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(1)Distribution. Chrysanthemum. of. MAEEAWA. ye20enSe. ----11一三1生芹｢i什-1手打n甫hT】｢--ま牙¶汀¶｢------. By. Shuzo. Tbe. (NAGAMI,. papers. present. 1954,. Most. data. of the. out. carried. in. at. the. basic. SIIIMOTOMAI,. chromosome 1933). The. of Chr.. number. is. therefore,. species,. is 45. yezoense. it, readers. KITAMURA. (1940),. a. series. Our. material. follow･ing. tubular. showing of heads herbaceous. and provided part. at. plant. in. this. paper. (1) The. will. llead. 且orets. (yellow) and. corymbose. style. The. truncate-cuneate. long. margin,. few. spine-shaped. are. is. at. base, in. and. winged bai一s beneath･. Distributional. Ⅲorizontally follows.･. as. tion. to. 4･5cm. over. folユnd. on. a. irregularly. 1914-'15,. petiole,. ′21;. is. in. length,. cm. creepers. and. stipule,. (2) Scores. of. perennial. at. erect,. form,. covered. and. lower. the. ronnded at. acuminate-dentate and. the. posessing. (white).. in its total. by. the. Distribution. Area. the. south. exhibiting. which. some. incised. absent. one. diameter,. ones. stem,. 15-25. in. the in. 10 ligulate. distributional area of the whole 1at. 36o32'-43o20′ N., long. 139o50′-145o50'E.. is Nemuro,. author. one. con丘ned. is about. moreover,. stem,. be. about. ⅠⅠ. ⅠIorizontal A.. the. of the decaploids the original descriptions of (1921), (1912), MAEKAWA. few branches at the upper part and with (3) In general, the leaf is broadly ovate-cuneate. apex,. its. Japan･. etc.. characters:. numerous. Honshu,. (TAIiARA,. apparently. As for of Chrysanthemum-polyploids. to MIQUEL (1866), MATSUMURA will refer. in. the. 】班aterial Plant. I.. The. of. to. years.. several. past. The. characters. tlle･丘eld-observations. on. poly-. previous. belonging. North-East. to. his. problem･. distributional. of the Hokkaido. based. are. this. decaploids. (one. here. into. the. in. and. the. showing. from. coasts. presented. between. relations. inquiries. some. made. aims. the. on. grows. which. genus),. of. MAEKAWA. yezoense. on. studying. Chrysanthemum-species;. ′55) were. however,. report,. been. years,. _-. Chrysanthemum. has. for. distribution. the. and. ploidy. has,. author. NAGAM:I. Kuzi,. west. unknown,. and. east. will be described limit of distribu; north Nemuro (NAGAM=, 1954)･. species.

(2) S. Nagami. 10. These. results. related. lJ140opOm?lakil142o,tI that. GongenzaきIAIt.Fukkoshi-. the. species. The. Japan.. ●ヽl Aomori■ヽヽ/l. the. on. served. is, therefore,. ⅠⅠachin10thetlttt40｡. one′. genera】】y. in. seacoast. the. of. the. Tbe. 凹t.Urcratl. Honsbu-population. is. in spread 36o29'-41o30'N.,. IwaizumilI･TJodILisaki･. of the. folloving. tlle. north Gongenzaki,. Hiyatozii,a/,lN//Ilpresent. sendaioL･J/′. and. Shiogama. 38o. the. lation.. /_I.. TakaTrlatSu(. the. Eironゾ!. At. of the. Il /I //. of it. Ea*arago,/I/ 1k叫､-_”Izuki 3占o. is. yes:OenSe. 36o. from. follows. as. is endemic. is. Kuzi. Fig.. 1.. Map. in. of. and. Chr.. (Y). yezoense. North-East. former:. thick-lined. that to. the the. (DoBZHANSIくY, Among. populations･ and. between. is obtained･ But. especially. In Fig･ in. of. coast. (3). coast.. Pacific. the. limit south (5) The Honshumay. be divided. populations-the. north-, (6) The. south-populations.. on the coastal appears north-population from area Wakimoto to Deto, the middle from Hirono to Kuzi. south. discontinnity Tsugaru. 1951). is. are. these. are. seen. 2 shows. us. included long. Thus. the partial. three sandy. discontinuity. of Honshu.. the the. is continuously. spec''es. north-population. distribution. between. found. latter. the. of. Straits･. the. general. the. the. and. small. and. middle-,. latter:. side.. of the species. three. Japan,. of Distribution. owing. population. 1ine･. The. to Shiogama,. lt is evident. above,. into. The. Japan.. in the. is the. Japan-Sea. the. wbic壬1 We : (1) Chr.. Paci丘c. the. on. limit. the. population (N). n軸onicum. coast.. Continuity. isolation. of. coast.. Hachinohe B.. horizontal-. Chrysanthemum. Hohshu,. oblique-lined. -from. the. showlr)g areas. are. plant to North. the Japan-Sea. on. south Wakimoto (4). side.. 142o. most. tlle few. and. distribution. the. occnrrlng. foresaid.. as. I. Choshi. clones limit. south. of the coast. one. (2) It. genus.. Tokyo. the. 2,. 1 and. able to conclude. are. Nakaminato. few. a. of tbe丘eld-studies. results in Figs.. glVen. Kawarago,. of. llim. by. to. popu-. found Thus. revised. Tbe. The. is designed. south. decaploid. was. west. above-mentioned. recently. ;. Todosaki.. cotlrSe,. Kuzi,. author. Kuzi,. south. east. of. paper,. investigate. plant 1at.. area:. long. 140o14'-142oO4'E.. limit is Omazaki,. 140o. It. coast-cbrysan-. I▲omoto. ー. ob-. area.. tbemuⅢ1S.. 40o. 38o. North. to. is. species. wakimotoー､ヽ▲eboshti. indicate. above. is endemic. happens. in. species-. small scaled geographic HokkaidoHonshuand. small beaches. populations where. distl･ibutedalong in. the. the. no. as noted decaploid. the. coast-. decaploid-population,.

(3) Distribution. Chrysanthemum. of. Maekawa. yezoense. l140ooⅡ1aZaki. in. (n-45). Honshu,. 141o. ll. Japan. 142o/ / /. I. ohatashiriyazaki/. ＼. ′. Chikagawa′. ､ヽ･Takanozlaki. Gon7enzakiヾTo. zawaoippe/. 41oWakimoto､_lio. minato▲Mt･Fukkoshi-ebos. ヽヽ. 41E'. Detoi. ■---I■. iヽ一l Nohejiy. .Nona. Aomo■ri. hi. I. ＼､＼､ Aomoriprefecttlre. ヽ且achir,.he. ＼.1Tan-es:ihi Y... ･:'●････-･･'･･.:･･ ●Ip. ･.･:=＼乱. Akitaprefecture. ＼ニl. 40o. 40o. loも. 肌口rera▲rd,n.t.i. ■■●. I. oAkita:Morioks′Iw.aizumlBl I-I 'Todosaki'I. Iwateprefecture.一il. Fig.. 2･. (Y). yezoense. 140o,.1｡IJl.{.i. Map. Chr.. and. horizontal-distribution. the. showing. (N). m'J,1,one.Cwm. ⅠⅠⅠ. Vertical A.. Distributional. Chr.. ye20enSe. 1.. by. revealed was. formed. by. 1955, One. also. i)istriblltion. (hill type),. on. and. sandy. coasts. top. of Mt.. in. 丘elds. (丘eld type),. (sandy-coast. on. rocky. type).. C. TAI(AEAS‡‡Ⅰ, Forestal. Fukkoshi-Eboshi. OfBcial,inthe. (507･8m), which of 1955. This was. autumn. Mt. Urera Town. (625m) of lwaizumi also found S. MuRAI, Forestal Technical OfBcial, and confirmed is indebted. of the. most. frequently. and. to. their. distinctive :. plant-body. branches. short-stemmed. the. on. appears. on. who. diminutive. oてnO. hills. On. type),. decaploid. species. in. of Chrysanthemum Japan･. Hill Type:. The was. areas. Honshu,. Area. grows. (rocky-coast. coasts. North. in. one. observed, bushes on. (1) The or. a. stem. few. occupying. the. summit. (2). the. allthor. is its this type yezoense･of cm in length, bearing few. 10-15. Chr.. of. is about. heads･. the. in-. kindnesses.. tbougbtful features. by. The. Dwarf. hill-top. plants ground.. (Mt. Fukkoshi-Eboshi). head no with It grows in or. on. ･the. are. the. rocky･.

(4) 12. S.. cliffs (Mt. the. external. geological Mt. Urera.. Fig.. Chr.. the. coast. 丘eld of. of the. 3.. Upper. and. cm). slender.. lanky. their. in. well. as. is short,. a. Urera rock. mother. the. and. constituting. (Y). yezoense. Urera.. to. the. the. thefields. roadside. underbush of. in. this. as. plants. exist. the. underbushes. of. area. the. orin. rocky-cliffs. from. the. covering. the. on. especially. coast. south-population.. following. features. The. branches. and. subterranean. of. forests. underbush. coast. are. observed. are. stems. bear. is long. the stem. :. Most. numerous.. few. a. (20-. beads. and. show. Type.･. As. ye20enSe. Mt.. of. the. the. Rocky-coa占t. itse一f. Mt.. form.. 3.. Chr.. the. between. relationship. Chrysanthemum. cliff. frequelltly. half. south. occurring. plants. is. Lower:. the. some. distributiI唱On. :-limestone. Urera. of. appears. Morphologically 30. Mt.. :. of h'ills down Such forests.. the. place. is. there. Type:. yezoense. half. that -noticeable of the decaploid. limestone-crevice. the. Field. lower. is. characters. conditions. at. 2.. It. Urera).. Nagami. as. the grows. case On. of Chr. the. bacijicum. sea-cliffs.. dominantLSPeCiesinthe. tenacious,. and. few-branched,. The. coastal while. NAI(AI. (n-45), the of. plant habitats. the. this Of. course. subterranean. great type. majority often. the stem. reveals. aerial runs. of. stem very.

(5) Distribtltion. Fig･. Chrysanthemum. of. 4･. ye2fOenSe (Y). Chrysanthemum. in. 4.. the. the. Type. species. is also. relatively. small. The covers. individuals, Takahagi,. Such. Fig.. 5.. (Aomori. ln to. the. or. on. of. the. on. observed. that. seems. glossier･. upper-surface. coast. sandy. the. under. is usually. Its plant-body. areas.. a. it. Thus. plant-body･. their. (Aomori. TAI(A壬IASBT･). well-developed,. plant. is characterized. winged. ;. and. follows:. as. its aerial. or. it. sea-cliff, though has. met. indicating. Omoto,. Tanesashi, its lamina. subterranean. tall. some. is broad, stems. nu-. branched.. merously. B.. C･. :. brilliant ; petiole. and. Oippe. of. 13. Japan. Honshu･. author effects of the salt spray. on 70cm in length･ the.coasts of about. each etc.. the. on. in. The. above-mentioned. thick,. falls salt spray leaves thicker and. the. Sandy-coast. the,field. (Courtesy. prefecture). Sometimes rarely. the spray makes. (l]-45). Maekawa. yezoense. hillside. the. yezoense. (Y). on. the. sandy. (Courtesy. Prefecture)･. Continuity general. Chrysanthemum. of. coast C･. of Tomari. TAKA‡iASIiI･). of Distribution. Chr. But. hill-tops.. is. yezoense Lit occupies. The. isolated. continuously. distributed. discontinuously decaploid. on. of. Mt･. the. from. the sandy coast inland-cliffs, in the丘elds. Fukkoshi-Eboshi. is. found.

(6) S,. 14. at. 6km's. aboし1t. 16km. from. almost. distance. the. same. IV.. and. Natural. in. shown. Kuzi.. we. are. that. and to. able. Mt.. of the. observe. facing. inland-cliffs. Other. tlle 1,. Fig.. Urera. about growing. plant River. Omoto. the. an. has. area. overlapping. between. tJeZOenSe. a. I. distribution. of. MATSUMURA. found. ever. Chl･y8anthemum. Chrysan-themllmS-YezoellSe. (FRANCtI.). one. no. betweeII. flybrid盲zation. n軸om'cum. But. the. along. and. Chr.. But. coast,. Omoto.. and. As. Paci丘c. the. coast.. conthuously. lwaizumi. from. Nagami. (n-9). ranges. inter-speci丘c. natural. Chr.. of from. yezoense. Hachinohe. to. between. the. hybrid. tlVO.. At. Mizuki,. (n-45)･Chr. mixed. We. is the. Paczjicum. depressed. have. in. been. the. from. The. the. Chr.. an. overlap. of. cultivated two. has. the. first. describing. that. the. essentially. it. Chr.. as. similar. one. The. present. other. south Kuzi,. (1) The. north. Japan-Sea. side. linlit of. (3) the. the west. the. the. and. is Gongenzaki,. plant. (the. coast. chrysanthemums genetically'.. It is. Chr.. cliffof. a. fact. be-. yezoense. the second. others; reporting. hybrids. two. the. is dotted. and. between. hybrid. that. Chr. across. (1949), and. and. Mizuki. came. author. natural. KITAMURA,. or. crowded. chrysanthemums.. Surnrnary. Honsbu-population. Horizontally:. the. the. mixed growth NAl{AI (n-9) or. (1940), OKABE. is 72.. The. other.. llOrizontally. and oll.. a. between. hybrid. mかaioj2'mense. both. paper,. hand,. plant is Chrysanihemumye20enSe the Chrysanthemum-polyploid. in of the decaploids Tile distribution of. three,. in breadth,. m2. and. MAEKAWA. material. 2.. this. 100. Various. et. Miyatozima. KITAMURA. V.. l.. these,. pac2jicum-area. Honsllu. other. natural. by. and. the. obtained. is abotlt. MATSUMURA the. At. each. cultivated. short-stemmed,. cytologically. North-East. number. to. hybrids,. vigorous. both. On the. chromosome. Of. of. future.. (n-9). and. place.. from. Chr.. the. of Mizuki. place. the. cbrysantbemums. also been observed. somatic. these. and. species. the Paci丘c Ocean.. in nowhere Chr. Makinoi. distribution. the. with. found. MAKINO. the. are. in. the. never. Judging. from. investigated. reported. ye20enSe. (MAEINO). boreale. being. are. results will be Tbe atltbor has. tween. facing. is. yezoense. example. there. its small-cloned,. the. Chr. an. offer. and. overlapping. of Choshi).. The. of coast-cliffs. portion. for. to Mizuki. of Cbosbi-Omaezaki-Torishima). occupying. here. see. portion,. (north. (n-9),. this. with. remarkable. transplanted. one. Chrysanthemum,. of. to. contact. Besides. of Hasaki. coast. species. moreover,. able,. most. features.. northern must. are. coming. cllrySantbemums. three. (n-45)･Chr.刀妙om'cum. pacl'jicum. growth.. Chr,. Kuzi,. of. north. (n-45),. series.. of the. species. is investigated. in. vertically. limit is and. of. Wakimoto,. (4) the. di.stribution and east. that. is. Omazaki,. of. the. is Todosaki.. Paci丘c. (2) the side. is.

(7) Distribution. Vertically: coast,. the the. on. and. and Urera. the. decaploid.. Honshu,. the. The. north-,. are. able. is. one. On. area.. of the the. of. 丘eld,. the. on. Fukkoshi-Eboshi. isolated. small. w-ill be. species. coast. Japan. the. Mts.. the. observe. majority. great. plant. the to. Honshu-population. middle-,. 4.. ､15. rocky. (507.8m) of. poptllations. found. the. on. sea-. North-East. in. chrysanthemums. hybrid. ranges. betw･een. Chr.. But. one. into. of the. coast. three: forests,. south-population. Chr.. no. Chr.. and. yezoense. but. niP-. inter-specific. natural. obtained.. and Chr. boreale (n-9) are has ever the natural revealed. (n-9) no. been. ever. the. Kuzi,. to. divided. underbushes. between. Hachinohe. has. two. the. Makz'noi. ense-area.. from. be. may. the in. found. distribution. overlapping. (n-9). ponicum. species. In. frequently. are. plants. The. the. of. south-populations.. and. the丘eld-type. found. Chr.. the. near. hybrid. between. yezo-. the diploids. decaploid.. the. Moreover. following. the. Chr.. (n-45),. pacijicum various. hybrids. between. Mizuki.. Among. them,. four. taxonomically. and. cytologically. the. materials. examined. are. nil)1)om'cum, and them. are. found. are. reported Miyatozima. of. by. Th｡. Genetics. 1951.. the. and. Of. course,. 100. m2). and. yezoense. to. tllOSe Which. b女KITAMURA. (1949), each. of. of the. two. (1940) using. Cited. Literature DoBZIiANSKY,. K汀AMURA. OE-ABE. Chr.. Similar. Chr.. yezoense,. (about. between. essentiallyT. m13)aiojimense. 2n-72. as. sea-cliffs. hybrids. are. Chr.. :. chrysanthemums. the. on. vig･orous. plants. Chr.. named. Mizuki. at. cultivated. observed. the. Such. chrysanthemums.. cultivated were. coast. we. The. in. hill, in. the. on. appears. in Honshu,. (n-45). Japan.. 3.. and. sandy. Maekawa. yezoense. species. (625m),. Thus. clifEs.. Chrysanthemum. of. origin. of. New. species,. York.. Columbia. Urliv.. Press. KITAMURA,. S. 1940.. Compositae. MAEKAWA,. T.. Trans.. 1921.. MATSUMURA, MIQロEL,. N. 1866.. W.. NAGAMI,. 1912.. OKABF_,. S.. Sci, Rep.. 1949.. SHIMOTOMAI,. STEBBIBS,. Jap. N.. G.L.. Musei. S°i. Rep｡. 1955.. Botanici. Genet.. 24:. Variation. Ludguno-Bata∇i. National. See.. Univ. Univ｡. 1-12. Tokyo. (ⅠⅠ). (ⅠⅠ).. Sec･. ⅠⅠ,No･. ⅠⅠ,No･. 3:. 36-48･. 4･: 12125･. 79.. Hiroshima. Jour.. 8:. Soci･. National. YokohaITla. Jr. 1950.. Hist.. Japonicarum. Yokobama. Jour.. 1933.. Nat.. Plantarum. Index. Annales. S. 1954.. Kyoto. (ⅠⅠ).. Japonicae. Sapporo. Ser,. Univ. and. evolution. B, in. Diy. plants.. 2,. 2: New. 11100. York. Columbia. Univ.. Press.. TAIiARA,. M.. 1914J15.. Bot.. (8), (9), (15), No･ 1921.. Jour.. Coil.. Mag. 340,. Tokyo.. 29:. No1. 336,. (491-493),. Univ.. (7),. 337-339,. _(6),. (48-50)･. Sci. Imp.. No･. Tokyo.. 43:. 1-57..

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