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Accepted by J. Klimaszewski: 18 Jul. 2011; published: 28 Oct. 2011

ZOOTAXA

ISSN 1175-5326 (print edition)

ISSN1175-5334(online edition) Copyright © 2011 · Magnolia Press

Zootaxa 3075: 1–151 (2011)

www.mapress.com/zootaxa/

Monograph

ZOOTAXA

World catalogue of the species of the tribe Lomechusini

(Staphylinidae: Aleocharinae)

PETER HLAVÁČ

1

, ALFRED F. NEWTON

2

& MUNETOSHI MARUYAMA

3

1Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamýcká 1176, CZ-165 21 Prague 6, Czech Republic. E-mail: phlavac@stonline.sk

2Zoology Department/Insect Division, Field Museum of Natural History, 1400 South Lake Shore Drive, Chicago, IL 60605 USA. E-mail: anewton@fieldmuseum.org

3The Kyushu University Museum, Fukuoka, 812-8581 Japan. E-mail: dorylus@hotmail.com

Magnolia Press

Auckland, New Zealand

3075

(2)

HLAVÁČ ET AL.

2

· Zootaxa 3075 © 2011 Magnolia Press

Peter Hlaváč, Alfred F. Newton & Munetoshi Maruyama

World catalogue of the species of the tribe Lomechusini (Staphylinidae: Aleocharinae)

(Zootaxa 3075)

151 pp.; 30 cm.

28 Oct. 2011

ISBN 978-1-86977-811-8 (paperback)

ISBN 978-1-86977-812-5 (Online edition)

F

IRST

P

UBLISHED IN 2011 BY

Magnolia Press

P.O. Box 41-383

Auckland 1346

New Zealand

e-mail: zootaxa@mapress.com

http://www.mapress.com/zootaxa/

© 2011 Magnolia Press

All rights reserved.

No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any

means, without prior written permission from the publisher, to whom all requests to reproduce copyright

material should be directed in writing.

This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose

other than private research use.

ISSN 1175-5326 (Print edition)

ISSN 1175-5334 (Online edition)

(3)

Zootaxa 3075 © 2011 Magnolia Press ·

3

WORLD CATALOGUE OF LOMECHUSINI

Table of contents

Abstract . . . 3

Introduction . . . 4

Methods . . . 4

Nomenclatural and taxonomic corrections and changes . . . 6

1. Replacement names . . . 6

2. Reinstated junior secondary homonyms . . . 7

3. Genera moved or excluded from Lomechusini . . . 7

4. New synonymies and assignments . . . 8

5. Other taxonomic problems . . . 10

Natural history . . . 11

Catalogue of Species . . . 12

Subtribe Lomechusina. . . 12

Subtribe Myrmedoniina. . . 16

Subtribe Termitozyrina . . . 120

Acknowledgments . . . 122

References . . . 122

Appendix 1 . . . 144

Appendix 2 . . . 148

Appendix 3 . . . 153

Abstract

All names of species and higher taxa proposed in the tribe Lomechusini (Staphylinidae: Aleocharinae) up to the end of the year 2010 are cataloged, with a full bibliography of original descriptions. The original combination of the described spe- cies, original citation of their type localities and their world distributions are given. The following replacement names are proposed for newly discovered primary and secondary junior homonyms: Brachysipalia elgon nom. nov. (= B. elgonensis Pace), Drusilla sculpticollides nom. nov. (=Drusilla sculpticollis Pace), Macrogerodonia bolivianides nom. nov. (=M. bo- liviana Pace), Orphnebius curticornides nom. nov. (= O. curticornis Pace 2008), Pachorhopala ituri nom. nov. (= P. ge- rardi Bernhauer 1932) , Zyras (Camonia) aethiopicides nom. nov. (=Zyras (Camonia) aethiopicus Pace) and Zyras (Zyras) neoparageminus nom. nov. (= Zyras (Zyras) parageminus Pace). The following secondary homonyms are rein- stated as valid: Drusilla opacicollis Cameron (= D. vedda Pace), and Zyras (Grammodonia) erraticus Last (= Z. (G.) ad- jectus Last). The following new synonymies are proposed (junior synonym first): Aenictobiina Kistner (= Aenictoteratini Kistner), Athexeniina Pace (= Myrmedoniina Thomson), Thoracophagus Kistner (= Drusilla Leach), Urodonia Silvestri (= Pedinopleurus Cameron), Lomechusoides hosodai K. Sawada (= L. amurensis (Bernhauer)), Diplopleurus notabilis Pace (= Trachydonia incredibilis Bernhauer), and Acanthoglossa badia Motschulsky (= Zyras (Glossacantha) affinis Kraatz) (the last-mentioned being a reversal of precedence). The following new combinations are established: Aenictocleptis antennarius (Pace) ex Zyras (Glosacantha); Drusilla congoensis (Kistner) ex Thoracophagus; Pedinopleurus notabilis (Silvestri) ex Urodonia; Pedinopleurus reptans (Pace) and Pedinopleurus tumorfrontalis (Pace ) ex Zyras (Diaulaconia); Pseudastilbus fortispina (Pace) ex Zyras (Glosacantha) and Deroleptus draco (Assing) ex Orphnebius. A lectotype is designated for Lomechusoides ganglbaueri (Bernhauer) to fix this name as a synonym of L. amurensis (Wasmann). The following genera are newly included into Lomechusini, subtribe Myrmedoniina: Aenictophila Seevers, Athexenia Pace, Cephaplakoxena Pace, Malaybergius Kistner, Mimaenictus Kistner & Jacobson, Procantonnetia Kistner & Jacobson, Steysborgia Kistner & Jacobson and Weissflogia Kistner. The genus Myrmigaster Sharp is removed from Lomechusini to Oxypodini. Nomenclatural problems with the names Lomechusa Gravenhorst, Lomechusoides Tot- tenham and Atemeles Dillwyn, and recent taxonomic changes to the genus Orphnebius Motschulsky, are also discussed. Lomechusini as currently constituted includes 207 genera and 2205 species or subspecies placed in the subtribes Lom- echusina (3 genera, 40 species or subspecies), Termitozyrina (11 genera, 16 species) and Myrmedoniina (193 genera and 2149 species or subspecies). Their regional distribution patterns and extensive associations with ants and termites are briefly summarized.

Key words: Coleoptera, Staphylinidae, Aleocharinae, Lomechusini, catalogue, world, myrmecophile, termitophile

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