜙᥇㞟䛧䛯 2 ಶయ䛛䜙ᚓࡽࢀࡓ᰾ 28S rDNA ࡢሷᇶ㓄ิࢆ

᪤Ꮡࡢࢹ࣮ࢱ࣮࣋ࢫ࡛ẚ㍑ࡍࡿ࡜, ྎ‴ࡢࢭࢲ࢝࣊ࣅ㢮 Pareas formosensis ࠿ࡽᚓࡽࢀ

ࡓಶయ࡛ Mesocoelium ᒓ࡜ྠᐃࡉࢀࡓ✀㢮ࡸ, ࣮࢜ࢫࢺࣛࣜ࢔ࡢ Mesocoelium sp. ࡜ࡋ

࡚グ㘓ࡉࢀࡓ✀㢮࡟㏆⦕࡛, ᐩᒣࡢ࢖ࣔࣜ࠿ࡽ᥇㞟ࡉࢀࡓ M. elongatum ࡜ྠࡌ࡛࠶ࡗ ࡓ (ᾆ㒊 ᮍⓎ⾲)

≉グ஦㡯

ᮏ✀䛿༸㯤⭢䛜⹸య๓᪉䛻఩⨨䛧, ୧⏕㢮䛾⭠Ⳣ䛻ᐤ⏕䛩䜛Ⅼ䛛䜙, Bray and Blair

(2008) 䛻ᚑ䛳䛶 Mesocoeliidae ⛉䛸䛧䛯. 䜎䛯, ⢭ᕢ䛜䜔䜔ᩳ䜑䛷ᶓ䛻୪䜆䛣䛸䛛䜙,

Pojmanska (2008) 䛻ᚑ䛳䛶 Mesocoelium ᒓ䛸䛧䛯. ᮏ✀䜢, ᕷᕝ (1970) 䛷᪥ᮏ䛛䜙グ㘓䛥 䜜䛶䛔䜛 Mesocoelium ᒓ䛾 4 ✀, M. ovatum, M. japonicum, M. brevicaecum, M. elongatum 䛸 ẚ㍑䛩䜛. ᮏ✀䛿⭡྾┙䛜ඛ➃䛛䜙 3 ศ䛾 1 䛾఩⨨䛻䛒䜛Ⅼ䛷 M. brevicaecum 䛚䜘䜃 M.

elongatum 䛸୍⮴䛧, ⭡྾┙䛜䜋䜌⹸య䛾୰ኸ䛻䛒䜛 M. ovatum 䜔⭡྾┙䛜ඛ➃䛛䜙 4 ศ䛾

1 䛾఩⨨䛻䛒䜛 M. japonicum 䛸༊ู䛷䛝䜛. 䜎䛯, ᮏ✀䛿⢭ᕢ䛜⭡྾┙䛾୰ኸ䜘䜚๓᪉䛻䛒 䜛䛯䜑, ⢭ᕢ䛜⭡྾┙䜘䜚᫂䜙䛛䛻๓ഃ䛻఩⨨䛩䜛 M. ovatum 䛚䜘䜃 M. japonicum 䛸༊ู䛷 䛝䜛. 䛥䜙䛻, ⭠⟶䛾ศᒱ䛜ဗ㢌䛾┤ᚋ䛷, 䛭䜜䛮䜜䛾⭠⟶䛜⭡྾┙䜘䜚ᚋ᪉䜎䛷ఙ䜃䛶䛔

䜛 M. japonicum 䜔, ⭠⟶䛾ศᒱ䛜୧྾┙䛾୰㛫䛷, 䛭䜜䛮䜜䛾⭠Ⳣ䛜⭡྾┙䛾ᚋ⦕䛒䜛䛔

䛿⭡྾┙䜘䜚ᚋ᪉䜎䛷ఙ䜃䛶䛔䜛 M. brevicaecum 䛻ᑐ䛧, ᮏ✀䛿⭠䛜ཱྀ྾┙䛛䜙⭡྾┙䛾

㊥㞳䛻ᑐ䛧๓᪉ 3 ศ䛾 1 䛾఩⨨䛷ᕥྑ䛻஧ศᒱ䛧, 䛭䜜䛮䜜⭡྾┙䛻䛿⮳䜙䛺䛔Ⅼ䛷ᐜ᫆

䛻༊ู䛷䛝, 䛣䜜䜙䛿 M. elongatum 䛾⭠䛾ศᒱ, ⭠䛾ᮎ➃䛾఩⨨䛸▩┪䛧䛺䛔. ௨ୖ䛾ᙧ

ែⓗ≉ᚩ䜔㑇ఏᏊゎᯒ䛾⤖ᯝ䛛䜙, ௒ᅇᚓ䜙䜜䛯ᶆᮏ䜢 M. elongatum 䛸ྠᐃ䛧䛯. ୍᪉, Goldberg and Bursey (2002) 䛿 M. elongatum, M. lanceatum, M. japonicum, M. ovatum, M.

pearsei, M. minutum 䜢䛩䜉䛶 M. brevicaecum 䛾᪂ཧ䝅䝜䝙䝮䛸䜏䛺䛧, Uchida and Ooi

(2018) 䜒䛣䜜䛻ᚑ䛳䛶䛔䜛. 䛣䛣䛷䛿䛣䛾ᢅ䛔䛻‽䛨䛶, ௒ᅇ䛾ほᐹᶆᮏ䜢 M. brevicaecum

䛸䛧䛯.

ศᕸ䛸ᐟ୺

ᮏ✀䛿᭷ᑿ㢮䛸↓ᑿ㢮䛻䜎䛯䛜䜛 20 ✀㢮䛾୧⏕㢮䛛䜙グ㘓䛥䜜䛶䛔䜛 (Uchida and Ooi

2018) . ᮏᕞ䛸஑ᕞ䛷ᗈ䛟▱䜙䜜䛶䛚䜚, 㟷᳃┴䛛䜙䛾グ㘓䛿௒ᅇ䛜ึ䛸䛺䜛.

Mesocoelium sp.

ᅗ 10

ほᐹಶయ

1 ಶయ, 2017 ᖺ 6 ᭶ 6 ᪥, 㟷᳃┴ᘯ๓ᕷᆏඖ, 䝲䝬䜰䜹䜺䜶䝹 1 ಶయ (㞝 1) , ኱⭠, ▼ 㒓ᒸ༓ḷ㡢䞉㛗ᑿ䛿䜛䛛᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䝲䝬䜰䜹䜺䜶䝹䛾ᑠ⭠䛷䜏䜙䜜䛯.ᐤ⏕ᩘ䛿 1 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿๓➃䛜ᗈ䛟ᚋ➃䛜ᑤ䜛㛗༸ᙧ䛷, ඛ➃䛛䜙 3 ศ䛾 1 䜎䛷య⾲䛻᫂░䛺Ჲ䜢䜒䛱, ༙ศ 䜎䛷Ჲ䛜⥆䛔䛶䛔䜛䛜ᚋ➃䛻䛿Ჲ䛿䛺䛔 (ᅗ 10) . ⹸య䛾ඛ➃䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 0. 85 mm, ⹸య䛾ᖜ䛿⹸య䛾ඛ➃䛛䜙 5 ศ䛾 2 䛷᭱኱ 0. 37 mm 䛻㐩䛩䜛. ཱྀ྾┙䛿⹸య䛾ඛ➃ 䛻఩⨨䛧, ┤ᚄ 145 μm 䛾෇ᙧ䛷䛒䜛 (ᅗ 10) . ⭡྾┙䛿ཱྀ྾┙䜘䜚䜒䜔䜔ᑠ䛥䛟┤ᚄ 100 μm 䛾෇ᙧ䛷, ⹸య䛾䜋䜌୰ኸ䛻఩⨨䛩䜛. ๓ဗ㢌䜢䜒䛯䛺䛔. ཱྀ྾┙䛻⥆䛟ဗ㢌䛿➽⫗㉁

䛷, 㛗䛥 35 μm, ᖜ 15 μm. 㣗㐨䛿▷䛟, ⭠䛿⹸య䛾ඛ➃䛛䜙 4 ศ䛾 1 䛾఩⨨䛷ᕥྑ䛻஧ศ

ᒱ䛧, ⹸య䛾༙ศ䜎䛷ᘏ䜃, ᥋⥆䛫䛪䛻⤊䜟䜛. ༸㯤⭢䛿⌫ᙧ䜎䛯䛿ᴃ෇ᙧ䛷, ཱྀ྾┙䛾 ᚋ᪉䛛䜙⹸య䛾୰ኸ௜㏆䜎䛷 100 ಶ⛬ᗘ䛜ᕥྑ䛻ᐦ㞟䛧䛶ศᕸ䛩䜛. ༸ᕢ䛿 1 ಶ䛷, 㛗ᚄ

120 μm, ▷ᚄ 85 μm 䛷, ⭡྾┙䜘䜚䜔䜔ᚋ᪉䛷⭡྾┙䛾ᕥᩳ䜑䛻఩⨨䛩䜛. ⢭ᕢ䛿 2 䛴䛒

䜚, ⭡྾┙䛾ഃ᪉䛻, ⭡྾┙䛻㔜䛺䜛ᩳ䜑๓᪉䛷, ᶓ୪䜃䛻㓄⨨䛩䜛. ᮏᶆᮏ䛷䛿ᕥഃ䛾

⢭ᕢ䛜ྑഃ䜘䜚䜒䜟䛪䛛䛻๓᪉䛻఩⨨䛧䛶䛔䛯. ᮏᶆᮏ䛷䛿, ༸䛿䜏䜙䜜䛺䛛䛳䛯. ⢭ᕢ䛿

༸ᕢഃ䛻఩⨨䛩䜛ᕥഃ䛜㛗ᚄ 105 μm ▷ᚄ 85 μm, 䜒䛖୍᪉䛾ྑഃ䛜㛗ᚄ 115 μm ▷ᚄ 80 μm 䛷, ༸ᆺ䛒䜛䛔䛿ᴃ෇ᙧ䛷䛒䜛. ᤼ἥᏍ䛜⹸య䛾ᮎ➃䛻 V Ꮠ≧䛻㛤䛟. ⏕ṪᏍ䛾఩⨨䛿

୙᫂䛷䛒䜛䠊

≉グ஦㡯

ᮏ✀䛿༸㯤⭢䛜⹸య๓᪉䛻఩⨨䛧, ୧⏕㢮䛾⭠Ⳣ䛻ᐤ⏕䛩䜛Ⅼ䛛䜙, Bray and Blair

(2008) 䛻ᚑ䛳䛶 Mesocoeliidae ⛉䛸䛧䛯. 䜎䛯, ⢭ᕢ䛜䜔䜔ᩳ䜑䛷ᶓ䛻୪䜆䛣䛸䛛䜙,

Pojmanska (2008) 䛻ᚑ䛳䛶 Mesocoelium ᒓ䛸䛧䛯. ᮏ✀䛿๓㏙䛾 Mesocoelium elongatum 䛸, య㛗, ⭡྾┙䛾఩⨨, ⭠⟶䛾㛗䛥, ༸ᕢ䛾఩⨨, ᤼ἥᏍ䛾ᙧ䛾㐪䛔䛷༊ู䛷䛝䜛䛯䜑, ✀ ᑠྡ୙ヲ䛾ู✀䛸䛧䛯. ௒ᅇほᐹ䛧䛯 1 ಶయ䛿༑ศ䛻ᡂ⇍䛧䛶䛔䛺䛔ಶయ䛰䛳䛯. ௒ᚋ, ⏕ Ṫჾᐁ䛾ヲ⣽䜢ᢕᥱ䛩䜛䛯䜑䛻, ᡂ⇍ಶయ䜢ほᐹ䛩䜛ᚲせ䛜䛒䜛.

Brachycoeliidae sp.

ᅗ 11 ほᐹಶయ

1 ಶయ, 2016 ᖺ 10 ᭶ 2 ᪥, 㟷᳃┴እ䞄὾⏫, 䝲䝬䜰䜹䜺䜶䝹 1 ಶయ (ND1) , ᑠ⭠, ୹⩚

ዋኴ᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䝲䝬䜰䜹䜺䜶䝹䛾ᑠ⭠䛷䜏䜙䜜䛯.ᐤ⏕ᩘ䛿, 2 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿ඛ➃䛸ᮎ➃䛜㕌䛟ᑤ䜛ⴥ≧䛷, ඛ➃䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 0. 48 mm, ⹸య䛾ᖜ䛿⹸య 䛾ᮎ➃䛛䜙 3 ศ䛾 1 䛷᭱኱ 172 μm 䛻㐩䛩䜛 (ᅗ 11) . ⹸య⾲㠃䛻Ჲ䛿䛺䛔. ཱྀ྾┙䛿⹸

య䛾ඛ➃䛻఩⨨䛧, ┤ᚄ 62. 7 μm 䛾෇ᙧ䛷䛒䜛. ⭡྾┙䛿ཱྀ྾┙䜘䜚䜒ᑠ䛥䛟㛗ᚄ 40. 3 μm

▷ᚄ 31. 9 μm 䛾ᴃ෇ᙧ䛷, ⹸య䛾ඛ➃䛛䜙 5 ศ䛾 2 䛾๓᪉䛻఩⨨䛩䜛. ཱྀ྾┙䛻⥆䛔䛶㛗

䛥 7. 38 μm, ᖜ 12. 3 μm 䛾๓ဗ㢌䜢䜒䛱, 䛣䜜䛻㛗䛥 13. 5 μm, ᖜ 12. 3 μm 䛾➽⫗㉁䛺ဗ㢌 䛜⥆䛟. 㣗㐨䛿⹸య䛾ඛ➃䛛䜙 4 ศ䛾 1 䛾఩⨨䛷ᕥྑ䛻஧ศᒱ䛧, ⭠䛻䛺䛳䛶ఙ䜃䜛䛜䛹䛣 䜎䛷⥆䛔䛶䛔䜛䛛ุู䛷䛝䛺䛛䛳䛯. ༸㯤⭢䛿୙ᐃᙧ䛷, ဗ㢌䛾ᚋ᪉䛛䜙⭡྾┙䜎䛷, ᕥ

ྑ䛻ᐦ㞟䛧䛶㓄⨨䛩䜛. ༸ᕢ䛸⢭ᕢ䛿ุู䛷䛝䛺䛛䛳䛯. Ꮚᐑ䛿⹸య䛾⭡྾┙䜘䜚ᚋ᪉䛾 య⭍䜢ᒅ᭤䛧䛺䛜䜙ᐦ䛻༨䜑, 䛣䜜䛻ຍ䛘䛶, ⹸య䛾ྑഃ䛷䛿⭡྾┙䛾ഃ᪉䛛䜙䜔䜔๓᪉ 㒊䛻䜎䛷ఙ䜃䛶䛔䜛. ༸䛿ᴃ෇య䛷㛗ᚄ 11 μm, ▷ᚄ 4. 9 μm. ᤼ἥᏍ䛿ุู䛷䛝䛺䛛䛳䛯.

≉グ஦㡯

ᮏ✀䛿༸㯤⭢䛜⹸య๓᪉䛻఩⨨䛧, ୧⏕㢮䛾⭠Ⳣ䛻ᐤ⏕䛩䜛Ⅼ䛛䜙, Bray and Blair (2008) 䛻ᚑ䛳䛶 Brachycoeliidae (ཎᩥ䛷䛿 Mesocoeliidae ⛉) 䛸䛧䛯. ᮏ✀䛿๓㏙䛾 Mesocoelium brevicaecum 䛚䜘䜃 Mesocoelium sp. 䛸, ⹸య䛾ᙧ≧䛸๓ဗ㢌䛜䛒䜛䛣䛸䛷༊ู

䛷䛝䜛䛯䜑, ู✀ Brachycoeliidae sp. 䛸䛧䛯.

Family Haematoloechidae

Haematoloechus japonicus (Yamaguti, 1936) ᅗ 12

Pneumonoeces sibiricus japonicus Yamaguti, 1936, p. 567-569, fig. 21.

Haematoloechus sibiricus japonicus (Yamaguti, 1936)

Haematoloechus japonicus (Yamaguti, 1936) . Goldberg SR & Bursey CR (2002)

ほᐹಶయ

3 ಶయ, 2018 ᖺ 7 ᭶ 15 ᪥, 㟷᳃┴ᘯ๓ᕷᆏඖ, 䝒䝏䜺䜶䝹 1 ಶయ (㞝 1) , ⫵, ▼㒓ᒸ

༓ḷ㡢, ῝⃝ᖾᜨ᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䝒䝏䜺䜶䝹䛾⫵䛷䜏䜙䜜䛯.ᐤ⏕ᩘ䛿 5 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿⫼⭡䛻ᡥᖹ䛺⣳㗽ᙧ䛷⹸య䛻Ჲ䜢䜒䛯䛪, ඛ➃䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 7. 9 mm, ๓

➃䛾ཱྀ྾┙䛾୰ኸ䛷䛾ᖜ䛿 0. 65 mm, ⹸య䛾୰ኸ䛷᭱኱ᖜ䛸䛺䜚 2. 2 mm 䛻㐩䛩䜛 (ᅗ

12) . ཱྀ྾┙䛿⹸య䛾ඛ➃䛻఩⨨䛧, ┤ᚄ 0. 4 mm 䛾෇ᙧ䛷䛒䜛. ⭡྾┙䛿ཱྀ྾┙䜘䜚䜒䜔

䜔኱䛝䛟┤ᚄ 0. 53 mm 䛾෇ᙧ䛷, ⹸య䛾ඛ➃䛛䜙 5 ศ䛾 2 䛾๓᪉㒊䛻఩⨨䛩䜛. ๓ဗ㢌䛿 ᣢ䛯䛺䛔. ྾┙䛻⥆䛟ဗ㢌䛿➽⫗㉁䛷, 㛗䛥 0. 15 mm, ᖜ 0. 06 mm. 㣗㐨䛿▷䛟, 䛔䛳䛯䜣

ྑഃ䛻‴᭤䛧䛯ᚋ, ⹸య䛾 8 ศ䛾 1 䛷ᕥྑ䛻ศ䛛䜜䛶⭠⟶䛸䛺䜚, ⹸య䛾ᮎ➃㏆䛟䜎䛷ᘏ䜃 䛶┣⟶䛷⤊䜟䜛. Ꮚᐑ䛿⟶≧䛷, ⭡྾┙䛾๓᪉䛷䛿ᕥྑ䛾⭠Ⳣ䛾ෆഃ䜢ᕥྑ䛻ᩘᅇ‴᭤

䛧, ⭡྾┙䜘䜚ᚋ᪉䛷䛿⭠⟶䛾ෆഃ䛸ᕥྑ䜢ᒅ᭤䛧䛺䛜䜙๓ᚋ䛻㉮䜚, ᚋ➃䛸⭡྾┙䛾ᕥྑ

䛷⎔≧䛻䛺䜛. ༸ᕢ䛿 1 ಶ䛷, 㛗ᚄ 0. 68 mm ▷ᚄ 0. 46 mm 䛷, ༸ᆺ䜢䛧䛶䛚䜚, ⭡྾┙䛾 ᚋ᪉ᕥഃ䛻఩⨨䛩䜛. ⢭ᕢ䛿༸ᕢ䜘䜚ᚋ᪉䛻䠎䛴䛒䜚, ศᒱ䛧䛯⭠⟶䛾ෆഃ䛻఩⨨䛩䜛䛜, ᶆᮏ䛷䛿ᙧ䛜୙᫂░䛷䛒䛳䛯. ༸㯤⭢䛿༸ᆺ䛷, 6-20 ಶ䛜䜎䛸䜎䛳䛶ⰼ≧䛾ᵓ㐀䛸䛺䜛. 䛣 䛾ⰼ≧䛾ᵓ㐀䛿 16-20 ಶ䛒䜚, ⭠䛾ศᒱⅬ䜘䜚䜔䜔ᚋ᪉䛛䜙, ⹸య䛾ᚋ➃㏆䛟䜎䛷୙つ๎䛺 㛫㝸䛷୪䜣䛷䛔䜛.

≉グ஦㡯

ᮏ✀䛿, ⏕ṪᏍ䛜⹸య䛾๓ഃ䛻䛒䜚, ⹸య䛻Ჲ䜢䜒䛯䛪, ୧⏕㢮䜢ᐟ୺䛸䛩䜛䛣䛸䛛䜙,

Bray (2008) 䛻ᚑ䛳䛶, Plagiorchioidea ୖ⛉䛸䛧䛯. 䜎䛯, ⹸య䛾ᚋ᪉䛻ẚ䜉䛶๓᪉䛜▷䛟,

」ᩘ䛾༸㯤⭢䛜ᐦ╔䛧䛶䛚䜚, ୧⏕㢮䛾⫵䛻ᐤ⏕䛧䛶䛔䛯䛣䛸䛛䜙, Bray (2008) 䛻ᚑ䛳䛶

Haematoloechidae ⛉䛻ྵ䜎䜜䜛. 䛭䛧䛶༸㯤⭢䛜⢭ᕢ䛾ᚋ᪉ᇦ䜎䛷ᗈ䛜䛳䛶䛚䜚, ⢭ᕢ䛜

⭠䛸⭠䛾ෆഃ䛻఩⨨䛧, ᡂ⇍ಶయ䛷⭡྾┙䜢᭷䛩䜛Ⅼ䛛䜙, Tkach (2008) 䛻ᚑ䛳䛶 Haematoloechus ᒓ䛻ྵ䜎䜜䜛. ᮏ✀䜢, Yamaguti (1936) 䛾 Haematoloechus japonicus 䛸 ẚ㍑䛩䜛䛸, ཱྀ྾┙䛜⭡྾┙䛻ẚ䜉䛶ᑠ䛥䛔Ⅼ, ༸ᕢ䛿༸ᆺ䛷䛒䜛Ⅼ, ⹸య䛻Ჲ䜢䜒䛯䛺 䛔Ⅼ, Ꮚᐑ䛿⭡྾┙䛾๓ഃ䛷䜒ᒅ᭤䛧䛶䛔䜛Ⅼ䛾䛔䛪䜜䛾グ㍕䜒▩┪䛧䛺䛔. 䜘䛳䛶, ᮏ

✀䜢 Haematoloechus japonicus 䛸ྠᐃ䛧䛯.

ศᕸ䛸ᐟ୺

ᮏ✀䛿䛣䜜䜎䛷໭ᾏ㐨䛛䜙୰㒊ᆅ᪉䜎䛷䛻ศᕸ䛩䜛 3 ✀䛾䜹䜶䝹㢮 (䜴䝅䜺䜶䝹, 䝖䜴䜻䝵 䜴䝎䝹䝬䜺䜶䝹, 䝖䝜䝃䝬䜺䜶䝹) 䛛䜙グ㘓䛥䜜䛶䛔䜛 (Uchida and Ooi 2018) . ௒ᅇグ㘓䛥 䜜䛯䝒䝏䜺䜶䝹䛿᪂ᐟ୺䛸䛺䜛.

Family Omphalometridae

Opisthioglyphe japonicus Uchida, Itagaki et Inoue, 1980 ᅗ 13

Opisthioglyphe japonicus Uchida, Itagaki et Inoue, 1980, p. 110-113, figs. 3-5.

ほᐹಶయ

1 ಶయ, 2017 ᖺ 8 ᭶ 6 ᪥, 㟷᳃┴῝ᾆ⏫, 䜰䝈䝬䝠䜻䜺䜶䝹 1 ಶయ (㞤 1) , ᑠ⭠, ụ⏣

⣫ኈ᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䜰䝈䝬䝠䜻䜺䜶䝹䛾ᑠ⭠䛷䜏䜙䜜䛯.ᐤ⏕ᩘ䛿 1 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

య䛿⫼⭡䛻ᡥᖹ䛷ඛ➃䛜㕌䛟ᑤ䜛䜋䛛ᖜ䛾ኚ໬䛜ᑡ䛺䛔ᖏ≧䛷䛒䜛 (ᅗ 13) . ⹸య䛾ඛ

➃䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 2. 58 mm, ⹸య䛾୰ኸ䛾ᖜ䛿 0. 89 mm. ⹸య䛿ඛ➃䛛䜙 3 ศ䛾 1 䜎䛷᫂░䛺Ჲ䜢䜒䛱, ୰ኸ௜㏆䜎䛷Ჲ䛜⥆䛔䛶䛔䜛䛜, ᚋ➃䛻䛿Ჲ䛿䛺䛔. ཱྀ྾┙䛿⹸య 䛾ඛ➃䛻఩⨨䛧, ┤ᚄ 185 μm 䛾෇ᙧ䛷䛒䜛. ⭡྾┙䛿ཱྀ྾┙䜘䜚䜒䜔䜔኱䛝䛟┤ᚄ 195 μm 䛷, ⹸య䛾ඛ➃䛛䜙 3 ศ䛾 1 䛾๓᪉㒊䛻఩⨨䛩䜛. ๓ဗ㢌䜢䜒䛯䛺䛔. ཱྀ྾┙䛻⥆䛟ဗ㢌䛿

➽⫗㉁䛷, 䛭䜜䛮䜜㛗䛥 68 μm, ᖜ 29 μm 䛾ᴃ෇ᙧ䛷䛒䜛. 㣗㐨䛿▷䛟, ဗ㢌䜘䜚䜔䜔ᚋ᪉ 䛷ᕥྑ䛻ศ䛛䜜䛶⭠⟶䛸䛺䜚, ⹸య䛾ᚋ➃䛛䜙 3 ศ䛾 1 䜋䛹䜎䛷ఙ䜃䛶䛔䜛䛜, ᮎ➃䛾఩⨨

䛿ุู䛷䛝䛺䛛䛳䛯. ⏕ṪᏍ䛿⹸య䛾ᕥഃ䛷, ཱྀ྾┙䛾ഃ᪉䛷እ⦕㏆䛟䛻㛤䛟. ༸㯤⭢䛿

༸ᙧ䛷, ဗ㢌ᚋ㒊䛾఩⨨䛛䜙⹸య䛾እ⦕䛻ἢ䛖䜘䛖䛻ከᩘ䛜ᐦ䛻ศᕸ䛧, ⢭ᕢ䛾ᚋ᪉䜢㉺

䛘䛶⹸య䛾ᮎ➃䛻㐩䛩䜛. ༸ᕢ䛿 1 ಶ䛷, ┤ᚄ 273 μm 䛾⌫ᙧ䛷, ⭡྾┙䛾ྑᐤ䜚䛾ᩳ䜑ᚋ ᪉䛻㓄⨨䛩䜛. ⢭ᕢ䛿 2 ಶ䛷⌫ᙧ䛛༸ᆺ䛷, ᕥഃ䛷䛿㛗ᚄ 253 μm, ▷ᚄ 243 μm, ྑഃ䛷

䛿㛗ᚄ 321 μm ▷ᚄ 243 μm. ⢭ᕢ䛿⹸య䛾ᚋ➃䛛䜙 3 ศ䛾 1 䛾఩⨨䛻䛒䜚, ᕥഃ䛜ྑഃ䜘

䜚䜒ᩳ䜑๓᪉䛻఩⨨䛩䜛. ᤼ἥᄞ䛿 Y Ꮠᆺ䛷, ⢭ᕢ䜘䜚๓᪉䛷஧ศᒱ䛩䜛.

≉グ஦㡯

㻌 ᮏ✀䛿, ⢭ᕢ䛜 2 䛴䛒䜚, ⏕ṪᏍ䛜ṇ୰⥺䜘䜚ᕥഃ䛻㛤䛝, ୧⏕㢮䛜ᐟ୺䛷䛒䜛Ⅼ䛛䜙

Bray (2008) 䛻ᚑ䛳䛶 Pleurogenidae ⛉䛸䛧䛯. ᮏ✀䛿䛥䜙䛻, ྾┙䛾఩⨨䜔༸㯤⭢䛜⹸య

䛾඲య䛻ศᕸ䛩䜛Ⅼ, ⢭ᕢ䛜ᮎ➃㏆䛟䛻䛒䜛Ⅼ䛺䛹䛾≉ᚩ䛛䜙 Opisthioglyphe japonicus 䛸

ྠᐃ䛧䛯. 䛯䛰䛧, ணഛⓗ䛺㑇ఏᏊゎᯒ䛷䛿ศ㢮Ꮫⓗ఩⨨䛻㛵䛩䜛ၥ㢟䛜ᣦ᦬䛥䜜䛶䛔䜛 (୰ᑿ ᮍⓎ⾲) .

ศᕸ䛸ᐟ୺

ᮏ✀䛿䛣䜜䜎䛷 Uchida and Ooi (2018) 䛻䜘䛳䛶㟷᳃┴䛸᪂₲┴, ⚟ᓥ┴䛛䜙グ㘓䛥䜜䛶 䛔䜛 (Uchida and Ooi 2018) . ᐟ୺䛿䝒䝏䜺䜶䝹, 䝖䝜䝃䝬䜺䜶䝹, 䝙䝩䞁䜰䜺䜺䜶䝹, 䝍䝂 䜺䜶䝹䛾 4 ✀䛷䛒䛳䛯. ௒ᅇ䛾㟷᳃┴῝ᾆ⏫䛛䜙䛾グ㘓䛿, ཎグ㍕௨᮶ึ䜑䛶䛷, ᐟ୺䛾 䜰䝈䝬䝠䜻䜺䜶䝹䜒᪂ᐟ୺䛷䛒䜛.

Family Pleurogenidae

Pleurolobatus lobatus (Ozaki, 1926) ᅗ 14

Pleurogenes lobatus Ozaki, 1926, p. 40-43, figs. 7-8.

Pleurogenes lobatus Ozaki. ᕷᕝ, 1970, p. 183-184, ᅗ 101.

Pleurolobatus lobatus (Ozaki, 1926) , Kaw, 1943, p. 106-107; Lotz and Font, 2008, p.

572, fig. 58. 12

ほᐹಶయ

1 ಶయ, 2019 ᖺ 4 ᭶ 13 ᪥, 㟷᳃┴ᘯ๓ᕷᗙ㢌▼, 䝲䝬䜰䜹䜺䜶䝹 1 ಶయ (㞝 1) , య⭍, ▼ 㒓ᒸ༓ḷ㡢᥇㞟

ᐟ୺䛸ᐤ⏕㒊఩

㻌 䝲䝬䜰䜹䜺䜶䝹 1 ಶయ䛾య⭍䛛䜙䜏䜙䜜, ᐤ⏕ᩘ䛿 4 䛷䛒䛳䛯. ᐤ⏕㒊఩䛻䛴䛔䛶䛿, య

⭍ෆ䛾⫶䛻௜╔䛩䜛≧ែ䛷, ⷧ䛔⭷䛷ᅖ䜎䜜䛯⌫య䛜䛒䛳䛯. 䛭䛾⭷䜢◚䜛䛸 4 ಶయ䛜ワ䜎 䛳䛶䛔䛯. 䛣䛾⌫య䛿ᐟ୺䛾ෆ⮚䛸⧅䛜䛳䛶䛚䜙䛪, ⊂❧䛧䛶䛔䛯䛣䛸䛛䜙, 䛣䛣䛷䛿ᐤ⏕㒊

఩䜢య⭍䛸䛧䛯.

ᙧែ䛾グ㍕

య䛿⫼⭡䛻ཌ䜏䛾䛒䜛ᗈᴃ෇ᙧ䛷, ඛ➃䛛䜙ᮎ➃䜎䛷䛾㛗䛥䛿 7. 6 mm, ⹸య䛾୰ኸ䛾

ᖜ䛿 5. 0 mm, ⹸య඲య䛜 6-10 μm 䛾Ჲ䛷そ䜟䜜䛶䛔䜛 (ᅗ 14) . ཱྀ྾┙䛿⹸య䛾ඛ➃䛻

఩⨨䛩䜛䛜, ᶆᮏస〇䛾㝿䛻ᅽᖹ䛾䛧䛩䛞䛷ṍ䜣䛷䛧䜎䛔, 䛭䛾ᙧ䜔኱䛝䛥䛿㆑ู䛷䛝䛺䛛 䛳䛯. ๓ဗ㢌䛾᭷↓䜒, ྠᵝ䛻ᶆᮏస〇䛜ୖᡭ䛟䛔䛛䛪, ㆑ู䛷䛝䛺䛛䛳䛯. ⭡྾┙䛿⦪

᪉ྥ䛻䜟䛪䛛䛻㛗䛔෇ᙧ䛷㛗ᚄ 0. 44 mm ▷ᚄ 0. 30 mm 䛷, ⹸య䛾ඛ➃䛛䜙 3 ศ䛾 1 䛾๓ ᪉䛻఩⨨䛩䜛. ဗ㢌䛿➽⫗㉁䛷, 㛗䛥 0. 2 mm, ᖜ 0. 12 mm. ဗ㢌䛻⥆䛟㣗㐨䛿, 㛗䛥 0. 6

mm, ᖜ 0. 1 mm. ⭠⟶䛿⹸య䛾ඛ➃䛛䜙 9 ศ䛾 1 䛾఩⨨䛷ᕥྑ䛻஧ศᒱ䛧, ⢭ᕢ䜢㉺䛘䛶

⹸య䛾ᚋ➃䛛䜙 3 ศ䛾 1 䜎䛷ఙ䜃, ஫䛔䛻᥋⥆䛫䛪䛻┣≧䛻䛺䛳䛶⤊䜟䜛. 㝜ⱼᄞ䛿⊃༸

ᙧ䛷ඛ➃䛜᭤䛜䜛䛜, ᶆᮏ䛾≧ែ䛜ᝏ䛟㛗䛥䛿䜘䛟䜟䛛䜙䛺䛛䛳䛯. ⏕ṪᏍ䛿ဗ㢌䛸⭡྾┙

䛾㊥㞳䛾୰㛫䜋䛹䛾఩⨨䛷, ⹸య䛾ᕥഃ⦕䜘䜚䛻㛤䛟. ⢭ᕢ䛿 2 ಶ䛷䛹䛱䜙䜒῝䛔ษ䜜㎸䜏 䜢」ᩘ䜒䛴. ⢭ᕢ䛿, ⹸య䛾୰ኸ㒊䛷, ⭡྾┙䛾ᚋ᪉䛻఩⨨䛧, ⭠⟶䛾ᕥྑ䛻䜋䜌ᑐ⛠䛻

఩⨨䛩䜛. ༸ᕢ䛿 1 ಶ䛷, ⭡྾┙䛾┤๓䛻䛒䜚, ⢭ᕢ䛸ྠ䛨䛟῝䛔ษ䜜㎸䜏䛜䛒䜛䛜, ⢭ᕢ 䜘䜚ษ䜜㎸䜏䛾ⴥ䛜ᑡ䛺䛔. Ꮚᐑ䛿⟶≧䛷, ⭡྾┙䛾ᚋ᪉䛛䜙ᒅ᭤䛧䛺䛜䜙ఙ䜃, ⢭ᕢ䜘䜚 ᚋ᪉䛾య⭍䜢Ṥ䛹඲䛶‶䛯䛧䛶䛔䜛. ༸㯤⭢䛿㣗㐨䛾ᚋ᪉⭡ഃ䛻఩⨨䛧, 8 䛛䜙 23 ಶ䛾ᑠ

⬊䛛䜙䛺䜛ᡣ䛜⭠⟶䛻ἢ䛳䛶㓄⨨䛩䜛. ᤼ἥᄞ䛿⹸య䛾ᚋ᪉䛻㛤䛟䛜, ᙧ䛿㆑ู䛷䛝䛺䛛䛳 䛯.

≉グ஦㡯

ᮏ✀䛿, ⏕Ṫ⭢䛻῝䛔ษ䜜㎸䜏䜢䜒䛱, ⢭ᕢ䛜ᒅ᭤䛧䛯Ꮚᐑ䜘䜚๓᪉䛻䛒䜛䛣䛸䛛䜙, Lotz and Font (2008) 䛻ᚑ䛳䛶 Pleurolobatus ᒓ䛸䛧䛯. Pleurolobatus ᒓ䛿 Pleurolobatus lobatus1

✀䛾䜏䛛䜙ᵓᡂ䛥䜜䛶䛔䜛 (Kaw, 1943) . Pleurogenes lobatus 䛿, Ozaki (1926) 䛜ᅄᅜ䛾䜹 䝆䜹䜺䜶䝹䛾⫹⟶ (Bile duct) 䛛䜙ᚓ䜙䜜䛯ಶయ䛷᪂✀グ㍕䛧䛯✀㢮䛷䛒䜛. Ozaki (1926) 䛿, Pleurogenes lobatus 䛜༸ᕢ䛸⢭ᕢ䛻῝䛔ษ䜜㎸䜏䜢᭷䛩䜛䛾䛻ᑐ䛧, ྠᒓ䛾௚䛾✀䛷䛿

༸ᕢ䜔⢭ᕢ䛻ษ䜜㎸䜏䛜↓䛔䛣䛸䜢༊ูⅬ䛸䛧䛶䛔䜛.

䛭䛾ᚋ, Kaw (1943) 䛿, ⏕ṪᏍ䛜⭠䛾ศᒱⅬ䛾ᚋ᪉䛻㛤䛟Ⅼ䜔༸ᕢ䛸⢭ᕢ䛻ⴭ䛧䛔ษ䜜

㎸䜏䛜䛒䜛Ⅼ, ༸㯤⭢䛜┣≧䛷, ᑠ䛥䛺ᑠ⬊䛛䜙Ⅽ䜛ᡣ≧䛷䛒䜛Ⅼ䛛䜙 lobatus 䜢䝍䜲䝥✀䛸 䛧䛶 Pleurolobatus ᒓ䜢๰タ䛧䛯. 䛣䛾ᢅ䛔䛿, Lotz and Font (2008) 䜒ㄆ䜑䛶䛔䜛. ⌧ᅾ䜎䛷, Pleurolobatus ᒓ䛿, Pleurogenes lobatus ௨እ䛾✀䛿ྵ䜎䜜䛶䛚䜙䛪, 䜎䛯, 㑇ఏᏊゎᯒ䜒䜎 䛰⾜䜟䜜䛶䛔䛺䛔 (Choudhary et al. 2015) . ᕷᕝ (1970) 䜔 Uchida (1975) 䛿, ᮏ✀䛾Ꮫ

ྡ䜢 Pleurogenes lobatus 䛾䜎䜎䛻䛧䛶䛔䜛䛜, ᒓ䛾ጇᙜᛶ䛻䛴䛔䛶≉䛻㆟ㄽ䛿⾜䛳䛶䛔䛺䛔

䛯䜑, ༢䛻ศ㢮Ꮫⓗኚ᭦䜢▱䜙䛺䛛䛳䛯䛯䜑䛸ᛮ䜟䜜䜛.

௒ᅇほᐹ䛧䛯ᶆᮏ䛿⢭ᕢ䛸༸ᕢ䛻ⴭ䛧䛔ษ䜜㎸䜏䛜䛒䜛Ⅼ, ⹸య඲య䛻Ჲ䜢䜒䛴Ⅼ, ⏕Ṫ Ꮝ䛜ဗ㢌䛸⭡྾┙䛾㊥㞳䛾୰㛫䜋䛹䛾఩⨨䛷, ⹸య䛾ᕥഃ⦕䜘䜚䛻㛤䛟Ⅼ, ༸㯤⭢䛜㣗㐨 䛾ᚋ᪉䛛䛴⭠⟶⭡ഃ䛻఩⨨䛧, ᑠ⬊䛛䜙䛺䜛ᡣ䛜䜎䛸䜎䛳䛶㓄⨨䛩䜛Ⅼ, Ꮚᐑ䛜ᒅ᭤䛧䛺䛜 䜙ᚋ㉮䛧, ⢭ᕢ䜘䜚ᚋ᪉䛾యෆ䛾Ṥ䛹඲䛶䜢‶䛯䛧䛶䛔䜛Ⅼ䛷 Ozaki (1926) 䛾 Pleurogenes.

lobatus 䛾グ㍕䛸▩┪䛧䛺䛔. ≉䛻, Ozaki (1926) 䛜✀䛾㚷ูⅬ䛸䛧䛶ᣲ䛢䛶䛔䜛⢭ᕢ䛸༸ᕢ 䛻ⴭ䛧䛔ษ䜜㎸䜏䛜䛒䜛Ⅼ䛿ᮏᶆᮏ䛷䜒☜ㄆ䛥䜜䛯.

Ozaki (1926) 䛾 Pleurogenes lobatus 䛿䜹䝆䜹䜺䜶䝹䛾⫹⟶䛛䜙Ⓨぢ䛥䜜䛶䛔䜛䛾䛻ᑐ䛧,

ᮏᶆᮏ䛿䝲䝬䜰䜹䜺䜶䝹䛾య⭍䛛䜙䜏䛴䛛䛳䛯. 䛥䜙䛻, Ozaki (1926) 䛻䜘䜛䛸, Pleurogenes

lobatus 䛾⭠䛿㛗䛟⢭ᕢ䜢㉺䛘䛶⹸య䛾ᚋ➃䛛䜙 4 ศ䛾 1 䛾఩⨨䛻㐩䛧䛶䛔䜛䛾䛻ᑐ䛧, ᮏ

ᶆᮏ䛷䛿⭠䛿⢭ᕢ䜢㉺䛘䛶䛔䜛䜒䛾䛾, ⹸య䛾ᚋ➃䛛䜙 3 ศ䛾 1 䜋䛹䛾఩⨨䛷⤊䜟䛳䛶䛔䜛.

䜎䛯, Ozaki (1926) 䛾 P. lobatus 䛾య㛗䛿᭱኱䛷䜒 5. 1 mm 䛷䛒䜛䛜, ᮏᶆᮏ䛿 7. 6 mm 䛸

኱䛝䛛䛳䛯. 䛣䛾䜘䛖䛺⭠䛾㛗䛥䛾㐪䛔䜔య㛗䛾㐪䛔䛜ಶయኚ␗䛺䛾䛛䠈ᅽᖹ䛾⛬ᗘ䛺䛹䛾

ேⅭⓗせᅉ䛻䜘䜛䛾䛛䠈䛒䜛䛔䛿䠈⣔⤫䛜␗䛺䜛䛯䜑䛺䛾䛛䛿䠈ᶆᮏᩘ䛜ᑡ䛺䛔䛯䜑䛻ุู

䛷䛝䛺䛛䛳䛯.

ศᕸ䛸ᐟ୺

ᮏ✀䛿䛣䜜䜎䛷, ᐑᓮ┴䛸㤶ᕝ┴䛷グ㘓䛥䜜, ᐟ୺䛿䜹䝆䜹䜺䜶䝹䛸䝒䝏䜺䜶䝹䛾 2 ✀䛷 䛒䛳䛯 (Uchida and Ooi 2018) . ௒ᅇ䛾㟷᳃┴ᘯ๓ᕷ䛷䛾䝲䝬䜰䜹䜺䜶䝹䛛䜙䛾グ㘓䛿, ศ ᕸ໭㝈䛾᭦᪂䛸᪂ᐟ୺䛾グ㘓䛸䛺䜛.

Cestoda㻌 ᮲⹸⥘

Cestoda㻌㻌 sp.

ᅗ 15 ほᐹಶయ

1 ಶయ, 2017 ᖺ 6 ᭶ 6 ᪥, 㟷᳃┴ᘯ๓ᕷᆏඖ, 䝙䝩䞁䜰䝬䜺䜶䝹 1 ಶయ (㞝 1) , ᑠ⭠, ▼ 㒓ᒸ༓ḷ㡢᥇㞟. 1 ಶయ, 2017 ᖺ 6 ᭶ 18 ᪥, 㟷᳃┴㫏䞄ἑ⏫, 䝲䝬䜰䜹䜺䜶䝹 1 ಶయ (㞤

1) , ᑠ⭠, ụ⏣⣫ኈ᥇㞟. 1 ಶయ, 2016 ᖺ 9 ᭶ 24 ᪥, ᒾᡭ┴す࿴㈡⏫, 䝰䝸䜰䜸䜺䜶䝹 1

ಶయ (ND1) , ᑠ⭠, ୹⩚ዋኴ᥇㞟. 1 ಶయ, 2018 ᖺ 7 ᭶ 22 ᪥, 㟷᳃┴༑࿴⏣ᕷ, 䝍䝂䜺䜶 䝹 (㞤 1) , ᑠ⭠, ▼㒓ᒸ༓ḷ㡢᥇㞟.

ᐟ୺䛸ᐤ⏕㒊఩

䝙䝩䞁䜰䝬䜺䜶䝹, 䝍䝂䜺䜶䝹, 䝲䝬䜰䜹䜺䜶䝹, 䝰䝸䜰䜸䜺䜶䝹䛾ᑠ⭠䛷䜏䜙䜜䛯.ᐤ⏕

ᩘ䛿 1-7 ಶయ䛷䛒䛳䛯.

ᙧែ䛾グ㍕

㢌㒊䛻䠐䛴䛾྾┙䜢ᣢ䛴 (ᅗ 15) . ほᐹ䛧䛯ᶆᮏ䛿඲䛶ᮍᡂ⇍ಶయ䛷, ⏕Ṫჾᐁ䜢䛿䛨 䜑䛸䛩䜛ෆ㒊ᵓ㐀䛿䜏䜙䜜䛺䛛䛳䛯.

≉グ஦㡯

௒ᅇ䛾ᶆᮏ䛷䛿, ᡂ⇍ಶయ䛜ྵ䜎䜜䛶䛔䛺䛛䛳䛯䛯䜑, ௒ᚋ䛾ㄪᰝ䛷ᡂ⇍ಶయ䜢᥇㞟䛧 䛶ᨵ䜑䛶ほᐹ䜢⾜䛖ᚲせ䛜䛒䜛.

Phylum Acanthocephala㻌 㖋㢌ື≀㛛 Class Palaeacanthocephala㻌 ྂ㖋㢌⹸⥘

Order Echinorhynchida㻌 䝁䜴䝖䜴䝏䝳䜴┠

Family Echinorhynchidae㻌 䝖䝀䝝䝅䝁䜴䝖䜴䝏䝳䜴⛉

Pseudoacanthocephalus toshimai Nakao, 2016㻌 䜶䝌䜹䜶䝹㖋㢌⹸

ᅗ 16-18

Pseudoacanthocephalus toshimai Nakao, 2016, p. 326-329, figs. 2-4.

ほᐹᶆᮏ

㞝 1 ಶయ, 2015 ᖺ 6 ᭶ 5 ᪥, ᖹᕝᕷ, 䝍䝂䜺䜶䝹 1 ಶయ (ND1) , ᑠ⭠, ୹⩚ዋኴ᥇㞟.

㞝 1 ಶయ, 2016 ᖺ 10 ᭶ 17 ᪥, す┠ᒇᮧ, 䝍䝂䜺䜶䝹 1 ಶయ (ND1) , ᑠ⭠, ୹⩚ዋኴ᥇㞟.

㞝 1 ಶయ, 2017 ᖺ 4 ᭶ 14 ᪥, ᘯ๓ᕷᆏඖ䛰䜣䜆䜚ụ, 䝲䝬䜰䜹䜺䜶䝹 1 ಶయ㻌 (㞤 1) , ᑠ

⭠, ▼㒓ᒸ༓ḷ㡢᥇㞟. 㞝 2 ಶయ, 2017 ᖺ 4 ᭶ 16 ᪥, ᘯ๓ᕷᆏඖ䛰䜣䜆䜚ụ, 䝲䝬䜰䜹䜺 䜶䝹 2 ಶయ (㞝 1 㞤 1) , ᑠ⭠, ▼㒓ᒸ༓ḷ㡢᥇㞟. 㞤 1 ಶయ, 2017 ᖺ 5 ᭶ 8 ᪥, ᘯ๓ᕷ

ⓒἑ Ỉ䛯䜑ụ, 䜰䝈䝬䝠䜻䜺䜶䝹 1 ಶయ (㞝 1) , ᑠ⭠, ▼㒓ᒸ༓ḷ㡢᥇㞟. 㞝 1 ಶయ,