J.Fac Educ Tottori Univ. (Nat Sci),41(1992)19-35
Pteropods from the Miocene Fugatti Mudstone Member of Wtti,
Tottori PreFecture9 Japan
Saburo AKAGI*1,I‐ Iiroshi SHIBATA*2 and Nobuhiroヽ
luRATA*3
(R￠c￠ル￠'4P万′2θ,7992)
Abstract
A pteropod fauna consisting of ninc species is reportcd and dcscribed frona the upper part
ofthe Fuganji Mudstonc Member of Waji,Totto PreFccture lt dosely resemblcs thosc of
thc carly middle Miocene Takakura Formation of the Katsuta Croup,Okayama Prefecture and Upper Shalc Mcmbcr ofthe Bihoku Croup,Hiroshima Prefccture
Thc occurrence of the ptcropods indicatcs inno、 vs of a 、varm occanic 、vatcr into the depositional sitc during deposition of the upper part Of the Fuganji Mudstonc Member
lntroduction
One of thc authors,Murata discovered ptcropodous moIIuscs flom the Mioccnc Fuganji Mudstone Mcmbcr of the Tottott Group at thc bank of thcヽVaji Rivcr about l km cast of
ヽ
Vaji, Totto Prcfccturc in the autumun of 1987, Subscqucnt conccting in thc Fuganti Mudstone of the Waji area by thc authors, espccially Murata has rcvcalcd threc additional locahties where ptcropods occur, and has yieldcd largc pteropod collcctions from thc four iocahtics, All pteropod spcciinens are moulds and casts. Ninc pteropodous species arc representcd in thc conectiOns. In addition onc heteropodous specics、vas found at one of thesc locahtics, Thc greater part of these pclagic molluscs,ho、 vever,rcmain unnamcd oMFing tO thcill statc of prcscrvation of thcir specirncns This papcr dcscribcs pclagic moHuscs frona thcsc
locahtics, and mcntions the agc and sedilncntary environmcnt of thc pteropod‐ bcaring strata
Thcre is no published study on the pteropod fauna of thc Tottori(3roup.
All spcciFnCnS treated in this paper exccpt those inustrated and mcasurcd are deposited in Faculty of Education,Tottori H」 niversity Of thc inustratcd and mcasurcd spcchncns one of
thc illustratcd spccirnens of C,:ο G,r,ヵ,どα is in the Tottori Prefcctural Muscum, and the
rcmaining spcci:mens arc depositcd in Conege of Gcneral Education,Nagoya Univcrsity.
ヽVe thanks MI, とζiyoharu HOshilni of the Tottori Prcfcctural Muscum who ano、 vcd us to
cxa■line ptcropod spccilnens in thc Tottori Prefcctural Muscunl collcction,and hclpcd usin the
ncld investigation,
*l Facu■y of Education,Totto University 率2 conegc of ceneral Education,Nagoya Univcrsity +3 Kokufu― cho,Iwanli―gun,Tottori Prcfecture
AKACと,S,SHIBATA,H AND MURATA,N
Stratigraphy
Thc carly to middle Mioccnc Tottori Croup(Murayama and othcrs,1963)ね cxtCns cly
cxposcd in thc castern part of Tottori Prcfecturc, Many 、vorkers including Nishiwaki and
lmamura(1956),Uemura and others(1979),MatSumOto(1986,1991)and thC TOtto
Grccn
Tuff Rescarch Group(1989)propOSCd stratigraphic classincatiOns of thc group morc or lcss different from onc another So far as the stratigraphy of this arca is conccrncd, ho、 vever,thcrc is no rcmarkablc diffcrcncc among prcvious studics, This report has foHo、 ved thc classincatiOn and nomenclaturc of Uemura and others who stratigraphically mapped thc ヽVakasa district in dctail.
Thc Fuganti Mudstonc Mcmber consists chieRy of mudstoncs.Itis morc than 400 m in total thickncss.Thc lower part of the FuBanii Mudstonc consists of laminated mudstoncs and siltstoncs,and land plant rcmains arc plcntifRll in thcm at Fuganji,Okamasu and somc othcr places,bcing mostly rcprcsented byんrttdθ?′ο
''7miOCι η
'c,, S,′
テχ sp, CοttρサοrT''/T,フ陶,71々
',
デ,=肋 冷∫sp.and L′ ?ガカ 胞うι′″ 脇 οd,服 (HOjO,1973),but nO fOS載にof ma ne organttms havc been found in thcm. Fronl thc lithofacies and the fossil contcnt they seeHi to bc lacustrinc
sedilnents.
Thc lnaiddlc part of the Fuganii Mudstonc Mcmber consists of interbcddcd mudstoncs and
siltstoncs、vith intcrcalations of acidic tuff, Marine nshcs such as S,′ ′,7T?Jr'and οs/7Eι′,S OCCur
abundantly in its lo、 vcr part at Miyanoshita,Mitani and Shichiyama in addition to thc plants mcntioncd abovc A Japancsc hving rcprcsentativc of Odttι r"∫, 0, '9猾t℃x inhabits a cold 、vatcr
Thc upper part of thc mcmber is composed of massivc black mudstoncs intercalatcd with
acidic tuffs and turbidites, Marine mo■ uscs are fatrly conlm6n in its lo、 vcr half,but no fossils havc been found in its uppcr half.
L6cations of iocalitics from 鞘′hich pteropods 、vcrc obtaincd arc sho、
vn in Fig l The
stratigraphic rclation of these iocahtics is deter■ nablc on thc basis of markcr tuffs Thcy
Senzanguchi
一一一︻一
一
Pteropods from he wi∝cnC Fuganii MidStonc しegend BlaCI Shttle FuffJCeoiS Shale AcidiC‐ t'ff Tuffuceous Sand FosSil loCality PterOpods
Fig.2. StlatigFaphic scctio■s at the ptcropod
10calhics in Wait.
stratigraphitaly are、vithin a 13:n thick soquence of rnudstoncs and turs in thc!ppcr part of the Fuganii Mudstonc(Fig.21.ThiS Sequence hes ib6ut 50 m aboVo tulo top Ofthe ulldcllying CO'810meratcs exposcd in the acor Of the WaJl R cr betwccn Kallliaraftlne and tte Waif Bridge.
Age
A Checklist of ptOropods iom Wajttis shown in Tablc生 . Ofthe nine speoes ypg腕9r,sp.is the most abundant spedies.
Seven pteFOpOdOus specics reported by Shibata a■
d othors(1989)と
oⅢthC Takakura
Formation of tho Katsut―
a Croup,0(ay4血
a PArefeCturo and SiX species repoied,y them fl・ o節日 Ｉ ■ Ｅ Ｅ Ｅ ′
AKACI,S,SHIBATA,H AND MURATA,N
Table l Chcckhst of hcteropodous and ptcrOpodous molluscs from thc Fuganii MudstOne Mcmber of Waji
R:1-20,F:21-50,Ci 51-100,AI Inore than 100 spccilnens
Species Localities 1 2 3 4 C,r,4α河
,sp R
Lどれ,cJη,sp R C
エンガOυ7iF sp R
?予石,=BrTι′肋 すIPttdd,Daudin R R F
I Sp. A R R A
Cιtο c,rilt,ど,Audenino R R ?C ,λrマヵηοゼChCCChia―RispOh R R R R
C・ テ翅を,″,'(Shibata) A R R R
C,フたriヵ,sp R R R R
C,フοJrヵ力 う汚′ん,筋(Kittl) R R F
thc Upper Shalc Mcmbcr of thc Bihoku Group, Ihroshiina Prpefecturc arc included in thc ptcropod fauna,and 1/pgJη ιtt sp.is thc donlinant spccics in thc both formations. This closc faunal rcsemblance suggests that pteropod― bcaring strata of thc thrcc areas are synchronous. The ptcropod―bcaring strata in the Takakura Formation and the Uppcr Shale Mcmbcr occur in the uppcr part ofthcス
甲ρ肪じ
οりれ
,sc″α
ri9-t/bを?r力 ,Cr,∫∫
たο
∫
簿″
zonulc of Tai(1959)(Shibata and othcrs, 1989),the upper boundary of which agrees with thc boundary bctwccn thc planktonic foraIIlinifcral zoncs N9 and N10 of BIow(1969)(Tai,1985).The pterOpod― bcattng strata of this arca thus sccna to be assigncd to thc N9 zone in age. A silnlar pteropod fauna characterizcd by thc prcdon nance of yagテη♂′′α sp. occurs in thc Konoura Shale Mcmbcr of Uchiura Croup,Fukui Prcfccturc(Shibata and Ujihara)1990)。 Nakagawa and othcrs(1985) suggested thc lo、vcr part of thc N9 zone for thc Konoura Shale. It sccms to support thc aforcmentioncd agc asttgnmcnt for thc pteropod― bca ng strata of WatiTai(1959)repOrtcd thc samc bcnthic foraninifcral zonule as the pteropod‐ bca ng strata in thc Takakura Formation and thc Uppcr Shale Membcr ttom the Fuganii Mudstonc of Waji As hc gavc no locanty data for his sampling sitcs,their stratigraphic relation、 vith our pteropod
localitics is unkno、v■.
Palecenvironment
Caッοと,れ,t7うな
,励
occurs in thc carly to n ddlc Mioccnc Mizuna■li Group of(3ifu Prcfcc‐ture in association with such tropical,shallow sca,benthic molluscs as,形
′
ι
う筋
rip sp,C力θ
,cο一
/Jレ∫
Sp.and No',″
rθう胞 ο∫α〃,乃οどれdね (ItOigawa and others, 1981). It Suggests that thcpteropodous spccics is an inhabitant of a、 varm、 vatcr,and the other ptcropodous species which occur associatcd with itin thc FuBanil Mudstone Mcmbcr seem to be also warm watcr spccies
Table 2 sho、vs a chccklist of benthic monuscs collectcd frona the pteropod locahtics. DoHli… nant species by locahty are as folowsi locality l,と ,陶9psね sp.andと,れ,テレ′,sp localitics 2 and
3;PFOp♂α “
囲 ∫′捌町 ヵ
`,η
力ηdね and D♂′9cr9p?c″乃Pιじた力,翻′and iocality 4;と,脇?〃力,c,肋 sp.
Ptcropods frOm the Miocene Fuganji MudstOne
Tablc 2 Checklist of bcnthic moluscs from the pteropod iocalties in
Wati Species Locahtics 1 2 3 4 /1c力,rしv′οた,ヵ,=,ど
(Yokoyama) * * *
と,μ￠′rv/P,c,榜sp * *
ス0カsp *
Ⅲr,〃αヵsp. キ Y9′′ザrl sp * Pο′r肋々′J,sp *
L,脇 `"υ ∫ねsp
キ*
キ Pr?P￠α//B,dd'レ胞 立″たユ″,どKanehara 半P,曳 ″gα/P/J￠力dね
Omoh and lnoue(MS) *
半*
P sp
キ D￠ルo9χcttИ P♂cた力αれ,(Gabb) * *
半*
Lけれ,ど,う2 sp
キ*
と,,4ο れ,,c,万″々￠,r,脇(COnrad)
キ*
Lθ,たοttοど 'a cf ηJppο々,c′d(OgaSawara) *
Pppカ テ,sp *
TFt′,cJ,sp.
ホ D￠ηれ2万,p sp
十 几ダd″θ刀筋力,//1 cfノοたοノ '//B所(MaHyama) * * *
P力οs sp *
associated bcnthic mo‖uscs suggcst batyal depths as thc dcpth of dcposition for thc pteropod― bearing strata On thc ba s of thc assodatcd bcnthic molluscan fauna Matsumoto(1986) suggestcd that thc iniddlc to uppcr part of the FuganjI Mudstone Mcmbcr was dcpositcd in a
cold、vatcr. Thc occurrcnce of ptcropods frolla it,ho、 vevcr,indicates that the surfacc laycr of thc sea of the dcpositional site was subjcct to a warm watcr during deposition of thc uppcr part
of the member,though thc bottom laycr of the sca of the assumcd depths、vas naturany nllcd
、vith a cold、 vatcr.
Systematic descriptiOns Phylum Mo■usca Subphylum Conchifcra Class Gastropoda Subclass Prosobranchia Order Mcsogastropoda Supcrfalllily Atlantacca Family Carinariidac Ccnus C,rJれ,r力
Lamarck,1801
C,′′ηαr,,sp. (Pl.1,Figs.1-2)24
Fateriali Two impcrfect spccilnens.
Descriptioni Shc■
smal, hoodhkc,
、vith a carina; apcx coiled back, situatcd a httlc postcrior of thc posterior cnd of aperturc of the shcn standing on thc aperturc in sidc vic、vGrcatest hcight of the shcn shghtly Smaler than the lcngth of thc aperturc, Carina 、ve‖
dcvcloped, about 1/4 as broad as the lcngth of the apcrturc near thc apcrturc EFnbryOnic shcll consisting of about thrcc whoris, sman, helicinoid; thc rcmaining part of thc shc■ transvcrscly phcated, phcations narrower than their intcrspaccs, slightly curvcd do、 vn、vard,
continuing along thc carina turning upward, about 18 prilnary phcations some of Ⅵ′hich arc
bipartitc and a fc、v sccOndary ones onc an adult spccirnen.
Mcasuremcnt(in mm):
(Figured spccimcn) ( ″ ) Grcatcst height 8,0 3.0 Length of aperturc85
40
Comparison: This spccies rcscmbles C.P,r?サ ο
'Maycr from thc Langhian of ltaly in outlinc,
but thc shcn is lcss dcpresscd and transversc phcae arc lcss cro、 vdcd C. n, sp Stancu from
thc uppcr Bandcnian of Romania sccms to bc allicd to this spccics,but thc specilncns ngurcd arc too poorly prescrved for detailcd comparison, C sp. froEl thiS arca differs from thc Rcccnt C′αれ,′θた
'(PCrOn and Lesucur)in thC highcr shcll and the highcr ca
na, Occurrencc: Locanty 4. と,′η,c'rt,sp Shibata,Taguchi Subclass Opisthobranchia Ordcr Thccosomata Subordcr Euthccosomata Fa■lily Lirnacinidac
Genusと
'脇 ,c'々,Bosc,1817
と,脇αOれ粉sp. (Pl l,Figs.3,4) and Ujihara,1989,p.34,pl.4,Ilgs.1-3.Materiali 53 axiany cOmpresscd inner moulds.
Description: SheH vcry small, largest specirncn being 2 2 1nnl in 、vidth, sinistral Whoris
aboutfour and a half,rather rapidly enlarging. Spirc seems to projcct frona the outcr Ⅵ′horl in
side vie、v,for■ling a lo、v cone. Suturc dceply imprcssed. Surfacc smooth Umbilicus dcep
and broad.
Pteropods from thc Mioccnc Fuganji Mudstone
Width
2 2 (Figurcd spccimcn) ( ク )Comparison: Froni the sillailarity of thc sizc, thc modc of cnlargemcnt of、 vhoris and thc shapc of umbilicus this small Lirnacina seems to be conspccinc、vithと,″η,c,々,sp describcd by
Shibata and othcrs(1989)と om thC middlc Mioccnc Takakura Formation ofthc Katsuta Croup, Okayama Prcfccturc and thc Uppcr Shalc Mcmbcr of the Bihoku Croup,卜 nroshilna Prefcc‐
turc L sp.rcscmblcs L
ソガソαttη,(Rcuss)仕
om the Miocene of Poland in apical vicw,butthc umbiLcus is broadcr Availablc spccilncns arc insufnciently prcscrvcd for spccinc dctcr_
■lination
Occurrence: Locantics 3 and 4
FamJy Cavohn dac
Subfa■lily Chonac Ganus rFy,わεノ′たFol,1875
rF/α
′
。
9ノね
sp.(PlattC l,Fig.5)
Matcrial T、 vo transvcrscly compressed spccl=ncns
Dcscriptioni Shcn sman,conical,annulatcd;apical angle about 30° Latcral pronlcs faintly
convcx Annulations inc, a httle narrower than thcir interspaces, bcco■ ling mOrc closcly
spaccd to、 vard thc apcxゥ about 40 in total,numbcring in 13 in 2 mm length ofthc n ddlc part of the shcll.Apcx sccms to bc bulbous.Length 5 6 mm,diamcter 2.4mm(ngurcd spccimcn)
Remarks: Thh spccics closcly rcsembles the Rcccnt, sr′ 'α
r,(Rang)and打
力,テ″ヵ∫なCollins froln thc Mioccnc of Haiti)but annulations arc much incr and grcatcr in numbcr 声
sp.somcwhatc rcscmblcs P泡
″りα′οり ′ね ,刀打,′,筋 (TatC)仕om the Eoccnc of Australia insurfacc sculpturc, but thc shcn is smaler and faintly convex conical in outhnc, and thc apcx appcars to bc bulbous This spccics is probably unnamcd spccics,but availablc speciincns arc not good cnough for typc spccilncns,
Occurrencc: Locahty 4
Gcnus yag,刀ι′′α Daudin, 1800
?798'/1θJJp′宅pr3s∫
,Daudin,1800
(Pl.1,Fig.6) げα =,Ю θ/Jp ′▼pres∫ ' Daudin, 1800, p.145, pl ll, ng.1; Bastcrotラ 1825, p.19, pl.4, ng,16;AKAC●I,S,SHIBATA,H AND MURATA,N
Bellardi, 1873,p 64;Tibcri, 1880,p 37;I(itd, 1886,p.57,pl.2,ngs.17-22,Benoist, 1889, p.28,pl.2,flg.4a―c;Audcnino,1897,p.108,Sacco, 1904ラ p.15,pl,4,AB,10,Bcnini, 1905, p.40,Chccchia―Rispoli,1921,p.15,ng.6;Zilch(in Wenz),1959,p.51i ng.170;Sirna,1968, p.424,ng,12,Ctyroky,Papp and Stciningcr,1968ラ p.132,pl.4,ngs.1,2,Shibata,1970,pl
4,Ilgs,13a,b;Robba,1971,p.89,pl.4,ngs.6,7;Robba,1972,p.502,pl.59,ag.12,pl.60,
ng.8:Shibata(in ltOigawa,Shibata and Nishimoto),1974,p.191,pl.60,ngs 9,lo,Shibata) 1977, pl.13, ngs 3-6; Shibata, 1980, pl.3, ngs.13-17; Itoiga、 va, Shibata, Nishirnoto and Okumura, 1981,p1 47,ngs.6,7;Itoigawa,Shibata,Nishilnoto and Okumura, 1982,p.295; Ujihara and Shibata,1982;pl.6,ng.21,Spano,1983,p.257,pl,16,ng 9,p.17,ngs,1_7,pl 18,ngs,1_5; Shibata, 1983,p 77, pl.2, Ilgs,7,8;Jansscn, 1984a,p.75,pl.3,ng。 11,pl.4, ngs,9_13;Japanesc, 1984b,ngs lo_13,Jansscn, 1986,ng 5;Shibata and Ujthara, 1989, p 25,pl.1,ags l_3;Jansscn,1989,p.51,pl.7,ngs 8-12,p1 8,ags l_8;Shibata,Taguchi and
Ujihara,1989,p.35,p14,ngs.4,5,
Matcrial: 28 transvcrsely comprcsscd specilnens.
Descriptioni Shcl smaH,bottlc‐ shaped Greatest transversc diamctcr situatcd at about thc
n ddle of the shcl,from this point the shel rapidly tapcring postcriorly. Latcral keels distinct in the posterior part of the shcll, Post― apertural constriction of thc shcⅡ faint. Apcrtural rims convex.Apcx sharp.Apical angle about 40°
.Lcngth 6.Ommぅ width 2.4mm(agurcd
specimcn).Rcmarks: All availablc specilnens arc poorly Daudin in gcneral outlinc and size, though thcy spccilncns are questionably assigned toレ1 '9ρ/θdd,.
Occurrcnce: Localties l,3 and 4.
prcscrvcd. They resemble y. ダ9prθsS'
appcar to bc some、 vhat sicndcrcr. Thc
予 `α
gカプカ sp
(Pl.1,Figs 7-10)
ypg,れ
θ
JJp sp.Shibata,Taguchi and Ujihara,1989,p.36,pl.4,ngs,9_12.Matc al: More than 1700 spcdmens were coHcctcd from Waji An spcdmcns are trans‐ vcrscly aattcncd
Dcscriptioni Shel largc, slcndcr, tapering convcxly to、 vard a pointed apcx, with a 、vcl dcnncd kecl on cach side of thc shcl Both dorsal and vcntral pronles gcntly convex,grcatcst
dorso‐vcntral diameter bcing situatcd at about the■liddlc of thc shcl lcngth, Post― apcrtural constriction of the shell shght. Dorsal and vcntral apcrtural rilns convex Ernbryonic shel bulgcd,constricted antcriorly;tip pointed. Extcrior smooth Apical angle about 20° .
Ptcropods ttom the Miocenc Fuganji MudstOnc
Mcasurcment(in mm):
(Figurcd spccimen) ( ク ) Lcngth 14,3 11,4 11.3145
140
13.9113
Width
3,7 3.537
35
3.834
Remarks: This species closcly resemblcs y,α ,∫rr,IIc,Kittl from the Mioccne of Austria,but
is iarger and slcnderer lt also rescmblcs T/. α,cた肋η′,c,Clarkc fronl thc Mioccnc of New Zcaland,but its apcrture is wider and sccms to be clongated clliptical
y,sp is thc most common ptcropodous spccics in thc Waji arca.It occurs widcly in thc
early H ddlc Mioccnc of thc Chugoku and Sanin rcgions including this area, Numcrous
specilnens arc availablc frona thcsc rcgions,but thcy al arc casts and are strongly comprcsscd transvcrsely Bcttcr material must bc availablc in ordcr to makc dcflnitc spccinc identinca‐
tion.
Shibata and others(1989)suggCStCd that this spccies scrvcs as a good indicator of thc N9 zone` Occurrencc: Localities l-4. Gcnus Crrο Linnacus,1767 crio εα′,/t,ど
,Audenino, 1897
(Pl l,Figs.11,12) (Vテο crrrttαすρ Audcnino,1897,p,102,p1 5,ag,3,1)icci,1961,P 41,pl 15,ng.5,pl 16,ng,8; Robba, 1971, p 81,pl.2, ag 17,pl.3, ngs.1_3;Stancu, 1974, p.186, p1 3, ng.lo; Robba and Spano, 1978, p.772, p1 80, ag,4; Shibata and Ujihara, 1989, p 29, pl.3, ngs 3, 4; Shibata,Taguchi andとテ
jtharaぅ p.37,p1 4,ngs 15,16.Bttrrr乃 ガ,脇 (Frrvう ♂′′,′,/PT)cα′Jれ ,ど坊r71(Audenino),bCllini,p.39,ags 4(),41
Matcrial: T、 vo impcrfcct specilncns coⅡ ectcd by us frOm iocahty 4 and onc outer lnould of
the dorsal side in the Tottori Prcfcctural Museunl conection frOm locality 3.
Descriptioni Shell small,rhombicゥ ncarly as、vidc as iong. Latcral inargins shghtly divcrg‐
ing,prolongcd anteriorly,for■ling a short lateral spine. Dorsal sidc marked、 vith iongitudinal
ribs and transverse folds,Iongitudinat ribs ive,a broad,pronlinent rib in thc mcdian part and
t、vo narro、v, lo、v ribs on cach sidc of thc median rib, inncr iateral oncs running close by thc
mcdian onc,convcrging postcriorly,transvcrse folds curvcd, their convcx sidc looking toward
apcrturc, narro、ver than thcir intcrspaccsi ventral side with a pronlincnt, bipartitc, median
transvcrse folds in aぢ ttcned areas bct、 vccn thc mcdian iongitudinal rib and thc lateral lnargins; in addition faint iongitudinal striations arc visible on thc median part of eithcr side Apcrturc 、vidc;apcrtural rilns convcx. EInbryonic shcl bulbous Apical angle about 45° .
Measurcment(in mm):
(Figured spccimcn)( ク )
(Figured specimcn)
( ク )
RcmarksI This species is charactcrizcd by thc shghtly concavc latcral rnargins,thc bipartite,
mcdian lo■ gitudinal rib on thc vcntral sidc and thc strongly curvcd transvcrsc folds on thc
dorsal sidc
Reportcd Japanesc occurrcnccs of C. c,′ ,Ю ,ど, are from the Shirnosato Sandstonc and
Siltstone of the Kumano Croupぅ Wakayama Prefecture and thc Takakura Formation of thc Katsuta Croup Thc Shiinosaro Sandstone and Siltstone is assigned to thc N8 zonc,and thc Takakura Formation is to thc N9 zonc. It has bccn kno、 vn from thc Langhian and thc Scrravanian in Europc
Occurrence: Locahtics l and 2
?C′Jο βttι中 /1ο 'Chccchia― Rispoli,1921 (P,.1,Figs 13,14) (〃 'ο (Cr,ο),などゅ 狩ο 'ChCcchia‐
Rispoh,1921,p 20,ng 10
0,ο ′ね′夢2れοr ChCCchia_Rispoh;Sima,1968,p421,ng 7;Robba,1971,p82,p13,ngs 4-6:
Robba and Spano, 1978, p 773, P1 79, ngs l, 2; D'Alcssandro, Laviano, Ricchctti and Sardcna,1979,p 84,pl 16,ngs 55-61;E)'Alessandro and Robba, 1981,p.636,p1 70,ags l-5.
cr,。 sp Shibata,Taguchi and Uiihara,1989,p.38,pl.4,ags.4-6
Matcria1 12 poorly prescrved specilncns.
Description: Shcl small,trigonal, with apical angle of about 25° Latcral margins nearly straight,kceled and comprcssed, ヽrcntral sidc aattish,with a lo、 v,InCdian longitudinal ridge, 、vhich is a little broadcr than 1/3 of thc shcl 、vidth ncar aperturci arcas bct、 veen the longitudinal ridge and thc latcral inargins aattcncd,curvcd vcntratty ncar thc margins Dorsal side sccms to be swollcn, but dctails arc unkno、 vn o、ving to thc poor prcscrvation of the specimcns Extcrior smooth
Mcasurencnt(in mm):
Length130
61
Length100
120
Width
12.6 5.5 ヽVidth 5.0 6.5Ptcropods from thc Miocenc Fuganji Mudstone
Remarks: Thc shen outhne and thc fcaturcs of thc vcntral sidc of this spccics agrce、 vith
thosc of C,′ねケ中 れοF Checcia― Rispoh fI・om thc Mioccnc of ltaly Wcl prcscrvcd mattrial must be availablc for positivc dctcrolination of thc spccinc namc This spccics differs from C. =α
rg'々 'Cク
Sirna from thc samc Mioccne that C.′など中 ヵο,Was Originally described ttom in thc largcr apical anglc, Comparison、 vith thc specilnens of C sp.described by Shibata and othcrs (1989)仕 om the Bihoku and Katsuta Groups shows thc Waji specimens are conspecinc with
thosc frona the t、vo groups
Occurrcncc: Localities l-4
9,ο
力οを
,″α
,(Shibata,1983) (Pl l,Figs 15,16)E′,テοうβ肋れど力陶
(Rang)(nOn Rang,1834),Shibata,1970,p81,p14,ngs.16,171 Shibata(in
ltoigawa,Shibata and Nishimoto),1974,p.192,pl.60,ng,4;Shibata,1977,pl.13,∩ gs10-12.
β′じ力ο cf.う ,肋μttЙ
“
(Rang);ItOigawa,Shibata,Nishimoto and Okumura,1981,p147,ngs
8-10,Itoigawa,Shibata,Nishilnoto and Okumura, 1982,p.296.
ど1/cど,ο ,rοをα″α,Shibata,1983,p.73,pl.2,ngs l_3
α
,ο′
ゎを
,″所 (Shibata),Shibata,Ishigaki and Uiihara,1986,p46,p17,ig,7,Shibata,
Taguchi and Ujihara,1989,p.38,p14,ngs 13,14.Matcrial lore than 100 spccilncns wcrc obtaincd from this arca Most of thcm are incomplctc and are dcformcd.
Description: ShcH triangular, slightly curvcd dorsany in thc pOStcrior part Lateral ribs shghtly diverging in thc postcrior partラ faindy converging in the anterior part, doublc,
gutter―shapcd in thc antcrior part Cross― section near the apcx round Sculpture consisting
of iongitudinal ridgcs and transvcrsc folds,longitudinal rigdes on the doisal sidc three,of、 vhich
the median onc is thc narro、 vcst, converging postcrioriy, broadcr than their interspaccs; a
broad iongitudinal ridge present in the mcdian part of the dorsal sidc,thc breadth of the ridgc about a half of that of the sheH near aperturc,arcas bctlvecn the longitudinal ridgc and latcral margins nattcncdi transvcrse folds closcly spaccd,curvcd,their convex side looking to、 vard thc apcrturc, Apcrturc lip― shapcd. Apical anglc about 60°.
Mcasuremcnt(in mm):
(Figurcd spccimcn) ( ク ) Length 16.8 11.4Width
95
90
Rcmarksi This specics is closcty allicd to thc h
ngじ
′?ο,rフ,(Childrcn),but thc shcll issmalcr, transvcrse folds arc nncr and morc closcly spaccd,and thc mcdian iongitudinal ridge on thc dorsal side is narro、 vcr than thc latcral oncs lt somc、 vhat rcscmblcs C =1/J′οrr′
Silnonclli from thc Miocene of ltaly, but the aperture is narro、 vcr, and transvcrsc folds arc nncr
C・ どわをク〃
''has a bio‐chronological rangc ofthc N8 zone to the N10/12 zonc,occurring most
coHunonly in the N8 and N9 zoncs.
Occurrence: Locahtics l-4.
Subfanllly Cuvicrininae Ccnus C,ソ,ι/肋
,boas,1886
(デ,ソ′θ′ι/1,Sp.
(Pl l,Π gs.17-19) Matcriali 28 transversely compresscd spccilnens`
Descriptioni Shen largC, largcst spcciIIlcn rcaching 18 m■ l in length including a part of
juvcnile spine,bottlc―shapcd, shghtly bulging in thc middle part,posterior cnd of pcrmancnt shcn c10sed by a convcx scptum. Grcatcst diamctcr situated at about thc nliddlc of thc shcn
fl・om which thc juvcnilc spinc is cxcludcd. Bclow thc apcrturc the sheli shghtly consticted in
frontal vic、v. Surfacc smooth
Mcasurcmcnt on mm):
Length
ヽVidth(Figured spccimen) 115 3.1
(
ク) 139 4.6
(
ク) 178 4.3
Comparison: This specics most cioscly rcscmblcs C.gr,Ю βなD'Alessandro and Robba from thc Mioccnc of ltaly,but thc post apcrtural constriction is stronger. It inore or less resemblcs
the l inB C.cο ′,“ηι〃,f.θ ο′,躍η,肋 (Rang),but thc shcll is iargcr and its greatest diameter is
situated morc antcriorly. C.sp frona this arca diffcrs from C,sp. dcscribcd by Shibata and Ujihara(1989)from the Mioccnc Kumano Croup of Wakayama Prcfccturc in that thc postcnor
half of thc shcn is naOre strongly bulgcd.
Occurrence: Locahtics l-4.
Subfa■ ly Cavohn dac Gcnus C,ッο′′/P,,Abildgaard, 1791
C,ソο力η,α うな,た ,″ (Kitd,1886)
(Pl.1,Figs.20-22)
ry/αヵ9α ぅたЙrc′どα Kitd,1886,p.65,pl.2ぅ ngs.29-32;Jansscn,1984a,p.68,pl.5,ngs.6-8.
PteroPods from the Mioccnc Fuganti Mudstone
Cαフο′,れ ,α ,Erβι/P,ヵοどヽ/inassa dc Rcgny,1898,p.84;BcHini,1905,p.34,ngs.21,22;Robba and Spano, 1978,p.778,p1 76,ng.9,pl.80,ng l,pl.81,ng.5;D'Alcssandro and Robba,1981, p 651,pl.75,ngs.1_4,pl.76,ags.1,2.
CαフοJi/P,α αrv′ι々,れοゴvar,う οれο々,θη∫た ヽ■nassa dc Rcgny,1898,p 84,ng.
Cαフοど,々,, c9/,〃 ,, Checchia―Rispol, 1921, p.24, ng.11, sirna, p 428, ng,19; 。bba, 1977, p 612,p1 25,ngs 5-7;D'Alcssandroぅ Laviano,Ricchetti and Sardella,1979,p.91,pl.15,ngs 16, 17,23,28
C,ソο′ど/1,α ,,′ι/tFttο′ど′,々力,ris Rutsch,1934,p.311,pl.8,ngs 6-8.
C'ツ0′ '7Tρ
′α′′rσrね Nomura and Zinbo,1935,p 169,pl.15,flgs.25,26.
C,ソο力刀力 cf.′ρ/″α胸む(Nomura and Zinbo), Shibata, 1970,pl,4, ngs.14, 15,Shibata(in
ltoiga、va,Shibata and NishiFnOtO, 1974,p.192,pl,60,ngs.5-8,12.
Cαッο′,々α(Cαフοrittα
)audeninoi(Vinassa de Regny),Robba,1971,p.100,pl.4,ng 15,pl.5,
ngs l-5,
Qッ
οカカ,α 翻′′″ris(Nomura and Zinbo)ゥ Shibata,1977,pl,13,ngs.16-18,Shibata,1980,p13,ags.19_21;Itoiga、va,Shibata,Nishirnoto and Okumura, 1981,pl.47,ngs 12-15;Itoiga、 va,
Shibata,Nishimoto and Okumura,1982,p.296;Ujihara and Shibata,1982,pl.6,ng 22;
Shibata,1983,p.81,pl.2,ng。 13Cα
ッ
ο
力
,デ, うな
,たρ
″
(Kittl), Shibata, Ishigaki and Ujihara, 1986, p.49, pl,8, ngs.12, 13;Ujihara,Shibata and Saito,1990,p 319,pl.2,ng,7.
Cα
ッ
ο
力η
,,うなレ
rf″,var ′
α
万″
rね (Nomura and Zinbo)i Shibata and Ujihara,1989,p.32,pl.3,ngs.6-8,Shibata,Taguchi and Ujihara,1989,p.39,pl.4,flgs,17-19. MatCrial: 55 poorly preservcd spccirnens.
Descriptioni ShcH sman,lateral inargins roundcd, Ventral sidc as long as、 vidc,inaated,
、vith two radiating BroOVes and faint transverse undulations; ventral lip sharpラ not rccurved.
Dorsal sidc longcr than wideラ slghtly inaated in thc mcdian part, provided with thrcc, very
lo、v, longitudinal ridgcs 、vhich convcrgc posteriorly and conccntric undulations; dorsal lip
slightly bent vcntraHy,projccted far bcyond thc riln of thc vcntral lip. Posterior spinc sman.
Latcral spincs not wcll dcvcloped.
Mcasuremcnt on mm):
(Figurcd specimen;dorsal platc) ( // ;Ventral plate)
(
ク; // )
Lcngth123
83
4.3 Width 8.0 8.855
Rcmarks: Dorsal longitudinal ridgcs of spccimens from the Fuganti Mudstonc arc lower
than thosc Of thc lectotype and para‐ lcctotypc of C.う
ね冴たα
″
(Kitd)ngured by Jansscn(1984a,p15,ngs.4 and 5).Shibata and Ujhara(1989)disCriminatcd Japanesc Miocene spcdmcns with which thc Fuganji spccimens arc identical ttomう
た
,どθ
α
″
, and idcntincd them with c,/,/,勉どね(Nomura and Zinbo)fl・ Om thc Mioccnc Yanagawa Shcll Bed of Fukushima Prefecturc, placing /α/Jrαrぢd in the rank of the varicty of う,s,′ cρチα Thc differencc mcntioned abovc,
AKAGI,S.,SHIBATA,H.AND MURATA,N
however,may fall within thc va ation ofうた,た ,″.
OnC(Shibata)of the authors reccndy exammed four topotypes of r,ガ カrね and 12 spccimcns of it fro■l the Yanaga、va Shel Bcd near thc type locanty. All the speciinens have proHlincnt
longitudinal ribs on thcir dorsal sidc,and thus′ αrirpris sccms to be a synonym ofうね,′θ,´,
Occurrencc: Locanties l,3 and 4.
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Plate l
Figs l,2 CarFrt,′J,sp Loc 4 1a,inner cast, ×3;lb,outcr lnould, ×3;2,inner cast, ×6
Figs 3,4 とどれ,cJヵ,sp Loc 4 3,apical view, ×15;4,umbilicai vicw, ×11
Fig 5 rFyα ′οαガλ sP, Loc 4 ×
61
Fig 6 ?να8JηιrrFゴqpress,Daudin Loc 4 ×5
Figs 7-10 1/f=力￠肋 sp 7-9,Ioc. 1, ×3, 10,Ioc 4, ×15
Figs ll, 12 CrJο θ,7,々pr,Audcnino ll,dorsal vicw,Ioc 3,outer mould, ×3; 12,vcntrai view,loc 4, inncr cast,×3
Figs.13,14 ?C′′ο′r5た丁2′,ο
'Checcia―
Rispoli Loc l Ventral vicw 13,× 35,14,×3
Figs 15,16 α′οめ を
'″'ど (Shibata)Loc l 15,dorsal view,× 24;16,ventral vicw,× 3
Figs 17-19 C,ソJι′,/И sp 17,18,loc l, x3;19,loc 4, ×2
Figs,20-22 C,ソ οriPrl′ うる,rt・,筋 (Kitti) Loc 1 20,dorsal view view,× 22;21,ventral view,× 28,22,
Plate l
