අਬȆକ̥͙̹ͣͺΐͺȆͺές
ᨆ͞ˠ˽ɹॶᩃᨆɴȲȿʳᰖɹᅊਖ਼ݭଘգᅎɭ
зɹྫ֝༎֪
࣬ǵᅒǵ౩ǵχ⏋
*ǵ߂ǵីǵѵǵে⏋* ೠǵؤǵᅓǵ᱗**
Habitat Use of Fishes and Fishing Activity
in Plain Area of Southern Laos
IWATA Akihisa,* OHNISHI Nobuhiro* and KIGUCHI Yuka**
Fish fauna, habitats of aquatic bodies, habitat use of fi shes, and local people’s fi shing activities were researched in the Ban hiang River system, one branch of the Mekong River, southern Laos.
Fish fauna of this area comprised 10 orders, 31 families and about 158 species. Aquatic bodies were divided into two large water areas, : permanent water areas and : temporary water areas, and habitats were classifi ed into 14 types. About 56 species of fi shes were found that clarifi ed relation between their ecology and habitat, and 12 habitat use patterns were recognized. Fishes inhabiting paddy fi elds were noteworthy in that they were not spontaneous arrivals from permanent water areas, but particular species specifi c to this habitat.
A total of 26 types of fi shing gear were found in this area. Local fi shing practices are based on a sound indigenous knowledge of the ecology of fi shes. There were some differences in sex and age among fi shing gear users related with safety, distance from village and so on.
The implications of expansion of irrigation and aquaculture toward aquatic resources management system were discussed.
1ȅ͉ȁ̲ȁ͛ȁͅ
ǵആˏ˻ˏɻ͐Ͳ˽ͻͲຩҌɴఌᨗɇʶ⏋ϗ࡚ɭᮒ࡚ȵȲʱɲɐៀհɌȦᕶးɹ̀͌͠ɹ μɫзȵᅊ༎Ɍɩȷɛ⏏ᮒ࡚ȵʢɛʰɐ߂ᩄɹອɻɄɹܩݖɴȲȿʳΥ๖ᅋɹݜॻᅊɫ
* ϴᨊ߂߂ˏ˻ˏˏ̶͠ˡܩݖድድዀ⏋Graduate School of Asian and African Area Studies, Kyoto University
ȡʳዝсˁఌȭ⏋ɗʶɭؾಁɴօʱՏɇʶɛዛɲອݖᄩޅɻ⏋߃းອᏙɭȡȦʚɤɩտ ᰩɭɌɩɹອᅋ႒ᅊᅋɴჼܧɫʢ߂ɲᢨˁɌɩȦʳ⏏ ǵആˏ˻ˏɹʛɲʰɑ⏋ɗɹៀฐɴȲȦɩγᅠధɹ߂ໜाɫȡʳ͍˱Ͳໜɻ̜̍̾̒ɴ ྇༑ˁеᒞɌɩᮖˁάʱ⏋ɗɹ༑ݖɴμܕ⏋͊͑Ͳ͇ͻ⏋͞ˠ˽⏋̊˓⏋ˡͲ̓˻ˏ⏋̾ ̜̠͌ɹ 6 ˡܕˁيʛɲȵʰɴ̜̠̾͌ɹ͍˱Ͳ̊̚͡ɴɗɹໜؤˁᬖȼ⏏যᬏᤂᮐ 4,620 Ρ⏋༑ݖᯆዩᏝ 795,500 km2 ɹμɴᏝ 6,500 ΨϺɹϺǾȵಬʰɌɩȦʳ 1996; ̠̊Ͳ 2002⏏ ǵ͍˱Ͳໜɹ༑ݖɴيʚʶʳ 6 ˡܕɹȩɠ⏋͞ˠ˽ɻɗɹܕܤɹᏝ 90⎾ȵ͍˱Ͳໜɹᮃອݖ ɴيʚʶɩȦʳ⏏Ʉɹܕɻ˓Ͳ̝˹̠ץऄԒ᭜ᨆɹആɴеᒞɌ⏋ɗɹܕޅˁμܕ⏋̜̠̾ ͌⏋͊͑Ͳ͇ͻ⏋̊˓⏋ˡͲ̓˻ˏɴஞɌ⏋ᯆዩ 236,800 km2 [Claridge et al. 1997]⏋1995 ॷɴ ȲȿʳϺؤɻ 450 ΨϺɫȡʳ⏏Ʉɹܕɹ 1998 ॷɴȲȿʳອᅋᅊᅋᩄɻ 42,800 t ɫȡʱ⏋ᕶ ᐌრɻෑʠɩᲞȦ⏏2000 ॷɫɹ͞ˠ˽ܕ 1 Ϻȡɛʱɹॷᬚᰖ༥ᢹᩄɻᏝ 8-10 kg ɫȡʳ [Modadugu et al. 2000]⏏ອᅋ႒ɹգᅎȵᲞȦɄɭɫؿɲೣɫ 1998 ॷ⏋ܕ 1 Ϻȡɛʱɹ ॷᬚອᅋ႒њᐌᩄɻᏝ 6.4 kg ɫȡʱ⏃ອᅋংకᨆНᅙᠩ֪كՕഓవ 2001ʭʱ፤Տ⏆⏋͞ˠ ˽ɴȲȦɩອᅋᣂ྇ȵෑʠɩᩂឹɲ৲ւˁഗɛɌɩȦʳɄɭȵࢅ౫ɴჿៗɫȷʳ⏏͞ˠ˽ܕ ɹອᅋ႒ɴࢪɐʳћࡔɻɄɹʭȩɴᲞȦȵ⏋ˁೳషɭɌɛᇍᨥᰩڃ̮ͻ̚˪ɻ႕ɴᩂឹ ɫ⏋Shoemarker et al. [2002] ɻȋ͞ˠ˽ϺɹᎦɴɦȽፀϠɹਖᮛᰩషڃȍɭᝠჼɌɩȦʳʓɰɫ ȡʳ⏏৳ʰɻ͞ˠ˽ᨆɹ̫Ͳ̶ˎ˓ा༑ݖɴȡʳˁёɴЂɌɩȲʱ⏋ɄɹᠶయȴʰᏤ ፤ɐʳɭ⏋Ʉɹܩݖɫɻ 1 ॷ 1 Ϻȡɛʱ 6.2-9.4 kg ɹ̮ͻ̚˪ˁᰩʏɩȦʳɄɭɴɲʳ⏏̮ͻ ̚˪ɹೳషɻзɹᅊ༎ܞɫʶɛ˱˓ᰖ⏋̝˻͚˗ᰖˁيʞμࢸ০ɹɫȡʳȵ⏋ɗʶʰ ɻʓɭ˃ɰɹݭغອᅒʦɗɹٞᦇɫᮃɇʶʳʢɹɫȡʳ⏏ ǵΥ⏋ອᅋᣂ྇ᅊᅋᩄɴ߂ɲ৯ᯱˁؖʖɐຂϟɻ͞ˠ˽ɴȲȦɩʢɇʚɊʚɲ̺ͦ ˻˙˪̜ȵ⏋ςɴܕลᬠςࢷɹ০ɫࣙᬖɌɩȦʳ [Modadugu et al. 2000]⏏ЁɹɭɄʸ⏋˱˓ ʦ̠˓̘͡ˑ̴͞ˏፈȵຂɇʶɩȦʳɴɐȹɲȦȵ⏋Ё⏋ঃ፴ܑɴຂˁᦶଁɐʳɄɭ ɴɲʶɽ⏋ჼܧആˏ˻ˏࣳɹعܩݖɫሾʰʶʳ˛̲ຂɹʭȩɴගകᣂ྇ɴ߂ɲ৯ᯱˁ ؖʖɐɽȴʱɫɲȼ⏋ຂɹዛᗈȵᩃɴఔ༑ɇʶʳɄɭɭȡȦʚɤɩ⏋͍˱ͲອᏙɹອݖ ᅊᏙɴᩂ߂ɲ৯ᯱˁήȭ⏋ܧഀອᅋᣂ྇ȵᣒᇕᩄᇕɴ߂ȷȼࢅɐʳɄɭȵઉɇʶʳ⏏ ǵອᅋᣂ྇ɹ୫ᐚᇕգᅎˁᓒȭʳȩȭɫ⏋ʚɑࢪᢒᅊ႒ȵɰɄɫᅋɌɩᅊᔒɌ⏋ɰɹʭȩ ɲݭଘɫᅊ༎رˁԃȩɐʳɹȴɭȦɤɛݜኄݬȵέخ๕ɫȡʳ⏏ᦏॷɴȲȿʳ͞ˠ˽ɹᰖ ɹՕᰖᇕድɭɌɩɻ⏋Taki [1974; 1978], Roberts [1993], Roberts and Warren [1994] Ȳʭʃ Baird [1999] ȵȡɁʰʶʳ⏏ɌȴɌ⏋մ 2 ᓔɻ͡ˏͲ̺̫͞Ͳȴʰʹˑ˛Ͳ̍͑Ͳᦏ᧿ʚɫɹᨆɭ ̮˪́ȴʰ˱ͻͲྡٞᦇɹᨆܩݖ⏋ 3 ᓔɻ˱ͻͲྡٞᦇɹᨆܩݖɹʛɭȦɤɛʭ ȩɴ⏋ᠭീȵ႕ࡴɹܩݖɴᭉࡴɇʶɩȦɛȵ⏋Kottelat [2001] ɴȲȦɩ՟ʠɩɄɹܕԃкɹ
ᰖሂȵᐨᒧɇʶʳɴᕸɤɛ⏏عዛᰖɹᅊਖ਼ݭଘ᧧ିʦው֪ɴᬠɐʳᠭീɻ Taki [1978], Roberts [1993] Ȳʭʃ Roberts and Warren [1994] ɴʭʱɲɇʶɩȦʳʢɹɹ⏋ᅋݭଘʦІɹᔒଁ ݭଘɭᅊਖ਼ݭଘգᅎɴᬠɐʳݬɻӱՕɫȡʳɭɻៜȦᮑȦ⏏Ꮫ [1997] ʦቄພ̝͞͡ [1990] ɻആˏ˻ˏɹཁອɹᅊᇕៗᠦˁɤɩȦʳȵȦɑʶʢෙᠦᇕɲԒࢅɴᅥʚɤɩȦʳ⏏ Mekong River Commision [2001] ɻ͞ˠ˽ˁيʠɩ⏋͍˱Ͳໜ༑ݖɴᅊਖ਼ɐʳςឹອᅋዛ 48 ዛ ɹው֪ɭᅋɴɦȦɩݬٚɌɩȦʳȵ⏋ᅊਖ਼ݭଘɹᏳɲ៨ᥔɭɗʶʰɹգᅎɴɻៜؖɌɩȲ ʰɑ⏋᧧ՏɇʶɛዛʢзɹᅊΫᩂឹɫȡʳࢸμ݀ɹዛᰖɻʓɭ˃ɰيʚʶɩȦɲȦɹ ɫΫᦚɌɛʭȩɲݜኄᇕݬȵʓɭ˃ɰɲȦɭȦȩɹȵჼႥɫȡʳ⏏ ǵ͞ˠ˽ɫܩݖድˁȩ᭬ɴອᅋ႒գᅎɭȦȩᇿȵෑʠɩᩂឹɲድࢪᢒɭɲʳɄɭɻΫ ៨Ɍɛဣȴʰʢᠶˁ৺ɛɲȦ⏏ɗɹμɫʢྫ֝༎֪ɻϺᬚɹۊʛɭອᅊᅊ႒ɹᅊɭˁᐄʉஞ ဣɫȡʱ⏋ອᅋᣂ྇ȵᣒᇕᩄᇕɴɌɛʱ⏋ʚɛ⏋ᅊ০ʦອᅋᣂ྇գᅎ˹˽̘͌ɴ ȵᅊɍʶɽ⏋ɗʶʰɹȵྫ֝༎֪ɴሁஞᇕȴɦቁಁᬚɴ৯ᯱˁήȭʳ⏏ɇʰɴຂϟ ɻອᅋᣂ྇ɴࢪɐʳዩෑᇕɲϺဤɹɻɛʰȷȴȿɭɌɩ⏋ɗɹۊဤȵਃഀɹᅊ০Ȳʭʃອ ᅋᣂ྇գᅎ˹˽̘͌ɴɰɹʭȩɲ৯ᯱˁʢɛʰɌɩʪȼɹȴᇿɇʶʳ⏏ ǵೣዣɻᘿᓔʰȵɄɹ 6 ॷᬚᠭീˁɤɩȷɛᨆ͞ˠ˽ɹ̫Ͳ̯˛Ͳा༑ݖɹॶᩃᨆɴɰɹ ʭȩɲᰖȵᅊਖ਼ɌɩȦʳɹȴˁኍɐɭɭʢɴ⏋ɄɹܩݖɴȲȿʳອݖᄩޅɹךՕˁȦ⏋Υ ಁᇕອݖˁμਕɭɌɩ⏋ᅋݭଘɭІɹᔒଁݭଘɴᇿɌɲȵʰ⏋ɗʶəʶɹዛȵȦ ȴɴ҅Ǿɹᅊਖ਼ᄩޅˁգᅎɌɩȦʳȴ౩ʰȴɴɐʳɄɭˁፀ 1 ɹᇿᇕɭɐʳ⏏ፀ 2 ɴɄɹܩݖ зɹྫ֝༎֪ˁΫ៨ɹݬɴݜɧȦɩ៨ᥔɐʳ⏏ɇʰɴ⏋ᦏɹອݖᄩޅɹɭຂϟ ɹಔؖȵЁɰɹʭȩɲ৯ᯱˁʢɛʰɌɩʪȼɹȴɴɦȦɩៜؖɐʳ⏏
2ȅऔ༹༷̤͍͢औশ
ǵᠭീܩݖɴȲȿʳᰖɹᅊਖ਼ݭଘգᅎɭྫ֝༎֪ɹᬠѥˁ౩ʰȴɴɐʳɛʠ⏋߂աɌɩГά ɹʭȩɲᠭീˁɤɛ⏏ (1) ᰖɹᮃᠭീ ǵᘿᓔʰᕶʰȵହᐨ⏋ଝᐨ⏋լɌᐨፈɹྫԍɴʭɤɩອᅋ႒ˁᮃɌ⏋ฌೣˁсଁɐʳɭɭʢ ɴܔࡴሁɹฌೣˁԠሎ৯Ɍɛ⏏႕ɴଝᐨɻᐨᇿɹࢸɇȦʢɹʢьᅎɌɩІɹᮃɴ ֗ʠɛ⏏ؾಁɴܩݖзȴʰؿዌ⏋գᅎ⏋ѝൔ⏋য়᠍ᅊ႒ɹᅊ⏋႕ɴᅋݭଘʦଁᬏɭው ֪ɴᬠʼʳᅊਖ਼ݭଘፈɹݬˁʳɛʠɴᓬʱᠭീˁɤɛ⏏ (2) ྫ֝༎֪ɹჼݭᠭീ ǵᠭീܩݖɹзȵອᅋᣂ྇ˁɌɩȦʳჼݭɴȷ⏋႒⏋ᓔ⏋⏋ݭ ଘˁԠሎɫ៨᪼ɐʳɭɭʢɴ⏋ᓔȴʰؿዌ⏋գᅎ⏋ѝൔ⏋য়᠍ᅊ႒ɹᅊ⏋႕ɴᅋݭ ଘʦଁᬏɭው֪ɴᬠʼʳᅊਖ਼ݭଘፈɹݬˁʳɛʠɴᓬʱᠭീˁɤɛ⏏(3) ॑ݭᠭീ ǵᠭീܩݖɹɫ॑ݭɴȷ⏋ɗɹݭଘɴι˃ɫȦʳອᅋᣂ྇ˁԠሎɫ៨᪼⏋ؾಁɴɗʶʰˁ ȩϺȴʰؿዌ⏋ݭଘ⏋⏋ᓔ⏋գᅎ⏋ѝൔ⏋য়᠍ᅊ႒ɹᅊ⏋႕ɴᅋݭ ଘʦଁᬏɭው֪ɴᬠʼʳᅊਖ਼ݭଘፈɹݬˁʳɛʠɴᓬʱᠭീˁɤɛ⏏ (4) ᅊɴᬠɐʳɫɹᓬʱᠭീ ǵᠭീܩɹɴȷ⏋ɗɹɹзɴعዛɹᅊ⏋႕ɴྫ֝ɭᦅകɹᬠѥ⏋ອᅋᣂ྇፬ ჿɹ⏋ຂϟອᅒຂɹံ⏋ɗʶʰɴࢪɐʳзɹᓒȭɲɰɹݬˁʳɛʠɴ ᓬʱᠭീˁɤɛ⏏ ǵዛᰖɹؾࡴɻ৯ɇʶɛԠሎ⏋ʚɛɻฌೣɹᔀሗɫ៎࢘ɫȷʳ০ᣒˁʢɭɴ⏋Kottelat [2001] ˁؒᓒɴɌɩʼʶɛ⏏ɛɞɌࡸкᰒʦᲓൔɹ႕ਐˁᠭʏʳɛʠɹ᥉˛̒˪˽ᐻខᒞɫȡ ʳ̶̘̅̒˪˽ˁьᅎɌɲȿʶɽɲʰɲȦʭȩɲᏳɲ៎࢘ɻేᇕಁᬚᇕɴέخᔨɫȡɤ ɛɛʠ⏋Ё܌ɹዛɴᬠɐʳؾࡴᐄഗɻȡȼʚɫಫࡴᇕɲʢɹɫȡʳ⏏ฌೣɻɐʏɩЁɹԊ ؾድɹɛʠອᅊᅊ႒ᣂ྇ድ́Ͳ̊ͻ (Living Aquatic Resources Research Center, LARReC) ɴѳ ፬ɇʶɩȦʳ⏏ɲȲ⏋Ё܌ɹᠭീɻ͞ˠ˽ܕጓ߂ (National University of Laos)⏋͞ˠ˽ܕጓᦅ കድଘ (National Agriculture and Forestry Research Institute, NAFRI) ɴȡʳອᅊᅊ႒ᣂ྇ድ́ Ͳ̊ͻɹ៰خɹʢɭɴʼʶɛʢɹɫȡʳ⏏ ǵᠭീܩݖɹອݖᄩޅɹᅊਖ਼ݭଘȲʭʃໜा০ɹךՕɻܤೠЦᬠីఌᨆᑂ [1998] ɴȲȿʳ ៨ᥔϟˁݜኄݬɭɌɛ⏏ᇿȲʭʃዀɹೣᝠ៨ɻΫᩃܮೣ1999ɴਃɤɛ⏏ ǵᠭീܩ⏋ᠭീܩɹᄩޅɹෙᅭ⏋ᠭീಁȲʭʃᠭീɻГάɹɭȲʱɫȡʳ⏃ܒ 1⏆⏏ (1) ̩̠̜͌͞Ͳ ǵ͍˱ͲໜɹΥ߂ఌ༑ɫȡʳ̫Ͳ̯˛Ͳाά༑ݖɹ߂࣐ುܩဣɴеᒞɐʳɫ⏋̍͑ͲͅͲ ाɹغ༑ဣȴʰᏝ 2 km ά༑ɴȡʳ⏏ధॷɴ 1 ɻ༇ອɴʛʚʼʶ⏋ዝсʦɗɹЈɹс႒ɻ߂ ଠˁ؝ȿʳ⏏ೣ༑ɹໜा০ɻဈɹᨆՕɹອཉȵཉȼ⏋ອᯆȵጓɛɲȦ Bc ݀ɫ⏋ɗʶɴ ɦʰɲʳໜᤄྲྀʦȵȡʳ⏏̫Ͳ̯˛ͲाȴʰͅͲ̺ɫளອɌϗ࡚ဌྰˁȩ⏏ᮒ࡚ɫʢᅒɴ ອɹɲȦȵᐚȼɭʦɻʱளອဌྰˁȩ⏏1999 ॷ 7 25 ⏃ᮒ࡚⏆ɴჼݭᠭീ⏋1999 ॷ 12 17-18 ⏃ϗ࡚⏆ɴ॑ݭᮃᠭീˁɤɛ⏏2001 ॷ 3 10-12 ⏃ϗ࡚⏆⏋6 30 Ȳʭʃ 7 1 ⏃ᮒ࡚⏆⏋2002 ॷ 8 10-11 ⏃ᮒ࡚⏆ɴʍɹᓬʱᠭീ⏋ჼݭᠭീȲʭʃᮃᠭ ീˁɤɛ⏏ (2) ˰Ͳ˱˪Ȳʭʃɗɹᦏ᧿ ǵ̫Ͳ̯˛Ͳा༑ݖɴȡʱ⏋Υᅱά༑ɫүȴʰغ༑ɐʳɄɹाɹϠ๖ఌ༑̍͑ͲͅͲाɹά ༑ᨆɴеᒞɐʳ⏏ੈχᇕອݖɹ̍͑ͲͅͲाೣ༑ᨆɹʓȴ⏋ᮒ࡚ɴɻɄɹाɴɦɲȵʳ߂ࢸɹ ʦໜᤄྲྀ⏋ੈχᇕΥಁᇕᏳ༑⏋ະ⏋༇ອകȵȡʳ⏏႕ɴ˽˓ɭ٧ɽʶʳ߂ៀฐɲɻ
ອᅋΫᩂឹɫȡʳ⏏ອᅒɴɻᮒອ߃ອᅒɹʓȴɴϗ࡚ဌྰᅒʢࡔܧɐʳ⏏ೣ༑ᨆɹໜा০ɻ Bc ݀ɫȡʳ⏏1999 ॷ 7 21 ⏋7 26-27 ⏃ᮒ࡚⏆ɴ॑ݭჼݭᮃᠭീ⏋1999 ॷ 12 11-12 ⏃ϗ࡚⏆ɴ॑ݭᠭീˁɤɛ⏏ʚɛ⏋˰Ͳ˱˪ᦏ᧿ɹࢸाɫ 7 26 ⏃ᮒ࡚⏆⏋ჼ ݭᠭീˁɤɛ⏏ɇʰɴ⏋2001 ॷ 3 13-14 ⏃ϗ࡚⏆⏋6 26-29 Ȳʭʃ 7 1-2 ⏃ᮒ࡚⏆ ɴ͞˪˹ͻ⏋̠˱˗⏋˹ˏͲ̫Ͳ⏋͡Ͳ̜͌ा⏋̫̒˪ाɫჼݭᮃᠭീɌɛ⏏2002 ॷ 8 12 ɴɻ̠˱˗ɹᬚɹຂᓔɴᓬʱᠭീˁ⏋8 12-13 ⏃ᮒ࡚⏆ɴɻ̝Ͳ̝ ˪͇˓ɭˤ̝ͻाɫʍɹᓬʱᠭീ⏋ჼݭᠭീȲʭʃᮃᠭീˁɤɛ⏏8 13 ɭ 14 ɴɻ˰Ͳ˱˪ɫ॑ݭᠭീˁɤɛ⏏2001 ॷȲʭʃ 2002 ॷɴᠭീɌɛܩဣɻ˰Ͳ˱˪ ɹᦏ᧿ɫȡʳɛʠ⏋ᝠ 1 ɴɻ˰Ͳ˱˪ɭɌɩɗʶʰˁɭʱʚɭʠɛ⏏ (3) ˵̠͌˵˓ ǵ̨́ʭʱܕ᧑ 13 شᐻɫᏝ 5 km ആɴɤɛܩဣɴȡʳɫ⏋ᮒອ߃ອᅒɫϗ࡚ဌྰɻʼ ʶɩȦɲȦ⏏ʚɛ⏋ᦏ᭰ɴੈ।ᇕອݖɻɲȦ⏏̊ͻ̩ˠाɴ༑ԂɐʳΥఌ༑ɹΫ༑ᨆɴеᒞ ɐʳ⏏2000 ॷ 7 18-19⏋26⏋28 ⏃ᮒ࡚⏆⏋2001 ॷ 6 25 ⏃ᮒ࡚⏆⏋ε᭙ܩɹອᅒ⏋Ȳʭʃ ࢸाɫჼݭᮃᠭീˁɤɛ⏏2002 ॷ 8 15-16 ⏃ᮒ࡚⏆ɴʍɹᓬʱᠭീ⏋ჼݭᠭ ീˁɤɛ⏏ 1 ȁऔ౷ത
(4) ̨Ͳˡ̼͌ͻ̜ ǵ˵̠͌˵˓ˁ༑ʶʳ̊ͻ̩ˠाɹΥఌ༑ɹά༑ᨆɴȡɛʱ⏋˵̠͌˵˓ȴʰᏝ 1 km ɹ ܩဣɴеᒞɐʳ⏏ໜ᧑ᨆɴɇʶɛΥಁᇕອݖɫȡʳອɛʚʱɫ 2001 ॷ 3 16 ⏃ϗ࡚⏆ɴ ᮃᠭീˁɤɛ⏏ (5) ̊ͻ̩ˠा ǵܕ᧑ 13 شᐻɭɄɹाȵϯेɐʳܩဣɻϗ࡚ɫʢອɹ༲ʶɲȦੈχᇕອݖɫȡʳ⏏2001 ॷ 3 17 ⏃ϗ࡚⏆ɴᮃᠭീˁɤɛ⏏ᠭീಁɻቆঈɹཌɭቆኊঈɹဈȵࡔܧɌ⏋ອཉȵ༓ ȼ⏋ອᯆȵጓɦ Bb ݀ɹໜा০ˁɌɩȦɛ⏏ (6) ̊͌͠ͻ ǵ̍͑ͲͅͲाɹغ༑ဣȴʰᏝ 50 km Ϋ༑ɹ̫Ͳ̯˛Ͳाೣ༑ɹृࣳɴеᒞɐʳ⏏̫Ͳ̯˛Ͳ ाೣ༑ɹໜा০ɻဈɹᨆՕɹອཉȵཉȼ⏋ອᯆȵጓɛɲȦ Bc ݀ɫȡɤɛ⏏ྫ֝ɌɩȦʳ ܩݖзɴ 1999 ॷ 7 22 ⏃ᮒ࡚⏆ɴჼݭᠭീˁɤɛ⏏ (7) ̜̩̋̒˓́ ǵ̫Ͳ̯˛Ͳाೣ༑ɭܕ᧑ 23 شᐻȵϯेɐʳܩဣɴȡʳ⏏ໜा০ɻ Bc ݀ɞȵ࣬ဈ⏃˰ͻ Ͳ⏆ȵȡʳ⏏1999 ॷ 7 23-24 ⏃ᮒ࡚⏆ɴჼݭᮃᠭീˁɤɛ⏏ (8) ̅Ͳ̸͠ᦏ᧿ ǵᎢˁ୫ɤɩᤉΫˁȦɩȦʳܩݖзɴ⏋1999 ॷ 7 27 ⏃ᮒ࡚⏆ɴჼݭᠭീˁɤɛ⏏ (9) ˵ʹˎ̠˭̜̒ ǵ˵ʹˎ̠˭̜̒ላɹμਕܩ⏏̫Ͳ̯˛ͲाອᏙȴʰɇʶʳອᅋᣂ྇ɹʓȴɴ͍˱Ͳໜೣ ༑ʦ̊˓ȴʰɹᥥԂອᅋ႒ʢ༑ᦲɌɩȦʳ⏏1999 ॷ 7 28-29 ⏃ᮒ࡚⏆⏋1999 ॷ 12 19 ⏃ϗ࡚⏆⏋2000 ॷ 7 17 ⏃ᮒ࡚⏆⏋2001 ॷ 3 15 ⏃ϗ࡚⏆ɭ 7 3 ⏃ᮒ࡚⏆⏋2002 ॷ 8 17 ⏃ᮒ࡚⏆ɴ॑ݭᠭീˁɤɛ⏏ (10) ͞ˠ̫ˠ ǵ̜̠̾͌ɭɹܕޅɴеᒞɐʳᅘ⏏̫Ͳ̯˛ͲाອᏙȴʰɇʶʳອᅋᣂ྇ɹʓȴɴ̜̠̾ ͌ȴʰɹᥥԂອᅋ႒ʢ༑ᦲɌɩȦʳ⏏1999 ॷ 7 30 ⏃ᮒ࡚⏆ɴ॑ݭᠭീˁɤɛ⏏ (11) ̫Ͳ̝Ͳ ǵ͞ˠ̫ˠȴʰᏝ 20 km ά༑ɹ̫Ͳ̯˛Ͳाೣ༑ɹृࣳɴеᒞɐʳ⏏1999 ॷ 7 30 ⏃ᮒ࡚⏆⏋ ॑ݭᠭീ⏋Ȳʭʃྫკ႒ˁ୫ɤɩᤉΫˁȦɩȦʳܩݖзɴჼݭᠭീˁɤɛ⏏ (12) ̶ˎ͞Ͳ⏃̩͞Ͳ˵˓⏆ ǵܕ᧑ 9 شᐻɭ̫Ͳ̯˛Ͳाఌ༑ɹȩɠɹΥໜा⏋˵Ͳ̅˓ाɹϯेܩဣɴȡʳ⏏ 1999 ॷ 7 31 ⏃ᮒ࡚⏆ɴ࣬ဈɹ༑ᨆɫହᐨɴʭɤɩᰖˁɌɩȦʳܩݖзɴჼݭ ᠭീˁɤɛ⏏
(13) ̨́ ǵ̫Ͳ̯˛ͲाອᏙȴʰɇʶʳອᅋᣂ྇ɹʓȴɴ̊˓ȴʰɹᥥԂອᅋ႒ʢ༑ᦲɌɩȦʳ⏏ 1999 ॷ 7 31 ⏃ᮒ࡚⏆⏋Ȳʭʃ 2000 ॷ 7 26 ⏃ᮒ࡚⏆ɴ॑ݭᠭീˁɤɛ⏏ (14) ̶˝Ͳ̠͌ ǵ̍͑ͲͅͲाΫ༑ݖɹε᭙ᨆɴеᒞɐʳ⏏ɄɹɴԂʳଝմɹε᭙ܩՕອݖɹᤉΫɴȡʳອ ɛʚʱ⏋Ȳʭʃɗɹดɴȡʳࢸྂܩɫ 2000 ॷ 7 21 ⏃ᮒ࡚⏆ɴᮃᠭീˁɤɛ⏏ (15) ͅͻ˵˓ ǵ̶˝Ͳ̠͌ɭؾɴ̍͑ͲͅͲाΫ༑ݖɹε᭙ᨆɴеᒞɐʳε᭙ܩɹᢆ༅ɴȡʳອᅒɫ 2000 ॷ 7 22-23 ⏃ᮒ࡚⏆ɴᮃᠭീˁɤɛ⏏ (16) ̯Ͳ˹ˠ˗ ǵͅͻ˵˓ȴʰីɴᏝ 2 km ɹܩဣɴȡʳɫɄɹܩݖˁБᝠɐʳ॑ȵᬖȴʶʳ⏏1999 ॷ 11 15 ⏃ϗ࡚⏆ɴ॑ݭᠭീˁɤɛ⏏ (17) ̨Ͳ˱͌ ǵ˵ʹˎ̠˭̜̒⏋̨́ᬚɴȡʳɫੈ।ᇕອݖɭɌɩɹ߂ȷȦȵȡʳ⏏Ʉɹɫ 2000 ॷ 7 29-30 ⏃ᮒ࡚⏆ɴᮃᠭീˁɤɛ⏏
3ȅࠫȁض
3.1ȁऔ౷֖͈ݿ႒ ǵᠭീೝᬚμ⏋غɫ 10 ᇿ⏋31 ዀ⏋158 ዛɹᰖȵᮃʢɌȼɻᇿែተ᠗ɇʶɛ⏃ᝠ 1⏆⏏ ǵᲓᴏΫᇿȵ 74.1⎾⏃117 ዛ⏆ɭԃкɹᏝ 3/4 ˁװʠʳΥ⏋˽̀˥ᇿɻ 12.7⎾⏃20 ዛ⏆ɭ ࢹɲȦ⏏Ʉʶɻ⏋ᲓᴏΫᇿɴيʚʶʳ˱˓ዀȵ 72 ዛɫʢዛᰖధȵȼ⏋ԃкɹ 45.6⎾ˁ װʠ⏋ᐚȦɩ̝˻͚˗ዀ 16 ዛ⏃10.1⎾⏆⏋˧˧ዀ 9 ዛ⏃5.7⎾⏆ȵΫе 3 еɴеᒞɐʳɭȦ ȩჿᅓɴʭʳ⏏ʚɛ⏋Ʉɹȩɠ˱˓ዀɹ Cirrhinus cirrhosus⏋˱˓ Cyprinus carpio⏋̩˪ͤͲ Hypophthalmichthys molitrix⏋ˡͩ˽͍̀ዀɹ̠˓̘͡ˑ̴͞ˏ Oreochromis niloticus ɹ 4 ዛᰖɻ ຂɹɛʠɴውԂɇʶɛܕውԂዛɫȡʳ⏏ɇʰɴ⏋Pangasiidae ɹ Pangasius bocourti⏋̯̠ͤ ͇̀ዀɹ Clarias batrachus ɻ͞ˠ˽ᨆɹܧഀዛɞȵ̊˓ɫຂɇʶɛʢɹȵڧڃɭɌɩ॑ݭ ɫެɇʶɩȦʳ⏏ 3.2ȁऔ౷֖̤̫ͥͅݿ႒͈௳ાਫ਼͈ߊ ǵᠭീܩ 17 Ȳʭʃɗɹٞᦇɴࡔܧɐʳອݖˁ⏋ϗ࡚ɹອɹံ⏋ອᅠɹៀฐ⏋ঈᣒ⏋ອཉ⏋ ᮒ࡚ɴȲȿʳЈɹອᅠɭɹᦹᐚਸፈˁᓒૅɌ⏋Гάɹʭȩɴᰖɹᅊਖ਼ݭଘɭɌɩךՕɌɛ⏏ A⏏Υಁᇕອݖ A-1⏏Υಁᇕອݖນ 1 ȁΨϋΪͺϋକࠏ͈औ౷֖́ږ̯̹ͦݿ႒͈ςΑΠ ᠭǵീǵܩǵဣ 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 ǧOsteoglossiformesǵˏ̠ͦͩᇿ ǵǣ Notopteridae ̠˧̠̠͇̊̀ዀ 1 Chitala ornata D RD 2 Notopterus notopterus D RD ǧClupeiformesǵ̢˹Ͳᇿ ǵǣ Clupeidaeǵ̢˹Ͳዀ 3 Clupeichthys aesarnensis R R D ǵǣ Sundasalangidae ˽Ͳ̋˵͞Ͳ˪˽ዀ 4 Sundasalanx mekongensis R ǧCypriniformesǵ˱˓ᇿ ǵǣ Cyprinidaeǵ˱˓ዀ 5 Aaptosyax grypus R 6 Albulichthys albuloides R 7 Barbichthys laevis R 8 Barbonymus altus R R R 9 Barbonymus gonionotus R R RD RD R 10 Boraras micros R 11 Chela sp. R 12 Cirrhinus cirrhosus R R 13 Cirrhinus ornatipinnis R RD 14 Cirrhinus molitorella R 15 Cirrhinus sp. R 16 Crossocheilus reticulatus RD R R 17 Cyclocheilichthys apogon R 18 Cyclocheilichthys armatus R R 19 Cyclocheilichthys armatus? D D R 20 Cyclocheilichthys enoplos R 21 Cyclocheilichthys furcatus? D 22 Cyclocheilichthys repasson D 23 Cyprinus carpio RD RD 24 Esomus metallicus R R D R R R D 25 Esomus sp. R 26 Garra fasciacauda R R 27 Hampala dispar RD RD R D 28 Henicorhynchus siamensis R RD R 29 Hypophthalmichthys molitrix R 30 Hypsibarbus lagleri R 31 Hypsibarbus malcomi R D 32 Hypsibarbus sp. RD R 33 Labeo pierrei R 34 Labeo barbatula R 35 Labeo chrysophekadion R D 36 Labiobarbus leptocheila R R D R R 37 Lobocheilos melanotaenia R R R 38 Lobocheilos sp. R 39 Macrocirichthys macrochirus R R ɭ D ɻɗʶəʶᮒ࡚⏋ϗ࡚ɴՏჼɌɛɄɭˁᝠɐ⏏ᠭീܩဣɹᅱشɻГάɹݭଘˁኍɐ⏏1⏏̊͌͠ͻ⏩2⏏ ̜̩̋̒˓́⏩3⏏̩̠̜͌͞Ͳ⏩4⏏̅Ͳ̸͠ᦏ᧿⏩5⏏˰Ͳ˱˪⏩6⏏˵ʹˎ̠˭̜̒⏩7⏏͞ˠ̫ˠ⏩ 8⏏̫Ͳ̝Ͳ⏩9⏏̶ˎ͞Ͳ⏩10⏏̨́⏩11⏏˵̠͌˵˓⏩12⏏̯Ͳ˹ˠ˗⏩13⏏̶˝Ͳ̠͌⏩14⏏ ͅͻ˵˓⏩15⏏̨Ͳ˱͌⏩16⏏̨Ͳˡ̼͌ͻ̜⏩17⏏̊ͻ̩ˠा⏏
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 40 Mystacoleucus atridorsalis R D 41 Mystacoleucus marginatus R 42 Mystacoleucus sp. R 43 Opsarius koratensis R R 44 Osteochilus hasseltii RD R 45 Osteochilus lini RD R 46 Osteochilus melanopleura RD R R 47 Osteochilus microcephalus R 48 Osteochilus sp. R R 49 Oxygaster pointoni R 50 Parachela maculicaua R 51 Parachela siamensis R D 52 Parachela sp. R 53 Paralaubuca barroni R 54 Paralaubuca sp. R 55 Paralaubuca typus R R 56 Poropuntius laoensis R 57 Poropuntius sp. D 58 Probarbus jullieni R D 59 Puntioplites falcifer R 60 Puntioplites sp. R R R RD 61 Puntius aurotaeniatus R R R R D D 62 Puntius aurotaeniatus? D 63 Puntius brevis R R R 64 Puntius jacobusboehlkei R 65 Puntius orphoides R 66 Puntius sp. D 67 Raiamas guttatus R 68 Rasbora aurotaenia D R RD R R 69 Rasbora borapetensis RD R D 70 Rasbora paviei R D 71 Rasbora spilocera R R R 72 Rasbora rubrodorsalis R R R D 73 Rasbora trilineata RD R 74 Rasbora sp. R 75 Scaphognathops stejnegeri R 76 Thynnichthys thynnoides R RD R ǵǣ Balitoridaeǵ̢̨̊̓͠ዀ 77 Homaloptera sp. R 78 Nemacheilus sp. D 79 Nemacheilus sp. R ǵǣ Cobitidaeǵ̝˻͚˗ዀ 80 Acanthopsis sp. R 81 Acanthopsis sp. R 82 Acanthopsoides sp. R R 83 Botia eos R 84 Botia helodes R 85 Botia lecontei R 86 Botia modesta R 87 Botia morleti R R 88 Lepidocephalichthys hasseltii? R 89 Lepidocephalichthys sp. R R 90 Lepidocephalichthys sp. R 91 Lepidocephalichthys sp. R
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 92 Lepidocephalichthys sp. D 93 Lepidocephalichthys sp. D 94 Lepidocephalichthys sp. D 95 Pangio anguilaris R R ǧSiluriformesǵ̠͇̀ᇿ ǵǣ Bagridaeǵ˧˧ዀ 96 Bagrichthys obscurus R 97 Hemibagrus aff. nemurus R R D R R 98 Hemibagrus wyckioides R 99 Mystus albolineatus R D R 100 Mystus atrifasciatus R 101 Mystus bocourti R 102 Mystus mysticetus R D 103 Mystus sp. D R 104 Mystus sp. D ǵǣ Siluridaeǵ̠͇̀ዀ 105 Belodontichthys truncatus R 106 Hemisilurus mekongensis R RD 107 Kryptopterus apogon R R R 108 Kryptopterus sp. R R 109 Ompok bimaculatus R RD D 110 Wallago attu R D D ǵǣ Pangasiidaeǵ̮Ͳˤ˹˗˽ዀ 111 Helicophagus waandersi R R 112 Pangasius bocourti R 113 Pangasius concophilus R 114 Pangasius pleurotaenia R 115 Pangasius pleurotaenia? R 116 Pangasius polyuranodon? R ǵǣ Schilubeidaeǵ˽˥̾͡ዀ 117 Laides longibarbis R ǵǣ Clariidaeǵ̯̠͇ͤ̀ዀ 118 Clarias batrachus R R RD R D R ǵǣ Sisoridaeǵ˹̅͡ዀ 119 Bagarius sp. R 120 Glyptothorax sp. R ǵǣ Akysidaeǵˏ˥͖˹˽ዀ 121 Akysis sp. R ǧBeloniformesǵ̋̓ᇿ ǵǣ Adrianichthyidaeǵ͍̋ˡዀ 122 Oryzias pectoralis R 123 Oryzias sp. D 124 Oryzias sp. R R ǵǣ Belonidaeǵ̋̓ዀ 125 Xenentodon canciloides R R R ǵǣ Hemiramphidaeǵ˵͜͠ዀ 126 Dermogenys siamensis R ǧGasterosteiformesǵ̜˰˗ˠᇿ ǵǣ Syngnathidaeǵ͜˗˻˗ˠዀ 127 Microphis sp. D ǵǣ Indostomidaeǵ˓Ͳ̝˽̜͌˽ዀ 128 Indostomus sp. R ǧSynbranchiformesǵ̊˗̠˧ᇿ ǵǣ Chaudhuriidae ˡ˗̝˕͠ˏዀ
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 129 Chaudhuria caudata R R ǵǣ Mastacembelidae ̜˰˗̠˧ዀ 130 Macrognathus siamensis R R RD 131 Macrognathus semiocellatus R 132 Mastacembelus armatus D D 133 Mastacembelus sp. R ǵǣ Synbranchidaeǵ̊˗̠˧ዀ 134 Monopterus albus R RD R R ǧPerciformesǵ˽̀˥ᇿ ǵǣ Chandidae ̊ˡ˵˳˓˹͐̍ዀ 135 Parambasis siamensis D R R D ǵǣ Datnioididae 136 Datnioides undecimradiatus R ǵǣ Cichlidaeǵˡͩ˽͍̀ዀ 137 Oreochromis niloticus R RD R ǵǣ Nandidaeǵ̠Ͳ̝˕˽ዀ 138 Nandus oxyrhynchus R R ǵǣ Pristolepididae 139 Pristolepis fasciata RD ǵǣ Odontobutidaeǵ̝Ͳ˱ዀ 140 Neodontobutis aurarmus R ǵǣ Gobiidaeǵ̩̄ዀ 141 Brachygobius mekongensis R R D R 142 Gobiopterus sp. R D 143 Oxyeleotris marmorata RD D ǵǣ Anabantidae ˥̨̓͠˗ˠዀ 144 Anabas testudineus R RD RD R ǵǣ Osphronemidae ˠ˽̺̦ͦ͌˽ዀ 145 Betta smaragdina R R 146 Osphronemus exodon R 147 Trichogaster trichopterus R R 148 Trichopsis schalleri R R D R 149 Trichopsis vittata R R R D ǵǣ Channidae ̊˓ͩͲ̝˻͚˗ዀ 150 Channa gachua R R R 151 Channa lusius R 152 Channa maluria? R 153 Channa micropertes R D 154 Channa striata R RD R D R ǧPleuronectiformesǵˡͤ˓ᇿ ǵǣ Soleidae ˵˵˗˹̨˹̊ዀ 156 Brachirus harmandi R ǧTetraodontiformesǵ̶˫ᇿ ǵǣ Tetraodontidaeǵ̶˫ዀ 157 Monotrete suvattii R R 158 Monotrete sp. R
A-1-1⏏ອɛʚʱ ǵᮒೝɴᤉΫʦఔ႑ܩɴɫȷʳෑʠɩࢸៀฐɫʢέࡴɲΥಁᇕອݖɫȡʳ⏏ A-1-2⏏Პеອᅒ ǵϗ࡚ɴອɻɲȦȵ⏋ᮒ࡚ɹᓚсೝɴɻΥಁᇕອݖɭɲʳ⏏Њᦏɹੈχᇕອݖɭɻ᧙ᤂᮐɴ ȡʳ⏏ A-1-3⏏жеອᅒ ǵϗ࡚ɴອɻɲȦȵ⏋ᮒ࡚ɹᓚсೝɴɻΥಁᇕອݖɭɲʳဣɴȲȦɩᲞеᅒɭؾɞȵ⏋ᮒ ࡚ɻٞᦇɹໜᤄྲྀɭᏳ༑ˁЂɌɩᦹᐚɐʳ⏏ A-1-4⏏ဌྰອᅒ ǵϗ࡚ɴɻೣഀॵΫȵɤɩȦɛжеᅒȵளອͅͲ̺ˁьᅎɌɩԗʃླྀອɐʳ⏏ளອɹᬠѥ Ϋ⏋ੈχᇕອݖɫȡʳ͍˱Ͳໜఌ༑ʦɹٞᦇɴеᒞɐʳ⏏ A-1-5⏏ະອ໗ക ǵ͍˱Ͳໜೣ༑⏋ఌ༑⏋ໜᤄྲྀ⏋ঃȦ⏋ɲɰɹࣳʦɗʶɴᦏȦݭଘɴȡʱ⏋ᮒ࡚ɴɻອ ໗ɐʳΥಁᇕອݖɫȡʳ⏏ A-2⏏Υಁᇕ༑ອݖ A-2-1⏏ΥಁᇕᏳ༑ ǵϗ࡚ɴອɻɲȦΥಁᇕອݖɫ⏋ᮒ࡚ɴɲʳɭᅒʦ⏋ໜᤄྲྀʦఌ༑ɲɰˁᦹᐚɇɓʳลᔨ ˁɐʳ⏏ݭଘɴʭʱᮒೝɴྂᅊඨ႒ʦୈອඨ႒ȵᑢᗛɐʳ⏏ঈᣒɻቆɫ⏋ៀฐɻࢸɇȦȵ ໜा০ɻဈɹᨆՕɹອཉȵཉȼ⏋ອᯆȵጓɛɲȦ Bc ݀ɫȡʳ⏏ B⏏ੈχᇕອݖ B-1⏏ੈχᇕອݖ B-1-1⏏ໜᤄྲྀ ǵ͍˱Ͳໜೣ༑ʦఌ༑ɴᦏஞɌɛݭଘɴȡʳອɹੈχᇕອݖɫ⏋ࣳᦇٞᦇɴɻະʦ ༇ອകˁɐʳ⏏ᮒ࡚ɴɻ͍˱Ͳໜೣ༑ᨆʦఌ༑ᨆɭອᤉˁЂɌɩᦹᐚɐʳ⏏ঈᣒɻɫໍ ອୈອ༝ᘽ༝ᧉɲʰʃɴྂᅊඨ႒ȵᑢᗛɐʳ⏏̩̠̜͌͞Ͳɹ̨Ͳ̊˓̶ˏ⏋˰Ͳ ˱˪ɹ̨ͲˡͲ⏋̨Ͳ͍̩ͻͲȵɄʶɴȡɛʳ⏏ B-1-2⏏ ǵໜᤄྲྀɴᰖаɐʳȵ⏋ອᅠɹៀฐȵ߂ȷȼ⏋ϗ࡚ɴॵΫȵɤɩȦɛອᤉȵᮒ࡚ɹອеɹΫ ɴʭʱ͍˱Ͳໜఌ༑ɭᦹᐚɐʳʢɹɹᤂᮐᇕɴɻɗʶɭ᧙ȦݭଘɴȡʳອݖˁɄɄɫɻ ɭ٧ʉɄɭɴɐʳ⏏˰Ͳ˱˪ᦏ᧿ɹ˽˓ȵɄʶɴȡɛʳ⏏ B-1-3⏏ ǵໜᤄྲྀɴᰖаɐʳȵ⏋ᮒ࡚ɫʢٞᦇɹੈχᇕອݖɭᏳ༑ɲɰɴʭʱᦹᐚɌɲȦອݖˁɄ
Ʉɫɻɭ٧ʉɄɭɴɐʳ⏏ B-2⏏ੈχᇕ༑ອݖ B-2-1⏏ੈχᇕᏳ༑ ǵϗ࡚ʢອɹ༲ʶɲȦᏳ༑ɫ⏋ໜᤄྲྀʦɭᦹᐚɐʳ⏏ໜाᄩޅɻΫ៨ɹΥಁᇕᏳ༑ɭᰖа ɐʳ⏏ B-2-2⏏͍˱Ͳໜఌ༑ ǵϗ࡚ɫʢອȵɲȼɲʰɲȦੈχᇕອݖɫȡʳȵ⏋ݭଘɴʭɤɩɻɄɹಁೝ⏋ອཉȵᘿɌ ȼ༓ȼɲʱϺȵਂɫʢाˁดుɫȷʳʭȩɴɲʳ⏏ʚɛ⏋͍˱Ͳໜೣ༑ɴປʏɩा३ȵႷ Ȧ⏏ঈᣒɻቆɫ⏋ໜा০ɻဈɹᨆՕɹອཉȵཉȼ⏋ອᯆȵጓɛɲȦ Bc ݀ɫȡʳ⏏ B-2-3⏏͍˱Ͳໜೣ༑ ǵϗ࡚⏋ᮒ࡚ˁکʼɑອȵࡔܧɌɩȦʳɭȦȩઝ٢ɫ⏋ɄɹܩݖɴȲȿʳʢɤɭʢࡴɌɛ ੈχᇕອݖɫȡʳ⏏ɛɞɌ⏋ϗ࡚ɭᮒ࡚ɹອеेɻధ m ɴؖʉ⏏ঈᣒɻቆɫ⏋ໜा০ ɻဈɹᨆՕɹອཉȵཉȼ⏋ອᯆȵጓɛɲȦ Bc ݀ɫȡʳ⏏ B-2-4⏏࣬ဈ ǵ͍˱Ͳໜೣ༑ʦఌ༑ɻቆɹঈᣒȵযǾɭᐚȼ⏏ɌȴɌ⏋ჼܩؿɫ˰ͻͲɭ٧ɽʶʳݭଘ ɻອঈȴʰ࣬ኂȵ᭠ᣱɌ⏋ाɻອɌʉȷˁȡɁɩ࣬আˁ༑ʶάʳਸ਼༑ᨆɭɲʳ⏏Ʉɹά༑ᨆ ɴɻʼɑȴɴኊঈȵࡔܧɐʳɄɭʢȡʳ⏏ B-2-5⏏ཌ ǵ͍˱Ͳໜೣ༑ʦఌ༑ɫ⏋႕ɴອཉɹȡʳཌˁჼܩؿɫͩͲɭ٧ʉ⏏ 3.3ȁݿ႒͈ڰঃ͂௳ાਫ਼၌ဥ ǵᠭീܩݖɫተ᠗ɇʶɛᰖɹȩɠ⏋ɗɹᅊ༎رɭᅊਖ਼ݭଘɹգᅎɹᬠѥɴᬠɐʳݬȵȴ ʶࢹɲȴʶʰʶɛዛᰖɻᏝ 56 ዛɫȡʳ⏏ተɲዛధˁ៨ɇɲȦɹɻɹຍ᭥ɫ⏋ࣚɻؾ ࡴɫȷʳȵዛᰖʚɫɻჼܧɴȲȦɩɻέخᔨɲʢɹȵيʚʶʳȴʰɫȡʳ⏏Гάɴɗʶʰˁෙ ᠦɐʳ⏏ A⏏Υಁᇕອݖɹʛɴᅊਖ਼ɐʳᰖ ǵ˱˓ዀɹ Esomus metallicus ⏃Ԡሎ 1⏆ɻ ᮒ࡚ɴжеອᅒɻʢɠʸ˃ɹɄɭ⏋Პеອᅒɫʢ ҅кధɻෑʠɩȼ⏋᧑ᤉΫɹອɛʚʱɴʢՏჼɐʳ⏏ʚɛ⏋ᮒ࡚ɴᩄɹᮒȵᭇʱ⏋Ȧɛ ʳɭɄʸȵອ༡ɌɴɲʳʭȩɲႥɹಁ⏋ೣዛȵ᧑ᤉΫɫʢ⏋ອཉɹෑʠɩ༓ȦᇤᕖႥɹອ ɹ༑ʶɹμˁృɌȦΥಁᇕອݖˁິʠɩᇵ˃ɴ᧚ΫɌɩȦȼࠒȵ᠗ʠʰʶʳ⏏ອᅒԒʦɗʶ ɴ᭰ஞɐʳఔ႑ܩɹອɛʚʱፈȴʰ⏋ʈሁɹմೝІȴʰଁʚɫعଁᬏຍ᭥ȵؾಁɴ ᮃɇʶʳ⏏ɄɹɄɭȴʰ⏋ೣዛȵɄɹݭଘɫᑢຂɌ⏋ԃᅊ༎رˁȲȼɤɩȦʳɄɭɻተࡸ
ɫȡʳ⏏ϗ࡚ɻອᅒʦᏳ༑ԒɹॵΫȵʱȴȿɛອɛʚʱɴ᠗ʠʰʶʳʢɹɹ⏋ੈχᇕອݖɫ ೣዛɹᅊਖ਼ȵተ᠗ɇʶɲȦ⏏ܩݖзɹ᠌ɫɻ⏋ᮒ࡚ɹɻɍʠɴੈχᇕອݖɭɻʚɞᦹᐚਸ ȵɲȦອᅒɴອȵྊʚʱɻɍʠʳɭɐȽɴࢸ݀҅кȵʛʰʶʳɭȦȩ⏏ɄɹɄɭɻᘿᓔʰʢ ተ᠗Ɍɛ⏏Јɹອݖȴʰው֪ɌɩɄɲȦɹɴɄɹʭȩɲჼᢒȵȲȷʳɹɻ⏋ೣዛɹȵϗၥ ᓘχਸˁ୫ɠ⏋Ʉʶȵϗ࡚ɹᬚɻອᅒɹܤμɴȡʱ⏋ອȵྊʚʳɭؾಁɴɗʶȵʈɐʳɭ ȦȩخᔨਸȵᲞȦ⏏ ǵ˱˓ዀɹ Rasbora rubrodorsalis ʢᮒ࡚⏋ϗ࡚ˁɭȲɌɩੈχᇕອݖɴՏჼɓɑ⏋ϗ࡚ʢອ ɛʚʱɹʭȩɲέࡴɲݭଘɫᣴϗɐʳဣɫմᓔɭʼʰɲȦȵ⏋ᲞеອᅒԒɫɻ҅кధȵ ࢹɲȦ⏏
ǵ̝˻͚˗ዀɹ Lepidocephalichthys spp.⏃Ԡሎ 2⏆ʢ Esomus metallicus ɭؾɲᅊ༎رˁȲȼ ʳʢɹɭਲ਼ʼʶʳȵ⏋ೣዛɹݭغ⏋Esomus metallicus ɭᅵɲʱ⏋ଁɻྂȵȡʶɽᬏಁᬚ ᅊࡔخᔨɲɹɫ⏋ϗ࡚ɴɻອᅒɹঈཉȼ࿃ԂɌɩᣴϗɐʳخᔨਸʢوࡴɫȷɲȦ⏏ ǵΫ៨ 3 ዛɻϗ࡚ɹဌྰᅒɫʢተ᠗ɇʶ⏋႕ɴմ 2 ዛɻʦᗐʢᮃɇʶɛ⏏ B⏏ϗ࡚ɴɻੈχᇕອݖɴᅊਖ਼Ɍ⏋ᮒ࡚ɴɻΥಁᇕອݖɴʢଁȵᅊਖ਼ɐʳᰖ ǵɄɹʭȩɲᅊ༎رˁȲȼʳᰖɻ⏋գᅎɐʳᅊ༎ݭଘɹेɴʭʱГάɹ 4 ̊˓̺ɴךՕɇ ʶʳ⏏ B-1⏏ΥಁᇕᏳ༑⏋жеອᅒ⏋Პеອᅒɫᑢຂɐʳᰖ
ǵ̯̠͇ͤ̀ዀɹ Clarias batrachus ⏃Ԡሎ 3⏆⏋̜˰˗̠˧ዀɹ Macrognathus siamensis ⏃Ԡሎ 4⏆⏋˥̨̓͠˗ˠዀɹAnabas testudineus ⏃Ԡሎ 5⏆⏋̊˓ͩͲ̝˻͚˗ዀɹ Channa striata ⏃Ԡ ሎ 6⏆ȵɄʶɴȡɛʳ⏏Channa striata ɻᅤɹᗳȵອᯆɴុȦɇʳʭȩɲݭଘ⏋ʚɛྂਸ⏋ ୈອඨ႒ȵᑢᗛɐʳݭଘɴ༝ਸˁᅋ⏋៊ɻʈʢɄʶʰˁࡪʳ⏏ܩݖзɹݬɴ ʭʶɽ⏋Clarias batrachus ɻʦʦອཉɹȡʳࣳᦇɹອঈɴ៊ȵܤˁɲʰɌɩ͇˗Ͳ̝Ⴅɹ ुˁᦶʱ⏋ˁѳᡸɐʳɭȦȩ⏏
ǵMacrognathus siamensis ˁ᭑ȼʓȴɹ 3 ዛɻជ֖ዷຩ٧ُ۬ࡲˁɌ⏋ϗ࡚ɫʢྂȵӱ ՕɲݭଘɫȡʶɽܤμɫᣴϗɌɩȦʳخᔨਸȵȡʳ⏏ʚɛ⏋Macrognathus siamensis ʢкȵ ྂɤɩȦʶɽᬏಁᬚᅊࡔخᔨɲɹɫ⏋մ 3 ዛɭؾɹخᔨਸȵᓒȭʰʶʳ⏏
ǵɲȲ⏋Channa striata ɻϗ࡚ɹဌྰᅒɴȲȦɩɭॾȵᮃɇʶɛ⏏ B-2⏏ΥಁᇕᏳ༑⏋жеອᅒɫᑢຂɐʳᰖ
ǵ˱˓ዀɹ Puntius aurotaeniatus ⏃Ԡሎ 7⏆⏋̊˗̠˧ዀɹ̊˗̠˧ Monopterus albus ⏃Ԡሎ 8⏆ ȵɄʶɴȡɛʳ⏏
ǵ̊˗̠˧ɻዷຩμȴʰᨧᏬˁ٧ُɐʳɄɭȵɫȷ⏋кȵྂɤɩȦʶɽᬏಁᬚᅊࡔخᔨɲɹ ɫ⏋ྂɹȡʳঈᣒȵȡʶɽϗ࡚ɫʢжеອᅒʦΥಁᇕᏳ༑ɫᣴϗɌɩȦʳخᔨਸȵᲞȦ⏏ B-3⏏ΥಁᇕᏳ༑⏋ໜᤄྲྀᑉᦇᨆɹະɫᑢຂɐʳᰖ
ǵ˱˓ዀɹ Puntius brevis ⏃Ԡሎ 9⏆⏋˽̀˥ዀɹ Parambasis siamensis ⏃Ԡሎ 10⏆ȵɄʶɴȡɛ ʳ⏏ᮒ࡚ɫʢອᅒɴѢԂɐʳɄɭɻॕɫȡʳ⏏
B-4⏏жеອᅒ⏋ໜᤄྲྀᑉᦇᨆɹະɫᑢຂɐʳᰖ
ǵ༑ອݖˁ߫ʚɲȦዛᰖɫ⏋ໍອඨ႒ፈɹອᗳȵᑢᗛɌɛᄩޅɴȼՏჼɐʳȵ⏋ᮒ࡚ɴɻ ɗʶʰɹɲȦжеອᅒʦໜᤄྲྀᑉᦇᨆɹະɴʢᅊਖ਼ɐʳ⏏ʚɛ⏋ʓȴɭ᧓Ȧ⏋ϗ࡚ɫʢ ᑢຂɐʳ⏏
ৢ૯ 3ȁClarias batrachus ͈౽ݿ ৢ૯ 4ȁMacrognathus siamensis ͈౽ݿ
ǵ͍̋ˡዀɹ͍̋ˡࣚ ⏃Oryzias pectoralis ɲɰ⏆ɴيʚʶʳዛᰖ⏃Ԡሎ 11⏆Ȳʭʃˠ˽̺ͦ ̦˽͌ዀɹ Trichogaster trichopterus, Trichopsis schalleri ⏃Ԡሎ 12⏆, Trichopsis vittata ȵɄʶɴ ȡɛʳ⏏
ǵɲȲ⏋Trichogaster trichopterus ɭ Trichopsis schalleri ɻϗ࡚ɹဌྰᅒɫʢᮃɇʶ⏋႕ɴմ ᓔɻᅋআʦʢተ᠗ɇʶɛ⏏
C⏏ଁɻੈχᇕອݖɴᅊਖ਼Ɍ⏋ᮒ࡚ɴΥಁᇕອݖˁᅋݭɭɐʳᰖ
ǵɄɹʭȩɲᅊ༎رˁȲȼʳᰖɻ⏋գᅎɐʳᅊ༎ݭଘɹेɴʭʱГάɹ 2 ̊˓̺ɴךՕɇ
ৢ૯ 7ȁPuntius aurotaeniatus ͈౽ݿ ৢ૯ 8ȁΗ;Ά Monopterus albus ͈౽ݿ
ৢ૯ 9ȁPuntius brevis ͈౽ݿ ৢ૯ 10ȁParambasis siamensis ଼͈ݿ
ʶʳ⏏ C-1⏏ଁɻੈχᇕ༑ອݖɴᅊਖ਼ɐʳᰖ ǵ̠͇̀ዀɹ Wallago attu ⏃Ԡሎ 13⏆ȵɄʶɴȡɛʳ⏏ೣዛɹଁɻ͍˱Ͳໜೣ༑ʦఌ༑ɴ ᅊਖ਼ɌɩȦʳ⏏ᮒ࡚ɹɻɍʠ⏋߂ᩄɹᭇᮒɴʭʱອеɹਸ਼Ϋɫ⏋ϗ࡚ɴɻॵΫȵɤɩȦɛ Ᏻ༑ȵླྀອɌɩໜᤄྲྀɭ͍˱Ͳໜɹ༑ອᨆȵᦹᐚɐʳ⏏ೣዛɻɄɹಁˁʛɻȴʰȦ߂ᒴˁɲ ɌɩໜᤄྲྀɴѢԂɐʳ⏏ܩݖзɹݬɴʭʶɽ⏋ະʦອ໗കɫᅋɐʳɭȦȩ⏏ɗɹ ⏋ໜᤄྲྀˁᔒଁݭଘɭɌɩգᅎɌ⏋ଁᬏɭɭʢɴ͍˱Ͳໜೣ༑ɴʢɰʳ⏏ C-2⏏ଁɻੈχᇕອݖɴᅊਖ਼ɐʳᰖ
ǵ̠͇̀ዀɹ Ompok bimaculatus ⏃Ԡሎ 14⏆⏋̊˓ͩͲ̝˻͚˗ዀɹ Channa lusius ⏃Ԡሎ 15⏆⏋ Ch. micropertes ȵɄʶɴȡɛʳ⏏
ǵOmpok bimaculatus ɹଁɻʦໜᤄྲྀɴᅊਖ਼ɌɩȦʳ⏏ᮒ࡚ɴໜᤄྲྀٞᦇɹະɫ ȵᮃɇʶʳɄɭȴʰ⏋ɄɹݭଘˁᅋݭଘɭɌɩգᅎɌɩȦʳɴ᧓ȦɲȦ⏏ɇʰɴᗐ ɻжеອᅒʦੈχᇕΥಁᇕᏳ༑ɴʢѢԂɌɩଁᬏɐʳ⏏
ǵChanna lusius ɭ Ch. micropertes ɻঃȦᬖఔᇕɲອᯆˁ୫ɦʦໜᤄྲྀፈɹອݖɴȦ⏏ ະʦ༇ອകɫȵᮃɇʶʳɄɭȴʰɄʶʰɹݭଘˁᅋݭଘɭɌɩգᅎɌɩȦʳɴ ᧓ȦɲȦ⏏
ৢ૯ 13ȁWallago attu ͈ဘݿ
D⏏ଁɻੈχᇕອݖɴᅊਖ਼Ɍ⏋ᮒ࡚ɴΥಁᇕອݖˁᔒଁݭଘɭɐʳᰖ
ǵɄɹʭȩɲᅊ༎رˁȲȼʳᰖɻ⏋գᅎɐʳᅊ༎ݭଘɹेɴʭʱГάɹ 2 ̊˓̺ɴךՕɇ ʶʳ⏏
D-1⏏ଁɻੈχᇕ༑ອݖɴᅊਖ਼ɐʳᰖ
ǵ˱˓ዀɹ Hampala dispar ⏃Ԡሎ 16⏆⏋Henicorhynchus siamensis⏋˧˧ዀɹ Mystus mysticetus ȵ Ʉʶɴȡɛʳ⏏
ǵHampala dispar ɹଁɻ͍˱Ͳໜೣ༑ఌ༑ɴȼᅊਖ਼ɌɩȦʳȵʼɑȴɲȵʰៀฐɹ߂ ȷɲɴʢՏჼɐʳ⏏ɻ͍˱Ͳໜఌ༑ʦɴɦʰɲʳੈχᇕఌ༑ɫᮃɇʶʳɹɫɄʶ ʰɹݭଘȵᅋݭɫȡʳخᔨਸȵᲞȦ⏏ɗʶʭʱ߂݀ɹॾɴɲʳɭໜᤄྲྀʦɗʶɴɦʰɲ ʳжеອᅒɴѢԂɌɩᅊᔒɌ⏋ଁᬏɴЫɤɩଁɹᅊਖ਼ݭଘɴʢɰʳ⏏
ǵHenicorhynchus siamensis ɭ Mystus mysticetus ɹଁʢΫዛɭؾɲݭଘɴᅊਖ਼ɌɩȦʳ⏏ ᅋݭଘɻέ౩ɫȡʳȵ⏋ॾɻໜᤄྲྀʦɗʶɴɦʰɲʳжеອᅒɴѢԂɌɩɗʶʰɹݭଘ ˁᔒଁݭଘɭɌɩգᅎɌɩȦʳ⏏
D-2⏏ଁɻੈχᇕອݖɴᅊਖ਼ɐʳᰖ
ǵ̢˹Ͳዀɹ Clupeichthys aesarnensis⏋˱˓ዀɹ Rasbora aurotaenia⏋̋̓ዀɹ Xenentodon canciloides ȵɄʶɴȡɛʳ⏏
ǵClupeichthys aesarnensis ɻঃȦᬖఔᇕɲອᯆˁ୫ɦʦໜᤄྲྀፈɹອݖɴȦ⏏ೣዛɹ ˹͞˽݀Іɻᮒ࡚ɴະʦອ໗കˁᔒଁݭଘɭɌɩգᅎɌɩȦʳ⏏
ǵRasbora aurotaenia ɭ Xenentodon canciloides ɻໜᤄྲྀ⏋ɲɰɴᅊਖ਼Ɍ⏋Ʉʶʰɹݭଘɫᑢ ຂɐʳ⏏ᗐɻᮒೝɴΥಁᇕᏳ༑ɲɰɴѢԂɌɩଁᬏɐʳ⏏
E⏏ଁɻੈχᇕ༑ອݖɴᅊਖ਼Ɍ⏋ᮒ࡚ɴੈχᇕອݖˁІɹᔒଁݭଘɭɌɩգᅎɐ ʳᰖ
ǵ˱˓ዀɹ Barbonymus altus ⏃Ԡሎ 17⏆⏋Crossocheilus reticulatus ⏃Ԡሎ 18⏆⏋Cyclocheilichthys
ɹЗᬚ⏋Labiobarbus leptocheila, Lobocheilos melanotaenia, Mystacoleucus atridorsalis, Paralaubuca typus⏋̝˻͚˗ዀɹ Acanthopsoides sp., ˧˧ዀɹ Hemibagrus aff. nemurus, ̮Ͳˤ˹˗˽ዀɹ Pangasius pleurotaenia ȵɄʶɴȡɛʳ⏏ ǵΫ៨Ɍɛ˱˓ዀɹዛᰖɻІȵໜᤄྲྀɲɰɹອݖɫᮃɇʶʳɄɭȴʰ⏋ɄɄˁᔒ ଁݭଘɭɌɩգᅎɌɩȦʳɄɭɻተࡸɫȡʳ⏏ɌȴɌ⏋ᅋʢɄɹݭଘɫʼʶɩȦʳȴɰ ȩȴɻჼܧɹɭɄʸέ౩ɫȡʳ⏏Ʉʶʰɹዛᰖɹȵɰɹʭȩɲਸᣒȴʢៗ౩ɫȷɩȦɲȦ ȵ⏋ʢɌʢ༝ਸɫȡʶɽ⏋ވອಁɴອɭɭʢɴȵໜᤄྲྀɴԂʱ⏋ʈɌɛІȵɄɄɫ ᇍᔒɌɩȦʳخᔨਸʢȡʳȴʰɫȡʳ⏏
ǵHemibagrus aff. nemurus ɹଁɻςɴ͍˱Ͳໜೣ༑⏋ఌ༑ɴᅊਖ਼ɐʳ⏏ܩݖзɹ᠌ɫɻ⏋ ఌ༑ɹཌȵᅋݭଘɫȡʳɭȦȩ⏏ॾɻఌ༑ɹʓȴ⏋ໜᤄྲྀɲɰɴѢԂɌɩଁᬏɐʳ⏏ ǵAcanthopsoides sp. ɭ Pangasius pleurotaenia ɹᅋݭଘɴᬠɐʳݬɻɲȦȵ⏋⏋ॾ ȵໜᤄྲྀɫᮃɇʶʳɄɭȴʰ⏋ɄɹݭଘˁᔒଁݭଘɭɌɩգᅎɌɩȦʳʢɹɭਲ਼ʼʶʳ⏏ ǵɄʶʰɹᰖɴɻድɹᦽࣙɴʭʱᅋݭଘɹݬȵʰʶʶɽ⏋C-1 ɴךՕɇʶʳዛᰖ ʢيʚʶɩȦʳʢɹɭਲ਼ʼʶʳ⏏ F⏏ੈχᇕອݖɹʛɴᅊਖ਼ɐʳᰖ ǵɄɹʭȩɲݭଘɫᅊ༎رˁȲȼʳᰖɻ⏋գᅎɐʳᅊਖ਼ݭଘɹेɴʭʱГάɹ 2 ̊˓̺ɴ ךՕɇʶʳ⏏ F-1⏏ੈχᇕອݖɴᅊਖ਼ɐʳᰖ
ǵ˱˓ዀɹ Boraras micros ⏃Ԡሎ 19⏆⏋Rasbora boprapetensis, R. spilocera, R. trilineata⏋̝˻͚ ˗ዀɹ Pangio anguilaris⏋˵͜͠ዀɹ Dermogenys siamensis, カウドゥリア科ɹ Chaudhuria caudata, ̠Ͳ̝˕˽ዀɹNandus oxyrhynchus ⏃Ԡሎ 20⏆⏋̝Ͳ˱ዀɹ Neodontobutis aurarmus⏋ ̩̄ዀɹ Brachygobius mekongensis ȵɄʶɴȡɛʳ⏏
ǵɄʶʰɹᰖɹȩɠ⏋Rasbora boprapetensis ɻϗ࡚ɴॵΫȵʳɄɭɻɲȦʢɹɹ༓ȼɩࢸ
ៀฐɲᄩޅɴȦ⏏
F-2⏏ੈχᇕ༑ອݖɴᅊਖ਼ɐʳᰖ
ǵ ˱ ˓ ዀ ɹ Aaptosyax grypus ⏃Ԡ ሎ 21⏆⏋ Probarbus jullieni ⏃Ԡ ሎ 22⏆⏋ Raiamas guttatus, Scaphognathops stejnegeri, Hypsibarbus spp., ̠͇̀ዀɹ Kryptopterus spp., ˧˧ዀɹ Mystus albolineatus, M. atrifasciatus ȵɄʶɴȡɛʳ⏏
ǵProbarbus jullieni ɹଁɻ͍˱Ͳໜೣ༑ʦఌ༑ɴᅊਖ਼Ɍ⏋ϗ࡚ɴ࣬ဈɴᮃʚʱᅋɐʳ [Roberts and Warren 1994]⏏ࢸ݀ɹᗐɻ͍˱Ͳໜఌ༑ɫᮃɇʶʳ⏏Hypsibarbus ɴࣚɐʳዛ ᰖɻܩݖзɹ᠌ɫɻ⏋ᮒ࡚ɴ߂ᒴɫ࣬ဈɴᮃʚʱ⏋࣬ɹΫɴᅋɐʳɭȦȩ⏏ɗɹʓȴ⏋ ɄɄɴȡɁɛዛᰖɹᅋݭଘɴᬠɐʳݬɻɲȦȵ⏋ೣᠭീݖɴȲȦɩΥಁᇕອݖȴʰɹ៨ ᪼ɻɲȼ⏋ʢɌȼɻॾȵ͍˱Ͳఌ༑ᨆɫᮃɇʶʳɄɭȴʰ⏋Ʉɹݭଘˁᔒଁݭଘɭ ɌɩգᅎɌɩȦʳɄɭɻተȴɫȡʳ⏏ 3.4ȁਯྦྷ͈ݽႻڰ൲͂ݿ႒͈௳ાਫ਼၌ဥ ǵ࣬ᅒ [2002] ɻೣᠭീܩݖȴʰ 25 ዛɹྫˁݬٚɌɩȦʳȵ⏋ɗʶʰɭᰖɹᅊਖ਼ݭଘգᅎ ɭɹᬠᦹɴɦȦɩɻᦚʏʰʶɩȦɲȦ⏏ೣ፲ɫɻɗɹݬˁᏳɴ៨ᥔɐʳ⏏
ৢ૯ 19ȁBoraras micros ଼͈ݿ ৢ૯ 20ȁNandus oxyrhynchus ͈౽ݿ
(1) தȦᐨ⏃˵ʹˑͲ⏆⏃Ԡሎ 23⏆
ǵᦲ।ɻΥಁᇕອݖɹжеອᅒ⏋Პеອᅒፈɫьᅎɇʶʳ⏏ᮒ࡚ɴȲȿʳਸ਼ɲވອಁɴɻ Ϋ៨Ɍɛݭଘɹʓȴɴ⏋͍˱Ͳໜఌ༑ɹ᭝ɹࢹɲȦࣳᦇ⏋Ȳʭʃϗ࡚ɫʢʦໜᤄྲྀɫь ᅎɇʶʳɄɭʢȡʳ⏏ςɭɌɩଁϺߨਸɭӻጣȵᅎȦ⏋Rasbora, Esomus metallicus, Trichopsis, Trichogaster trichopterus, Anabas testudineus, Channa striata, Clalias batrachus ፈȵɇʶʳ⏏ (2) ؕଝᐨ⏃ˡ̜Ͳ⏆⏃Ԡሎ 24⏆ ǵ͍˱Ͳໜఌ༑⏋ፈɹੈχᇕອݖɫᅎȦʰʶʳ⏏ଁϺᅖਸȵьᅎɌ⏋ᖐɹʍɇȷɴܔࡴɌɩ ᖐˁᦽʠ⏋ᝠࣝˁᧉɌɩȦʳᰖˁɐʳɭ⏋ೠɫᦶɤɛݭɴɄɹᐨˁܔࡴɌɩ༑ άɌɩȼʳᰖˁʰȭʳ 2 ዛᰖɹȵȡʳ⏏ьᅎಁೝʦࢪᢒዛɴᬠɐʳݬɻʰ ʶɲȴɤɛ⏏ (3) ܋ଝᐨ⏃˵̝˕Ͳ⏆ ǵଁϺߨਸɴʭʱॷᬚˁɭȲɌɩьᅎɇʶʳȵ⏋႕ɴᮒ࡚⏋ᭇᮒɹވອಁɴᰖȵ༎ᇍɴ ֪ɐʳᅊˁգᅎɌɩΥಁᇕᏳ༑⏋ະ⏋ອ໗ക⏋ݩອɌɛжеອᅒɹʓȴ⏋ੈχᇕອݖɫ ȡʳ͍˱Ͳໜೣ༑⏋ఌ༑⏋ໜᤄྲྀ⏋⏋ፈ⏋ݭȵʭȼɄɹᐨɹఌ഼ᨆȵԃɴɻອ໗Ɍɲ Ȧʭȩɲ༓ଘɫᅎȦʰʶʳ⏏ᐨɴˁৄɐʳɛʠɹɻԂʶɲȦ⏏ອঈɴᐨˁᘺɭɌɩɌɽ ʰȼ৺ɠ⏋ȵԂʳغˁឿʰɤɩளɁʳ௭сˁᑵʱᦐɐ⏏ᐨɹΫˁᦲɐʳɐʏɩɹዛ ȵࢪᢒɭɲʳȵ˱˓ዀ⏋̝˻͚˗ዀ⏋̠͇̀ዀ⏋̜˰˗̠˧ዀ⏋̊˓ͩͲ̝˻͚˗ዀɲɰ ɹࢸμ݀ዛȵȼ⏋߂݀ȵɇʶʳɄɭɻॕɫȡʳ⏏ (4) ହᐨ⏃̼ͻ) ǵଁϺᅖਸȵ߫˃ɫьᅎɐʳྫԍɫ⏋ੈχᇕອݖɫȡʳ͍˱Ͳໜೣ༑⏋ఌ༑⏋᭭ࢀ႒ɹɲȦ ʦໜᤄྲྀፈ⏋͍˱Ͳໜೣ༑⏋ఌ༑ɫɻঈᣒɻکʼɲȦȵອཉɹȡʳᨆՕʭʱɻ༓Ȧ࣬ဈٞ ᦇȵ߫ʚʶʳ⏏ጓɠᦉʛɹʓȴɴᖐΫȴʰହԂɐʳʢȡʳ⏏ςɴɹᩄȵȦᮒ࡚ɴʼ ৢ૯ 23ȁ̞݉࿌Ȫόͻϋȫ ৢ૯ 24ȁए࿌ȪΠϋȫ
ʶʳ⏏ೣɫɻହᐨɹμ߈ᨆɴɄʶˁଝᑵʱࢊɓʳᏣȵȡʳȵ⏋ೣᠭീܩɫɻɄʶȵࡔܧɌɲ Ȧ⏏ହԂ⏋ହɁɛϺȵອɹμɴԂɤɩᐨˁ܌ؚɐʳɛʠᏣˁਖឹɭɌɲȦȴʰɫȡʳ⏏ɛɞ Ɍ⏋ອཉɹȡʳݭଘɫьᅎɐʳݭغɻᏣˁɦȿʳɭȦȩ⏏ᐨɴȴȴʳɻɐʏɩգᅎࢪᢒɭɇ ʶʳȵ⏋ᕶсɹɛʠᐨᇿˁࢸɇȼɐʳɹȵࢅ౫ɫɻɲȼ⏋ςɴμ߂݀ɹዛˁႰȩ⏏ (5) լᐨ⏃͐ͻͲ⏆ ǵହᐨɭؾɴଁϺᅖਸȵੈχᇕອݖɫьᅎɐʳྫԍɫ⏋ьᅎȵ 2 ɦɴ߂աɇʶʳ⏏ʚɑ⏋ ͍˱Ͳໜೣ༑⏋ఌ༑ɹ༑ອݖɹᝠࣝȴʰμࣝɫȩ༑Ɍլᐨɫȡʳ⏏ᖐȴʰᐨˁໜाԒɴହԂ Ɍ⏋ອ༑ɴʚȴɓɩΥࡴךᬚ༑άɐʳɹˁ৺ɤɩȴʰᐨˁளɁ⏋ʚɛΫ༑ᨆɴଖɤɩΥᦹɹ௭ сˁᑵʱᦐɐ⏏ᧉਸɹȦμ߂݀ɹ˱˓ዀ⏋̠͇̀ዀɲɰˁႰȩ⏏ʢȩ 1 ɦɻᐨˁЇȿ ʳеᒞȵΥࡴɴѳɛʶɩȦʳɫȡʳ⏏͍˱Ͳໜఌ༑ɹ༑ອݖɭʦɲɰɹອݖȦɑʶ ɫʢʼʶ⏋ହᐨɭᅵɲʱອᗳʦໍೠɲɰɹ᭭ࢀ႒ȵȡʳݭଘɫʢȵɫȷʳɭȦȩգဣ ȵȡʳ⏏մᦚɹɭؾɴࢪᢒዛɻᧉਸɹȦ˱˓ዀ⏋̠͇̀ዀɲɰɫȡʳȵ⏋ᒞ ݭଘȵອঈɴᦏȿʶɽঈᅊਸɹȦ̝˻͚˗ዀ⏋˧˧ዀȵ⏋ʚɛ⏋ɴᒞɌɩᓂȡɁʶ ɽᬚʦᚚ౩ᚚಬɴ༎֪ɐʳ̠͇̀ዀ⏋̊˓ͩͲ̝˻͚˗ዀȵɇʶʳ⏏ (6) ؘଝᐨ⏃̢˙Ͳ⏆⏃Ԡሎ 25⏆ ǵଁϺߨਸɹʓȴᅖࡎߨࡎӻጣȵ 2 Ϻ 1 ᏼɫьȦ⏋ଁϺᅖਸȵьȩɄɭɻॕɫȡʳ⏏௭сɴ ɻ 2 ɦɹȵȡʳ⏏ 1 ɦɻɄʶˁ୫ɤɩອμˁմᦽɐʳɫ⏋ʓȴɻጓɠʚɤɛʚʚɫ ອ༑ɭɭʢɴ༑άɐʳˁɐʳɫȡʳ⏏Ȧɑʶɹݭغʢݭɹተȴɲ༓ଘȵ᧧ɽʶ ʳ⏏մᓔɻςɴΥಁᇕᏳ༑⏋ʦٞᦇɹະ⏋ᓔɻᭇᮒɹވອɫ༎ᇍɴ༎֪ɐʳ ᰖˁࢪᢒɭɌɩੈχᇕᏳ༑⏋͍˱Ͳໜఌ༑ʦɹࣳᦇ⏋ȲʭʃɗʶʰɹݭଘɴɦʰɲʳΥಁ ᇕᏳ༑ɫʼʶʳ⏏մᓔɹɫɻ Rasbora rubrodorsalis, Esomus metallicus, Lepidocephalichthys, Anabas testudineus ፈɹࢸμ݀⏋ᓔɫɻ Rasbora aurotaenia, Xenentodon canciloides ፈɹʓ ȴɴވອಁɴϟఖᇕɴ༑άɌɩȷɛ Esomus metallicus ʦ Trichogaster trichopterus ፈɹΥಁᇕອ ݖɴᅊਖ਼ɐʳᰖʢជɇʶʳ⏏
(7) ᝬᐨ⏃˻̺̒⏋̜ͻͲ⏆⏃Ԡሎ 26⏋27⏆ ǵ̩̠̜͌͞Ͳɫɻ⏋Ʉɹྫԍɻ˻̺̒ɭ٧ɽʶ⏋͍˱Ͳໜఌ༑ɭໜᤄྲྀˁᐄʉΥಁᇕᏳ༑ ɴᒞɇʶʳ⏏ᮒ࡚ɹɻɍʠ⏋߂ᩄɹᭇᮒɴʭʳອеɹਸ਼Ϋɫ͍˱Ͳໜఌ༑ɹອȵɄɄˁɭ ȲɌɩໜᤄྲྀɴ༑ʶᦉʞ⏏ɄɹᐨɻɗɹݭଘɴጓɩʰʶɩȦʳೠɹఌ഼ɴܔࡴɇʶʳ⏏ᐨɹ ᒞȴʰɹፈ⏋ΥᦹɹсɴɻភధɹଁϺᅖਸȵਃϟɐʳ⏏ࢪᢒዛɻ Wallago attu ɹᅋ ᧚Ϋᒴɞȵ⏋ʓȴɴཌྷკɇʶʳ˱˓ዀɲɰɹࢸμ݀ዛʢ̮ͻ̚˪ɹೳషɭɌɩգᅎɇʶ ʳ⏏ ǵ̯Ͳ˹ˠ˗ʦ˵̠͌˵˓ɫɻ̜ͻͲɭ٧ɽʶʳ⏏ᮒ࡚ɹވອಁɴᒞɇʶʳȵ⏋Ʉʶʰ ɹɻ͍˱Ͳໜఌ༑ȴʰ᧙ᤂᮐɴȡʳɛʠ Wallago attu ɻɇʶɲȦ⏏ࢪᢒዛɻ˱˓ዀ⏋ ̊˓ͩͲ̝˻͚˗ዀ⏋̯̠͇ͤ̀ዀ ʦ Anabas testudineus ɲɰɹࢸμ݀ዛɫȡʳ⏏ (8) ፊ⏃ดᒞȷ⏆ Ʉɹྫԍɻ 3 ዛᰖ᠗ʠʰʶɛ⏏ 8-1⏏ͦͻ̺⏃Ԡሎ 28⏆ ǵᮒ࡚⏋ވອɌɩອ໗Ɍɛ͍˱Ͳໜೣ༑⏋ఌ༑⏋ፈɹੈχᇕອݖɹࣳᦇʦɗʶɴᦹᐚɐʳ Υಁᇕອݖɹະʦອ໗കɹɹᦲʱ᧑ɴᒞɇʶʳ⏏ଁϺᅖਸɹьᅎɐʳྫԍɫ⏋μɴ ˁৄɐʳɻԂʶɲȦ⏏μ߂݀ɹ Hemibagrus wyckioides, H. aff. nemurus⏋ಁɴɻ˱˓ዀɹ Labeo ፈȵɇʶʳ⏏Channa striata ˁࢪᢒɭɐʳ᭬ɻ⏋ͦͻ̺ɹߡᨆˁዷຩμɴՏɌ ɩȲȷ⏋ɄɹμɴԂɤɛȵዷຩˁُȭɲȦɛʠɴྔʶɵɹˁᬿȽ⏏˛̲ᅎɹͦͻ̺ɻੈχ ᇕອݖɹ͍˱Ͳໜఌ༑ɹाঈɴܔࡴɌɩᒞɇʶ⏋ങГɹᨆՕɻ˵͞Ͳន⏃͠ͅݾ̢̲͠ ̚ͲᏙɹغଁᑥᐣɫೣɫɻᐨକɹᐨɭɌɩьᅎɇʶɩȦʳʢɹ⏆ɫȡʳ⏏ 8-2⏏˵˓⏃Ԡሎ 29⏆ ǵᮒ࡚⏋Υಁᇕອݖɫьᅎɇʶʳ⏏ᒞݭଘɻອᅒˁᑦȽࢸອᤉʦᏳ༑ɲɰɹ༑ʶɹȡʳݭ ଘɫȡʳ⏏ອ༑ɴЫɤɩΫάው֪ɐʳᰖɹᅊˁգᅎɌɩȦʳ⏏ᦲ।ɻʢɰɌɹɦȦɛԂ ؤȵ 1 ɦɫ⏋ອ༑ɴࢪɌԂؤȵά༑ɴكȼʭȩɴᒞɐʳȵ⏋μɴɻԂؤȵ 2 ɦȡʱ⏋ɗʶʰ ৢ૯ 26ȁన࿌ȪΐΛίȫ ৢ૯ 27ȁన࿌ȪΠȜϋȫ
ɹكȷȵɗʶəʶΫ༑ά༑كȷɫ⏋᧚Ϋ༑άؘɹˁɫȷʳʢɹʢȡʳ⏏Rasbora rubrodorsalis, Esomus metallicus, Trichopsis ɹЗᬚ⏋Trichogaster trichopterus, Anabas testudineus, Channa striata, Clarias batrachus, Macrognathus siamensis ፈȵɇʶʳ⏏
8-3⏏˵˓˳̺̒⏃Ԡሎ 30⏆ ǵɄʶɻɫɻɲȼ⏋̤͇ˤ˛͡ᰖˁࢪᢒɭɐʳፊɫ⏋ᮒ࡚ɴᅤɲɰɹྂɤɛܩᯆɴԂؤ ɞȿՏɌɩץᨆˁݏʠ⏋μɴᔼɤɛˡ̢ʦ⏋̮ͻ̚˪ˁԂʶɩৄɐʳ⏏ (9) ፊ⏃ᑖᒞȷ⏆ ǵࢪᢒɭɐʳዛɴʭʱ০Ⴅȵᅵɲʱ⏋ʚɛ⏋ᒞݭଘʢ᧓ɤɩȦʳ⏏ɇʰɴ⏋ࢪᢒዛ ɹؿዌˁЊɌɩɄʶʰˁךաɌɩȦʳݭغȵȦ⏏ȦɑʶʢଁϺᅖਸȵьᅎɐʳ⏏ 9-1⏏̜˕͌˓ˏͲ⏃Ԡሎ 31⏆ ǵɄʶɻ̊˗̠˧⏃ჼܩؿɻ˓ˏͲ⏆ɹʛˁɐʳፊɫȡʳ⏏ᮒ࡚⏋ϗ࡚ˁɭȲɌɩଁϺᅖ ਸȵжеອᅒʦະ⏋ໜᤄྲྀፈອݖɹອཉɹ༓Ȧݭଘɴᒞɐʳ⏏Ʉɹዛᰖɻ̼̲ɹʭȩ ɲк݀ɫմᦽᦡˁᕶܧɴȩɹɫ⏋ፊɴԂɤɛკ႒ˁᦣȵɇɲȦɛʠɴʱՏɌؤˁᯃ।ɴ ৢ૯ 28ȁᡫȪ؍౾̧ȫȪυȜίȫ ৢ૯ 29ȁᡫȪ؍౾̧ȫȪͼȫ ৢ૯ 30ȁᡫȪ؍౾̧ȫȪͼΌΛίȫ ৢ૯ 31ȁᡫȪਸ౾̧ȫȪΠͽθͼͺϋȫ
ᬏȼɌ⏋ɇʰɴɗɹᨆՕɴঈɹᬔɍɛԓፏႥɹጭɫᑂ˃ɞፏˁԂɐʳ⏏ɄɹፏɻᙦɭɌɩล ᔨɐʳɭؾಁɴɄɹˁৄɐʳɛʠɹ͊͊̀ᰖ⏋ᔼɤɛˡ̢ʦ⏋̮ͻ̚˪ɭȦɤɛˁԂ ʶʳᨆՕɴʢɲʳ⏏ᒞɐʳ᭬ɻ⏋ྫԍȵԃɴອ໗ɌɲȦʭȩɴɐʳ⏏ፊɴԂɤɛკ႒ȵዷ ຩȴʰᨧᏬˁ٧ُɫȷɑɴྔʶɲɲȦʭȩɴɐʳɛʠɫȡʳ⏏
9-2⏏̜˕̮͌ͻ˱̜̒
ǵɄʶɻ Hemibagrus aff. nemurus ⏃ჼܩؿɻ̮ͻ˱̜̒⏆ˁࢪᢒɭɐʳፊɫȡʳ⏏ᮒ࡚ɴ ଁϺᅖਸȵອɹੈχᇕອݖɫȡʳɴᒞɐʳ⏏ፊɹүɴᘽɹɦȦɛഘˁɇɌɩˡ̶͐ ͞ͻ˻͖ɌɩອঈɴໍʠʳɄɭʢȡʳ⏏ৄɐʳɛʠɹɭɌɩ̓͌˧ˏ͠ɹॾᛚˁԂʶʳ⏏ 9-3⏏̜˕͌͞Ͳ⏃Ԡሎ 32⏆
ǵᮒ࡚ɴଁϺᅖਸȵ༑ອݖɹੈχᇕອݖɹ͍˱Ͳໜఌ༑ɹाঈɴᒞɐʳ⏏ອ༑ɹȼɩອཉ ɹȡʳཌȵᒞݭଘɭɌɩ᧧ɽʶʳɄɭȵȦ⏏ˁৄɐʳɛʠɴɵȴʦ˹ͦˏ͠ᰖ⏋̓͌ ˧ˏ͠ɹॾᛚˁɭɌɩμɴԂʶʳ⏏˱˓ዀɹࢸ݀ዛɫȡʳ Puntius, Opsarius, Mystacoleucus ɹ Зᬚʦ Barbonymus, Hypsibarbus ɹЗᬚɹᗐȵࢪᢒɭɲʳ⏏͞ͲɻɄʶʰɹᐵዌɫȡʳ⏏ (10) ፊ⏃ᘺɭɌ⏆⏃̅Ͳ⏆⏃Ԡሎ 33⏆ ǵЈɹፊɭ᧓ɤɩԂؤɴʢɰɌȵɲȦ⏏ᮒ࡚ɴଁϺᅖਸȵΥಁᇕອݖɴᅊɍɛࢸອᤉɫ⏋μɴ Ԃɤɛȵອ༑ɹɛʠ᧚ʶɲȦʭȩɲਸ਼־ᨗɲݭଘɴᒞɐʳ⏏˵˓⏃8-2⏆ɭᰖаɌɛዛ ȵɇʶʳ⏏ (11) ᕳᎢ⏃˽͌⏆⏃Ԡሎ 34⏆ ǵϗ࡚ɴɲɤɩອȵৄȷɻɍʠ⏋ອཉȵ༓ȼɲɤɛະ⏋⏋ɲɰɫьᅎɇʶʳ⏏ଁϺ ᅖਸɹʓȴ⏋ଁϺߨਸ⏋ᅖߨӻጣʢьᅎɐʳ⏏ԂɤɛˁΫᨆɴȡʳؤȴʰଝ摑ʛɫʱՏɐ ɭȦȩɄɹྫԍɹ႕ਸȴʰ⏋ࢪᢒɭɲʳᰖɻȡʳዐɹ߂ȷɇɹ Channa striata, Clarias
ৢ૯ 33ȁᡫȪ၂̱͂ȫȪΕϋȫ ৢ૯ 32ȁᡫȪਸ౾̧ȫȪΠͽθρϋȫ
batrachus, Anabas testudineus, Macrognathus siamensis ʦμ݀ɹ˱˓ዀᰖɫȡʳɄɭȵȦ⏏ (12) ാ༡ȿᎢ⏃ˡͻ⏆⏃Ԡሎ 35⏆ ǵᮒ࡚⏋ςɴʦໜᤄྲྀȲʭʃ༑ʶɹᑃʦȴɲ͍˱Ͳໜఌ༑ɫଁϺᅖਸɴʭɤɩʼʶʳ⏏߂ ȷɲጭនɹɊʳႥɹྫԍɫ⏋Ʉʶˁອμɴໍʠɛ⏋ɗɹΫɴೠɹഘˁధዩʞ⏏ೠɹഘɴ ᮃɌɩȼʳ˛̲ᰖȲʭʃࢸμ݀ᰖ⏋ɗʶˁᰩɌɴᮃʚʳ̠͇̀ዀ⏋˧˧ዀ⏋̊˓ͩͲ̝ ˻͚˗ዀɲɰɹᰖȵࢪᢒɭɲʳ⏏ (13) ᒘ⏃Ԡሎ 36⏋37⏆ ǵԂؤɴᙦȵȡʱ⏋μɴȵԂʳɭɗʶȵᬔʚʳʭȩɲ෯ᦶɴɲɤɩȦʳ⏏Гάɹ 2 ዛᰖȵȡ ʳ⏏ 13-1⏏̍͑Ͳ ǵ̍͑Ͳɻጭនɫ⏋ᮒ࡚⏋༑ອɹੈχᇕອݖɫȡʳ͍˱Ͳໜೣ༑⏋ఌ༑ɹࣳȦɴ⏋Ԃؤˁά ༑үɴكȿɩᒞɇʶʳ⏏ଁϺᅖਸȵьᅎɌ⏋߂݀ɹ Hemibagrus wyckioides, H. aff. nemurus, ˱ ˓ዀɹ Labeo ɹЗᬚፈȵࢪᢒዛɭɲʳ⏏ 13-2⏏̍͑Ͳ̮ͻ˱ͻ ǵ̍͑Ͳ̮ͻ˱ͻɻೠɹɫɫȷɛ፯Ⴅɹྫԍɫ⏋ϗ࡚ɴອཧȵᲞȼɲɤɛ᭬⏋Channa striata ⏃ჼܩؿ̮ͻ˱ͻ⏆ȵອཧɹжȦອঈɹ˹˙̊͡ͻɴ᭯ʶʳɭȦȩᅊˁգᅎɌɩȦʳ⏏ϗ࡚ ৢ૯ 34ȁݿٓᢌȪΑθȫ ৢ૯ 35ȁড়૫̫ᢌȪȜȫ ৢ૯ 36ȁȪΙλϋȫ ৢ૯ 37ȁȪΙλϋΩȜȜȫ
ɴଁϺᅖਸȵໜᤄྲྀɫьᅎɐʳ⏏ (14) ᩑʱ⏃Ԡሎ 38⏋39⏆
ǵᩑʱᩍˁьᅎɐʳྫɴɻГάɹ 2 ዛᰖȵ᠗ʠʰʶɛ⏏ 14-1⏏̜̮̾̒˪
ǵ̜̮̾̒˪ɻᅖࡎӻጣȵࢃȴʰᦏȦᤂᮐɴȡʳΥಁᇕອݖɹອᅒʦະɫᮒ࡚ɴȩ⏏ ɭɌɩςɴ͊͊̀ˁьᅎɌ⏋Channa striata, Clarias batrachus, Anabas testudineus ʦ⏋ಁɭɌɩ ˡ˛͡ᰖˁɐʳ⏏
14-2⏏̴̜̾̒ˏ˪
ǵ̴̜̾̒ˏ˪ɻଁϺᅖਸȵ͍˱Ͳໜఌ༑ɹ༑ອɹੈχᇕອݖɫᮒ࡚ɴȩ⏏ɭɌɩ 10 cm еɹ̯̠͇ͤ̀ዀᰖɹ༎ˁ߂ȷɲᩑʱᩍɴᏣɫᐄʃɦȿ⏋߂݀ɹ̠͇̀ᰖɫȡʳ Hemibagrus wyckioides ʦ H. aff. nemurus ˁɐʳ⏏
(15) ൯⏃͠ͻ⏆⏃Ԡሎ 40⏆ ǵ͠ͻɻࢸ݀ɹ൯ɫ⏋ΥಁᇕອݖɫȡʳᲞеອᅒɴᒞɇʶʳ⏏ᒞݭଘɻᅒᣴɌɹອȵᏺ ᇕɴᮃʚʳάеອᅒɹອɹ༑ʶՏɌؤȵಔᦲɫȡʳȵ⏋μɹອᅒɹᅮʦອᅒʭʱάଝɹΥ ಁᇕᏳ༑ɴᒞɇʶɩȦʳݭغʢȡʳ⏏ᮒ࡚ȵɻɍʚʱ⏋ᅒඨȭȵᏺʼɤɛɴᒞɐʳ⏏ ᒞсɻᅒɹଘᓔɫȡʳଁϺᅖਸȵႶɫȩȵ⏋ៀฐɹ߂ȷɲݭغɻភధɹଁϺᅖਸȵԊ ؾɫɐʳ⏏ᮒ࡚ɹᬖࠃɭɭʢɴອᅒԒɫଁᬏᑢຂɌɛȵᭇᮒɫވອɌɛ᭬⏋ອ༑ɴɹɤɩ ው֪ɐʳᓃਸˁգᅎɌ⏋ອᅒȴʰˠͻ̫ͻ̶ͦͻɐʳອɭɭʢɴ༑άɐʳˁ࣊ɴȡʳጭ នɹޭɹᨆՕɴɛʠʳ⏏ᦏɻɄɹᨆՕȵ˵͞ͲនɹᝬɴɲɤɩȦʳݭغʢȦ⏏Ʉɹྫԍȵ ȡʳᅒɹᅮɻᦲ।ɹɗʶɴປʏᲞȼɲɤɩȲʱ⏋༑άɐʳʢɹɻɐʏɩྫკɇʶʳʭȩɴɲɤ ɩȦʳ⏏˱˓ዀɹ Esomus metallicus, Rasbora rubrodorsalis⏋̝˻͚˗ዀɹ Lepidocephalichthys ɹЗ ᬚ⏋̯̠͇ͤ̀ዀɹ Clarias batrachus⏋̜˰˗̠˧ዀɹ Macrognathus siamensis, ˥̨̓͠˗ˠዀ ɹ Anabas testudineus⏋̊˓ͩͲ̝˻͚˗ዀɹ Channa striata ፈȵςɴɇʶʳ⏏
(16) ᎅɹࡎ⏃̶˙ˏ̒˪⏆ ǵᎅɹࡎˁႶɫьᅎɌɩˁɐʳɄɭɻɲȦȵ⏋ͦͻ̺ (8-1) ʦ̍͑Ͳ (13-1)⏋̅Ͳ (10) ɴȵԂʳʭȩɴࢷɐʳɛʠɴьᅎɇʶ⏋͠ͻ (15) ɹ̶˙Ͳ˽ɭɌɩʢਖɑᅎȦʰʶʳɹɫ ɄɄɫɻྫԍɹʁɭɦɭɌɩȡɁɛ⏏ (17) ࢊɓݭ⏃͌͡⏆ ǵΥಁᇕອݖ⏋ςɴອᅒɴᦶʰʶʳԓ০ɹཉʛɫಁɭɌɩɗɹܑٞˁɹܤଝɫᲞȼɌɩȡ ʳ⏏ᦅсɴਃϟɐʳଁϺᅖਸȵȩ⏏ϗ࡚ɴອеȵάȵʱ⏋ȵອɹȡʳɭɄʸɴᮃغɐʳ ᅊˁգᅎɐʳ⏏͠ͻ (15) ɫɇʶʳዛɭᰖаɌɛዛᰖȵࢪᢒɭɲʳ⏏ (18) ᬏനπᐨ⏃̋Ͳ̜Ͳ⏆ ǵᬏɇ 3 m ʓɰɹጭጯɹӴɴሁ৹Ꮭ 15 cm ɹπȦങɹᐨȵɦȦɩȦʳ⏏ᮒ࡚ɹɴ⏋ଁϺᅖਸ ȵ̼̝̒͞˓̜ˁဣတɌɩ⏋ΥಁᇕອݖɴȲȦɩˡ˛͡ᰖˁɐʳಁɴᅎȦʳ⏏ (19) ອɞɌ˽˱̺̒⏃ˡ̅⏆⏃Ԡሎ 41⏆ ǵςɴϗ࡚ɹɻɍʠ⏋ଁϺᅖਸȵອᩄɹȼɲȦΥಁᇕອݖȴʰອˁȴȦՏɌɩˁɐʳ ಁɴьᅎɐʳ⏏ȴȦՏɌɛອɻ˵ʹˑͲ (1) ɴȩȿɩˁ܌ؚɐʳ⏏͠ͻ (15) ɫɇʶʳ ዛɭᰖаɌɛዛᰖȵࢪᢒɭɲʳ⏏
4ȅაȁ݈
4.1ȁݿ႒͈௳ાਫ਼၌ဥ͂କ֖۪ޏ ǵആˏ˻ˏᡉܕɻཁອȵዛధჼࡔᩄɭʢɴȦɄɭɫγᅠᇕɴሾʰʶɩȦʳ [Kottelat 1989]⏏ೣᠭീݖɫʢዛɹᰖȵ᠗ʠʰʶ⏋ɗʶʰɹʓɭ˃ɰɻɇʚɊʚɲᅊਖ਼ݭଘˁው֪ ɌɲȵʰգᅎɌɩȦɛ⏏ɗʶʰɹᰖɻழᩆڧڃɭɌɩᩂឹɲڃᇿɫȡʳɹʛɲʰɑ⏋।ᅊ ༎ɴȲȿʳᕶࢃ༥ᢹɴȴȴɐɄɭɹɫȷɲȦອᅋᣂ྇ɫȡʳ⏏Ʉɹʭȩɲᰖɹᅊਖ਼ݭଘգᅎ ৢ૯ 40ȁၴȪςȜȫ ৢ૯ 41ȁକ̺̱ΑΛίȪΕȫɴࢪਝɌɩܩݖзȵɇʚɊʚɲྫԍˁьᅎɌɩྫ֝༎֪ˁɤɩȦʳɄɭɻ⏋ܩݖзȵɄ ɹܩݖɹᰖɹᅊɴᬠɐʳሾᡫˁמՕɴ୫ɠغʼɓɩȦʳ៷ɫʢȡʳ⏏Ʉɹʭȩɲϟёɻ ആˏ˻ˏɹЈɹܩݖɭʢԊᦲɐʳʢɹɫȡʳቄພ̝͞͡ 1990; Hubbel 1999; Claridge et al. 1997⏏ອᅋᣂ྇ɹਸȵɄʶʰɹܕɹయˁఌȭɩȦʳɹɻᬚ᧓Ȧɲȼ⏋͞ˠ˽ʢɗɹё ɫɻɲȦ⏏ ǵTaki [1978] ɻ͞ˠ˽ɹ͡ˏͲ̺̫͞Ͳȴʰʹˑ˛Ͳ̍͑Ͳᦏ᧿ʚɫɹᨆɭ̮˪́ȴʰ˱ͻ Ͳྡٞᦇɹᨆɹ 2 ܩݖɭആˏ˻ˏعܩɴȲȦɩዛᏼଁˁៗ౩Ɍ⏋ɗʶʰˁປᥒɌɩᅊ ႒ܩჿᇕᓒ࢘ˁɌɛ⏏ɇʰɴ⏋͞ˠ˽ȴʰʰʶɛݬˁμਕɭɌɩ͍˱Ͳໜμ༑ᨆɴᅊ ਖ਼ɐʳᰖɹᅊਖ਼ܩ᧧ି⏋࡚፲ᇕው֪⏋ᰩਸ⏋ᅋೝɹࡴɴᬠɐʳድˁɤɛ⏏ᅊਖ਼ܩ ᧧ିɴᬠɌɩɻ⏋ອݖᄩޅˁܩཙ༑⏋жܩᏳ༑⏋ᑃʦȴɴ༑ʶʳອᤉˁيʞອݖ⏋μ߂ ៀฐɲжܩໜाɹ 4 ɦɴךՕɌ⏋ɗʶɴ༡ອˁ֓ȭɩ⏋ɗʶʰɹᅊਖ਼ܩɴՏჼɐʳᰖˁ͠ ˽̜ˏ̺̒ɌɩȦʳ⏏࡚፲ᇕው֪ɹድɻжܩໜाਸᰖˁࢪᢒɭɌɩ 1 ॷᬚɹᐙᐚᠭീˁ Ȧ⏋ᠭീೝᬚμɐʏɩɹʢɌȼɻ 6 ܌ɹ᭧ᠭീɹȩɠ 5 ܌ГΫᮃɇʶɛዛᰖɭ⏋ᠭീೝ ᬚμ 3 ͶГάʢɌȼɻ 6 ܌ɹ᭧ᠭീɹȩɠ 2 ܌ᦹᐚɌɩᮃɇʶɲȴɤɛዛᰖȵȡʳɄɭ ˁ᠗ʠ⏋մᓔˁࡴመਸ⏋ᓔˁው֪ਸᰖɭࡴᒶɌɛ⏏ɇʰɴ⏋༡ອɴжܩໜा܌ᧉਸ ᰖɹʛɲʰɑ⏋Ᏻ༑ਸ⏋ອਸɽȴʱȴжܩໜाࡴመਸᰖʚɫʢȵѢԂɐʳɄɭˁተ᠗Ɍ ɛ⏏Υ⏋ɄʶʰɹᠭീೝᬚˁɭȲɌɩкᬏ 3 cm ГάɹІȵᮃɇʶɩȲʰɑ⏋ᅋ ֪ʦᅋݭଘ⏋Іɹᔒଁݭଘȵ៨ᦚɇʶɩȦɲȦ⏏ೣድɴȲȦɩው֪ਸɹ᠗ʠʰ ʶɛ Paralaubuca typus, Wallago attu, Ompok bimaculatus, Hemibagrus aff. nemurus ⏃Taki [1978] ɫɻ Mystus nemurus⏆ɻ Taki [1978] ɫɻжܩໜाࡴመਸᰖɴيʚʶɩȦʳ⏏Ʉɹჿᅓɹʁɭɦɭ Ɍɩ⏋ೣድȵΥಁᇕອݖɫɹᅋݭଘʦІɹᔒଁݭଘɹᠭീɴᩂဣˁȲȦɛɄɭʢᓒ ȭʰʶʳȵ⏋Ёᠭീܩݖɹ᧓ȦˁᓒૅɌɲȵʰɇʰɲʳድȵਖឹɭਲ਼ʼʶʳ⏏
ǵRoberts [1993] Ȳʭʃ Roberts and Warren [1994] ɻ˱ͻͲྡٞᦇɹᨆܩݖɴᅊਖ਼ɐʳᰖ ɹᅊˁݬٚɌɩȦʳ⏏Іɴᬠɐʳݬɴʓɭ˃ɰʈʶʰʶɩȲʰɑ⏋ອݖᄩޅɻໜा ɭອ໗കˁ߂աɐʳɴɭɰʚʱ⏋ᅊਖ਼ݭଘɹᏳɲךՕɻʼʶɩȦɲȦ⏏ɌȴɌ⏋ᰖɹው ֪ˁعɅɭɴ៨ᥔɌɩȲʱ⏋ɗɹμɫᯃ܌ᧉਸᰖɭ܌ᧉਸᰖˁךՕɌ⏋ɇʰɴ܌ᧉˁᑢ ຂɭᯃᑢຂɴךաɌɩȦʳ⏏ೣᠭീܩɴȲȦɩ⏋ੈχᇕອݖɫȡʳ͍˱Ͳໜೣ༑ʦ͍˱Ͳໜఌ ༑ɫ⏋ᰖɹው֪ɭᅊ༎رɹᬠѥɴᬠʼʳݬˁɇʰɴؚᮃɐʳɄɭɻ⏋Ёɹᠩᰍɫȡʳ⏏ ǵೣᠭീɹᐄഗ⏋ᠭീܩݖɹອᄩޅɻੈχᇕΥಁᇕອݖˁيʠɩਸɴʛ⏋ȼɹᰖ ȵᅊ༎رɹ႕ࡴɹଁᬏຍ᭥ɫɗʶʰɹݭଘˁգᅎɌɩȦʳɄɭȵ౩ʰȴɴɲɤɛ⏏႕ɴᲞеອ ᅒɹᰖᒴᮃɻዛ෯ଁɴȲȦɩႶᕶɹʢɹɫȡʱ⏋ɴੈχᇕອݖȴʰɹᦜԂɴʭʳʢɹɫɻ ɲȦȩȭɴ߂ីʓȴ 2001⏋Ʉɹܩݖɹᰖᒴᮃɹᅊ႒ਸɹᲞɇɻ⏋Υಁᇕອݖɭੈχ ᇕອݖɭɹᬚ⏋ੈχᇕອݖɹໜᤄྲྀɭ͍˱Ͳໜɭɹᬚ⏋͍˱Ͳໜɹఌ༑ɭೣ༑ɹᬚˁΫάው֪
ɐʳɲɰ⏋ɗʶʰɹᅊ༎رɹμɫᅵɲɤɛອݖᄩޅɹᬚˁው֪ɐʳɄɭɴʭɤɩѳɛʶɩȦʳ ɭȦȩϟࡸɻ႕ɴᇿɇʶʳ⏏͍˱Ͳໜɴᅊਖ਼ɐʳᰖɹው֪ɭΥಁᇕອݖɹᩂឹਸɻˡͲ̓ ˻ˏʦ͍˱Ͳ̊̚͡ɫೣᠭീɭᰖаɌɛᠭീˁࡸేɌɛᇐቄ [1967]⏋Ꮫ [1975] ʦ Taki [1978] ɹድଁഗɴȲȦɩʢ୭ɇʶɩȦʳ⏏ᰖȵᅵɲɤɛອݖᄩޅɹᬚˁ৵ው֪ɐʳɄɭˁخ ᔨɴɌɩȦʳɹɻ⏋ɴອɹ༑ʶȵᦹᐚɌɩȦʳɹɫɻɲȼ⏋ɗʶʰɹᬚɴᰖɹው֪ˁᭀࢀ ɐʳʭȩɲ᭭ࢀȵɲȦɭȦȩɄɭɫȡʳ⏏Ʉʶˁلݻ [1991] ɻᅊ႒ᇕອᄩɭዌɌɩȦʳ⏏ ǵჼܧɹೣɫɻ⏋ੈχᇕອݖɻʢɠʸ˃ɹɄɭ⏋࡚፲ᇕɴੈχᇕອݖɭᦹᐚɐʳອᤉʦອᅒ ɭȦɤɛΥಁᇕອݖȵ⏋ɇʚɊʚɲᰖɴᅋݭଘʦІɹᔒଁݭଘɭɌɩգᅎɇʶɩȦ ʳɭȦȩɄɭȴʰ⏋ɗʶʰɹݭଘȵཁອᰖᒴᮃɹᅊ႒ਸɹᐣ୫ɴɻɛɐᩂឹɲ৲ւȵ ᇿɇʶɩȦʳຽऊᅒμ 1998; ᚹټອᢆ 1999; ႅᩃʓȴ 1988; ఱᚹ 1997; ఱᚹʓȴ 1988; ᦅ ᄩޅҺ́Ͳ̊ͻ 1997⏏ ǵɌȴɌ⏋ჼܧɹೣɫɻݜᇻҺϟɴʭʳܘݭҺɹ᭬⏋ᅎອᤉɭகອᤉˁՕᮐɌ⏋ᅎ ອᤉɻ̮˓̺͞˓Ͳɇʶʳɭɭʢɴອᅒȴʰɹອɻᘺेɹᯃ।ɴ߂ȷȦகອᤉɴᘺɠʳɛʠ ᰖȵອᅒˁգᅎɐʳɄɭȵέخᔨɭɲɤɛ⏏ʚɛ⏋ອᅒʦᅎອᤉɴᦹᐚɐʳໜाɫɻᦅᅎ ອݫʦቆᬿݫݨɹᒞɴʭʱᰖɹው֪ȵᘿɌȼߺɁʰʶʳɄɭɭɲɤɛ⏏Ʉɹʭȩɴᅊ႒ ᇕອᄩȵᘿɌȼᭀࢀɇʶɛᐄഗ⏋ҺɹອᅒʦɗʶɴЊ᭦ɐʳᅎອᤉɫɻ౩ʰȴɴອᅒ ᅊᏙɹᰖɹᅊ႒ਸɻཝࢹɌɩȦʳຽऊᅒμ 1998; ጩ 1985; 2000; ႅᩃʓȴ 2001; لݻ 1991; ᦅᄩޅҺ́Ͳ̊ͻ 1997; ఱᚹ 1984; ᅒμ 1999⏏ ǵɇʰɴ⏋ᅎອᤉɹሁᐻʦ⏋˱Ͳ˪͠ͻ̜Ϊᯆᡸࣳȵɐɐʛ⏋ɗɹȩȭໜाఒѹɹ᭬⏋ໜ আɴʭʳाঈɹॶܹɴЊ᭦Ɍɩဈɭཌȵ༥߉Ɍɛʱ⏋жອᤉȵሁᐻɌɛɄɭɲɰɴ ʭʱ⏋ອᄩޅɹਸȵ߂ȷȼ༥߉Ɍɛ⏏ɄʶɴЫɤɩᰖɹዛధ⏋҅кధȵɭʢɴཝࢹɌɛ لݻ 1991⏏ ǵೣᠭീܩݖɴȲȦɩɻ⏋ᰖɴᬠɐʳᅊ႒ᇕອᄩɴࢪɌɩੲ৯ᯱˁؖʖɐʭȩɲϺဤᇕ ۊဤȵʓɭ˃ɰʼʶɩȦɲȦɄɭʢȡʱᖪ߫ɲႥɫȡʳɭȦȭʳ⏏ɌȴɌ⏋˰Ͳ˱˪ᦏ ᧿ɴȡʳੈχᇕᏳ༑ɫɻ⏋ဌྰᅎອɹɛʠɹᦅᅎອݫȵলɇʶɛɄɭɴʭʱɹው֪ȵ ߺɁʰʶɛɹɴ֓ȭɩ⏋ɗɹά༑үɫɻອȵྡྷᅥɌɩᄩޅȵੲɌɩȵɭʶɲȼɲɤɛ⏏1⏆ ǵЁɹ͞ˠ˽ɴȲȦɩᰖᒴᮃɴ৯ᯱˁήȭʳϺဤᇕۊဤɻ⏋ྫ֝༎֪ˁաɴɐʶɽဌྰϟ ɭ̋͌লɫȡʳ⏏ɄɄ 10 ॷഀ͞ˠ˽ɫɻᎦɹԃᕶᐌ᧒ଁɹɛʠ⏋ϗ࡚ɴʢဌྰˁɌɩ Ꭶˁᅊᅋɐʳకፑˁᦽʠɩȷɛ⏏ೣᠭീܩݖɫʢ̩̠̜͌͞Ͳʦ˰Ͳ˱˪ɹٞᦇɫဌྰᅒ ȵᦶଁɇʶɩȦɛ⏏ೣᠭീܩݖɫɹဌྰϟɫɻ⏋Ϋ៨Ɍɛʭȩɴອᇿᇕɹᦅᅎݫݨˁল ɐʳɄɭʢȡʳȵ⏋Ʉʶʭʱʢ͍˱Ͳໜఌ༑ʦៀฐɹ߂ȷɲȴʰளອͅͲ̺ˁьᅎɌɩဌ 1⏆ ᛚ౩੯ᅊລȴʰɹኽѵɴʭʳ⏏
ྰᅎອˁተѳɌɩȦʳݭغȵȦ⏏Ʉɹဣɫອݖᄩޅʍɹ৯ᯱɻࢹɲȦɭȦȭʳ⏏ᅎອᤉɻሁ ᐻᇕɫΪᯆ˱Ͳ˪͠ͻ̜ᡸࣳɞȵ⏋U ࡓྋɫɻɲȼ⏋θࣳɻᑃӃహɫȡʳ⏏ʚɛ⏋ອঈɴɻȦ ɑʶɹݭଘɫʢȵྊʚɤɩୈອඨ႒ɹᅊᔒɌɩȦʳᨆՕʢȡʱ⏋ᰖɹᅊਖ਼ᄩޅɭȦȩဣȴ ʰੲɭɻਲ਼ʼʶɲȴɤɛ⏏ᅎອᤉȴʰᅒʍɹᘺेɻࢹɲȼ⏋ȿ༑Ɍɇʶɛອɻɴɻܤ ᒽ⏃Ꮼʱ⏆ɹອᤉˁɭȲɌɩΥಁᇕᏳ༑ɴகອɇʶɩȦʳ⏏ჼܧ⏋ೣᠭീܩݖɫʼʶɩȦ ʳဌྰ০ɻᅊ႒ᇕອᄩɹែဣɴɛɩɽ߂ȷɲکᰍɻɲȦɭՠుɇʶɛ⏏ ǵဌྰᅒɻೣഀॵΫȵɤɩȦɛݭଘȵϗ࡚ɴȲȦɩԗʃླྀອɐʳ⏋ϺဤᇕɴᦶʰʶɛృɌȦ Υಁᇕອݖɫȡʳ⏏2001 ॷ 3 11 ⏋̩̠̜͌͞ͲɹဌྰᅒɫᮃᠭീˁɤɛɭɄʸ ȋ3⏏ᐄഗȍɹ፲ɫ៨Ɍɛʭȩɴ⏋˱˓ዀɹ Esomus metallicus ɹॾଁɭ Rasbora
rubrodorsalis ɹॾଁ⏋̝˻͚˗ዀɹ Lepidocephalichthys sp. ɹଁ⏋̊˓ͩͲ̝˻͚˗ዀ ɹ Channa striata ɹॾ⏋ˠ˽̺̦ͦ͌˽ዀɹ Trichogaster trichopterus ɹᅋআɭ ॾ⏋Trichopsis schalleri ɹॾଁȵᮃɇʶɛ⏏Lepidocephalichthys Гɻॾ⏋ȡʳȦɻ ᅋআʦȵ᠗ʠʰʶɛɄɭȴʰ⏋Ʉɹݭଘɫϗ࡚ɫʢɄʶʰɹዛᰖȵᑢຂɌɩȦʳɄɭɻ ተࡸɫȡʳ⏏Ʉɹȩɠ⏋Trichogaster trichopterus ɭ Trichopsis schalleri ɻϗ࡚ɫʢອਸɹੈχ ᇕອݖɫᑢຂɌɩȦʳȵ⏋Esomus metallicus, Rasbora rubrodorsalis, Channa striata ɹ 3 ዛᰖɻᘿ ᓔʰɹᠭീɫɻ⏋ਃഀɹᅊਖ਼ݭଘɫɻϗ࡚ɴȲȿʳᑢຂˁተ᠗ɌɩȦɲȦ⏏ృɌȼօՏɇʶɛ ΥಁᇕອݖɴѢԂɌɛɄʶʰɹዛᰖɴᅊ༎رɹȵᅊɍɩȦʳɄɭɻ⏋Ϻဤᇕۊဤɭᅊ႒ɹ ࢪਝɭȦȩဣȴʰᖂ٢ཉȦჼᢒɭȦȭʭȩ⏏ ǵʚɛ⏋Ʉɹϟࡸɻອᅋᣂ྇ɹވ֓ɭȦȩᯆȴʰʢᇿɇʶʳȵ⏋ᅥઝɌɲȼɩɻȦȿɲȦɄ ɭɻဌྰᅒɹᄩޅɫȡʳ⏏зɹ᠌ɴʭʳɭဌྰᅒɴɌɩȴʰዝɹࢀᛚ⏋႕ɴ˪̼͐͠ˡ͍͌ ˹ Leptocorisa chinensis ȵ߂ᩄɴᇍᅊɐʳɄɭɴɲɤɛɛʠ⏋Ʉʶˁᱤ᭑ɐʳɛʠɴຏᛚջˁь ᅎɐʳਖឹȵᅊɍɛɭȦȩ⏏ʚɛ⏋ᔋషˁత॒Ɍɛɴȵ˃ɫ༝ȴʃΫȵʳɭȦȩ⏏ᰩష ɭɌɩգᅎɇʶʳᰖɻ႒ᣒɴູയɇʶɩȦɲȦɄɭȵմரИɫȡʳȵ⏋ອᅒɴᅊਖ਼ɐ ʳᰖˁອᅋᣂ྇ɭɌɩգᅎɐʳ᭬ɹᠩᰍɭɌɩЁᩂ߂ɲઝˁɻʰɤɩʪȼਖឹȵȡʳ⏏ ǵ͞ˠ˽ɴɭɤɩᮚɻܕɹᢧ྇ˁʳɛʠɹᩂឹɲᇿɹʁɭɦɫȡʳ⏏ɗɹɛʠ⏋ჼܧȡ ʳʢɹɴ֓ȭɩЁȼɹ̋͌ˁলɐʳᅙȵȡʳ 1996⏏̋͌লɴЫȩᅊᣂ྇ʦ ܩݖзɹᅊ༎ʍɹ৯ᯱɻ߂ɲʢɹȵȡʱ⏋ອᅋᣂ྇ɭྫ֝༎֪ɴ߂ȷɲȵᅊɍɩȦʳ ϟёʢݬٚɇʶɩȦʳೠؤ 1999⏏ೣᠭീܩݖɫɻჼܧɹɭɄʸ̋͌ȵࡔܧɌɩȦɲȦ⏏ ǵೣᠭീܩݖɹ⏋ʓȴɹܩݖɴʛʰʶɲȦႥɭɌɩɻ 2002 ॷʭʱ̫Ͳ̯˛ͲाΫ༑ɹ́ͅ Ͳᦏ᧿ɫᩆɹȵᬖࠃɇʶ⏋ፀ 2 ຍ᭥ɭɌɩ᪅᩿ɹᬖᇍȵϜࡴɇʶɩȦʳɄɭɫȡʳ⏏ᩆ ɹᏄ᪹ɻ˹ˏͲغ႒ˁьᅎɐʳȵɭʰʶʳ [Oxiana 2001]⏏ɄɹɴࢪɌ⏋ᅣ [2000] ɻ̺͞Ͳ̜கອμɹ˹ˏͲغ႒ȲʭʃঘɇȦμɹ˹ˏͲ⏋ອ᪃⏋᪅⏋᩹⏋ϩ᩹⏋ˡ̝͊˗ ͌ȵᄩޅʍ༑Տɐʳکᰍˁ୭ɌɩȦʳ⏏̶ˑ̴͠Ͳɫɻࡸ᭬ɴɄʶʰɹ႒ᣒȵܑٞɴ༑Տɐ
ʳϟఖȵᅊɍɩȦʳ⏏ɇʰɴ⏋ܩݖзȵອ᪃ˁьᅎɌɩᩆˁᏄᑇɐʳˁȩʭȩɴɲʶ ɽ⏋ံลອ᪃ɹᙜຩˁُɤɩံลອ᪃μນɴɲʳɽȴʱɫɻɲȼ⏋ᕶးᅠɫลɌɛອ᪃ȵ ᰖˁيʞᅊ႒ɴᙢዩɐʳϟȵᅊɍʳخᔨਸɻᲞȦ⏏᪅ɹɭɌɩɻᮚៗȵᅎ ɇʶʳ⏏Ʉɹɻᩃɴ᩿ቄˁݡዩɌɩॕቘᨧˁΫȴʰత॒ɐʳɹɫ⏋ܩάອʦໜाɴཉխ ɲອᣒູയˁȼ᭝ਸȵȡʳᅣ 2000⏏Ʉʶʰɹᄩޅکᰍɴᬠѥɐʳ႒ᣒɹΥᨆɻອѶᆖ ʦ˓̊˓˓̊˓ᆖɹ܍ɭԊᦲɌɩȦʳ⏏ɄʶʰɹᆖຩɻᲞɴᅊ႒ᑘɇʶɛ܍႒ᣒˁ ɐʳອᅋ႒ˁᰩɌɩᇍᆗɐʳϟёȵȦ⏏Еɴ⏋̫Ͳ̯˛ͲाΫ༑ɫᲞɴູയɇʶʳ ᰖȵঃݖው֪ዛɫȡʶɽ⏋ອᅋᣂ྇ˁᅎɐʳɄɹܕɫ⏋ঃȦ፴ܑɫᆖຩȵᇍᅊɐʳخᔨਸ ʢوࡴɫȷʳʢɹɫɻɲȦ⏏Ʉʶʰɹ႒ᣒɹ߃းອݖʍɹ༑Տɭອᅋᣂ྇ᩄɹ֪ˁ̢͐̊͠ Ͳ˫ɐʳɭɭʢɴ⏋ɄɹອݖɴȲȿʳᰖɹው֪֪ˁيʠɛᅊˁਸ਼ɴៗ౩ɐʳਖឹȵȡ ʳ⏏ 4.2ȁူমު͈ຽݞ̷͈͂ऻ ǵອᅋᣂ྇ˁᅋɐʳݭଘɻԊܩɫȡʱ⏋еɫᅎᇰณȵࡴɇʶɩȦʳݭغȵʓɭ˃ɰɫ ȡʳ⏏͞ˠ˽ɫɻฌᲞɹΥᅱжȦाᑉɴеᒞɐʳȋाȍ⏋ɗʶɴᐚȼॶܩɹȋॶܩȍ⏋ɗɌ ɩॶܩɹΫଝɫεɴȡʳȋεȍȵȡʱ⏋Ʉʶʰ 3 ɦɴךՕɇʶʳǾȵ႒Ǿϯழ⏋ϯ౫⏋॑ ݭɲɰɹᡉ༎֪ˁɭȲɌɩȡʳ႕ࡴɹܩݖɹμɫʁɭɦɹ̢̜̒͘ˁɦȼɤɩȦʳɹȵಔᦲɫ ȡʳ⏏2⏆Ʉɹ̢̜̒͘ԒɫɻʓȴɹϺʢ͡ͻ͡ɇȭࡪʶɽয়᠍ܩɹգᅎˁኧɇʶɩȦʳʼ ȿɫɻɲȦ⏏ɲȴɴɻʚɤɛȼɭᑉɹɲȦϺᬚɴʢˁ᠗ʠɩȦʳݭغʢȡʳ⏏ʚɛ⏋ອ ᅋᣂ྇ɹѳᡸˁᇿᇕɭɌɛྫ֝ៀըɹʓȴɴ⏋႕ࡴɹ֪႒ɴࢪɐʳȲɗʶ⏋ᏄᮠѵЖፈɹɇʚ Ɋʚɲឹ܍ɴʭɤɩ⏋ᘺеʢɌȼɻܩݖеɫࡪʰʶɩȦʳៀըȵȡʱ⏋ᐄഗᇕɴອᅋᣂ ྇ˁѳԃɌɩȦʳɭȦȩ⏋ܧഀɹᣂ྇፬ჿȵࡔܧɌɩȦʳ [Iwata in press]⏏Ʉɹʭȩɲը ɹμɫܩݖзɻΫ៨ɹʭȩɲྫ֝༎֪ˁɤɩȷɛ⏏ ǵ͞ˠ˽ɫɻЁᇵ˃ɴຂϟȵࣙᬖɇʶɩȦʳ⏏ຂϟɴᬠɌɩ⏋ჼಁဣɫɻዛᗈˁՏ ᗼ˵˓̀ɴʚɫᔒଁɌެɐʳܕጓɹݭɻࡔܧɌɲȦ⏏ɌȴɌ⏋ຂɹ៊ˁᰲᔒɌ⏋ ɗʶʰɹዛᗈᅊᅋˁɤɩȦʳ́Ͳ̊ͻɻ˵ʹˎ̠˭̜̒ላɹݭغ⏋̮˪ʹ˝ͻɹላጓዛᗈ ᅊᅋ́Ͳ̊ͻɫȡʳ⏏ᬚɹຂ́Ͳ̊ͻʢȼࡔܧɌ⏋ɛɭȭɽ˰Ͳ˱˪ᦏ᧿ɹᬚᓔɻ ˱˓⏋̠˓̘͡ˑ̴͞ˏ⏋Barbonymus gonionotus⏋̩˪ͤͲ⏋Cirrhinus cirrhosus ˁຂɌɩȲ ʱ⏋ᕶՕɹຂɫዛᗈȵᅊᅋɫȷʳ˱˓⏋̠˓̘͡ˑ̴͞ˏ⏋Barbonymus gonionotus ɹ ʦ៊ˁٞᦇɹɴެɌɩȦʳ⏏ᦏɫɻ̊˓ȴʰዛᗈȵᇵ˃ɴᥥԂɇʶɩȦʳ⏏ܩӯɹ ϺɻܕᅋʭʱʢڃᣒȵᖪȦɭȦȩ⏏
ǵຂɭɌɩɻਃഀȴʰࡔܧɌɩȦʳɴ֓ȭɩ⏋ᦏɫɻ᧑ᤉʦဌྰᅎອᤉɹҺɹɛʠ
ɴʱάɁɩອȵྊʚɤɛݭଘˁգᅎɐʳϟёȵވȭɩȦʳ⏏ʚɛ⏋ဌྰɹɛʠɴɓȷʠɛ ᨆՕˁຂɭɐʳёʢ᠗ʠʰʶʳ⏏ ǵᘺɫɹຂɻ҅Ϻͤ̾͡ɭɹԊˁьɤɛϟɭɴ߂աɇʶʳ⏏҅Ϻɫຂˁȩ ϺɻዛᗈȵᣘԂɫȷʳɴʳᣂᅋˁɌɩȦʳ⏏ᘺɫຂˁȩݭغ⏋ዛᗈˁɹᣂᩆɫᣘ Ԃɐʳݭغɭ⏋ɲ˃ʰȴɹອᅋஹ֖̺ͦ˻˙˪̜ɴʭʱዛᗈȵံషɫఌᐌɇʶʳݭغɭȵȡ ʳ⏏ɗɌɩ⏋Ȧɑʶɹ˭ͻ˽ʢዛᗈȵఔ༑ɇʶʳອݖɫɻྫ֝༎֪ȵ؏ᩂɴៀըɇʶʳɄɭɴ ɲʳ⏏Ʉɹʭȩɲёɻೣᠭീܩݖɫɻ˵̠͌˵˓ɹᘺԊɫ᠗ʠʰʶɛ⏏͞ˠ̫ˠᦏҶ ɹ́ͅͲɫɻਃഀɹອᅋ႒ݭଘɞɤɛךݖȵຂɴɲɤɛɛʠ⏋зɹݭଘ ȵИɹੲȦଘɴೃˁоәɲȼɇʶɛ⏏Ʉɹ˭ͻ˽ɻܕ᭬ᇕ NGO ɹˠ̒˪˽̶ˎ͌ɹᢩܐ ࢪፑ̺ͦ˻˙˪̜ɴʭʳʢɹɫȡʳ⏏ ǵჼܧ͞ˠ˽ɫຂϟɹɇʚɊʚɲ᠅ʛȵʼʶɩȦʳȵ⏋Ȧɑʶʢ๘ᎦʢɌȼɻೣȵς ࢷɌɩȦʳ [Modadugu et al. 2000]⏏ɗɌɩ⏋ຂϟȵᬖࠃɇʶʳɭዛᗈʦɗʶˁຂɐʳ ɴ౩ተɲ҅ϺᇕଘณȵࢷԂɇʶʳɄɭɴʭʱ⏋ɄʶʚɫɴɲȴɤɛகЈᇕըȵᅊɍ⏋ਃഀ ʭʱࡔܧɌɩȦɛԊʦԊؾɹᅎᇰณȵ༥߉ɐʳ⏏ᓔɻ͞ˠ˽ɹᕶးᄩޅɹμɫᅵɲɤɛ ጓܩИɴ࣎зɌɩȦʳϺǾȵԊࡔɐʳɛʠɴᬏಁᬚȴȿɩօʱȡɁʰʶɩȷɛ˹˽̘͌ɫȡ ʳ⏏ΫᦚɌɛʭȩɴ⏋॑ݭςᒶᇕែဣȴʰʛʶɽѝғɹɲȦࢸμ݀ɹᰖɫʢ⏋зɹᕶࢃ ༥ᢹɹɛʠɴɻᩂឹɲອᅋᣂ྇ɫȡʳ⏏Ё⏋ຂϟɭਃഀɹըˁθጓɇɓʳʭȩɲը Ϋɹू߅ȵਖឹɫȡʳ⏏ ǵຂϟɴʭʳອݖᅊᏙʍɹ৯ᯱɴɻ⏋ɇʰɴ๖ɹʭȩɲʢɹȵȡʳ⏏Гάɴɗɹёˁ୲ Ɂʳ⏏ ǵ̊˓ɫᣱȷɩȦʳϟёɻᅊ႒ᇕᅊᇕɲែဣȴʰʛɩཉխɫȡʳ⏏̯̠͇ͤ̀ᰖɻ ̊˓ɫʢإȼʭʱᰩషɭɌɩঃȼգᅎɇʶɩȦɛ⏏͞ˠ˽ᯊ٨⏋ౙ̅ᦹɹࢭᬐࢃȵ߂кᬏ 1.5 m ɴʢɲʳȵ٢ȵੲȦˏ̶͠ˡᅋɹ Clarias gariepinus ˁ͞ˠ˽ܕԒɴࢷԂɌ⏋ຂˁᬖ ࠃɌɛ⏏᭰ܕɹ̊˓ɫɻ⏋٢ɻчȦȵଁᬏȵᧆȼɩᆖຩɴȦ Clarias macrocephalus ɭɄɹዛ ˁϯᮊɇɓ⏋ʭʱ߂ȷȼɩ٢ɹчȦዛˁօʳɛʠɹ᠅ʛɭɌɩ Clarias macrocephalus ɹᮇɴ Clarias gariepinus ɹᮁˁϯᮊɇɓɩຂዛᗈɭɌɛ⏏ɄʶʰȵᩃɴᦣɁՏɌɩܧഀዛɹ̢̒ ̍ˁߥȩɽȴʱɫɲȼ⏋ܧഀዛɭϯᨗɌɩዛȵगޙɌɩɌʚɤɛܩݖȵȡʳໜᩃ 2001⏏ ǵ͞ˠ˽ɫʢᇵ˃ɴᩃఔ༑ɇʶɩȦʳ˱˓ɻ⏋ɐɫɴؾɲɄɭȵʼʶɛȦȼɦȴɹܕ ɫɻ⏋ອᗳˁᰩࢀɌອɹˁވ֓ɇɓʳɭȦɤɛອݖᄩޅɹੲˁৄȷᣱɄɐɽȴʱɫɲ ȼ⏋ЈዛɹជᰩɴȲȦɩʢཉխɲ৯ᯱˁؖʖɐʭȩɴɲʱ⏋ˠͻ˽̜͞͠ˏɹʭȩɴᣌ ᩆˁȴȿɩɄɹዛᰖɹᱤ᭑ˁᦽʠɩȦʳܕɐʰȡʳ [Baird 1999]⏏ɇʰɴෑጩɲёɭɌɩ⏋1970 ॷБɴˏ̶͠ˡɹᰩషลˁఛȩᇿᇕɫ FAO ɴʭʱ⏋Lates niloticus ȵೣഀɻɄɹȵȦɲ ȴɤɛ̲˪̜͠ˏྲྀɴఔ༑ɇʶɛ⏏ɭɄʸȵ⏋Ʉɹɻ 200 ዛˁᣳȭʳܧഀዛˁޙྙɇɓɩ
Ɍʚɤɛೠά 1998⏏˱˓ɭؾɴ͞ˠ˽ɫᩃఔ༑ɇʶɩȦʳ̠˓̘͡ˑ̴͞ˏʢγᅠع ܩɫᅊᏙʍᢦɹ৯ᯱˁؖʖɐɄɭȵਥɇʶɩȦʳ⏏2001 ॷɴ IUCN (International Union for the Conservation of Nature and Natural Resources) ɻ⏋ೣഀɹᕶးᄩޅȴʰաɹݭଘɴውʳɄɭ ɫᄩޅɴੲ৯ᯱˁήȭʳ 100 ዛɹ֪ඨ႒ˁ͠˽̜ˏ̺̒Ɍɛȵ˱˓⏋Lates niloticus Ȳʭʃ̠ ˓̘͡ˑ̴͞ˏɭؾࣚɹˡͩ˽͍̀ Oreochromis mossambicus ȵɄʶɴيʚʶɩȦʳృ ᓬ 2001; Global invasive species database 2003⏏ɗʶəʶɹܩݖɫອݖᅊᏙɹ෯ᦶȵ᧓ȩɹɫ⏋ Ʉʶʰɹёˁሁஞ͍˱Ͳໜ༑ݖɴȡɩɻʠʳ៱ɴɻȦȴɲȦ⏏ɌȴɌ⏋Ȧȼʰ͍˱Ͳໜອݖȵ ঃ߂ɞȴʰɭȦɤɩ⏋ɗɹ৯ᯱȵᇖံɫɻɲȦɞʸȩ⏏Ёᩂ߂ɲᬠਕˁʢɤɩܧഀອݖᅊ Ꮩʍɹ৯ᯱˁ̢͐̊͠Ͳ˫ɐʳਖឹȵȡʳ⏏ ǵᰖˁఔ༑ɐʳɄɭɻᄩޅʦዛᰖ෯ଁɴ৯ᯱˁήȭʳɽȴʱɫɻɲȦ⏏ѳԃᅊ႒ɹែဣ ɫɻᅊ႒ਸɭȦȩɭȷ⏋ዛ෯ଁɹਸɻʢɠʸ˃ؾዛԒɹ᧨Шᇕਸɹѳԃˁʢɗ ɹែএɴȦʶɩȦʳᵠᢆሼ 1996⏏ᅊ႒ɻؾዛɫʢ᧨Шᇕɴᅵɲɤɛ႕ਐˁ୫ɤɛȦȼ ɦȴɹᮃȴʰଁጓɌɩȦʳ⏏႕ࡴɹ᧨Шᇕ႕ਐˁ୫ɦᮃˁɗʶɭɻᅵɲɤɛ႕ਐˁ୫ɦ ᮃɹμɴఔ༑ɐʳɭ⏋ɗʶʰȵϯᨗɌɩ߃းɴɻࡔܧɌɲȴɤɛ᧨Шᇕ႕ਐˁ୫ɦᮃȵ Տჼɐʳ⏏ɄʶɻϺဤɴʭɤɩᣱɄɇʶɛ᧨Шᇕɲఉϕɫȡʳᨙ 1997⏏ჼܧ͞ˠ˽ɫɻ Barbonymus gonionotus ɹఔ༑ȵعܩɫᇵ˃ɴʼʶɩȦʳȵ⏋Ʉɹɻ͍˱Ͳɴೣഀȴʰᅊਖ਼ ɌɩȦʳɲɹɫഀɫȡʳ˱˓ʦ̠˓̘͡ˑ̴͞ˏɹёɴປʏʶɽܧഀᅊᏙʍɹ৯ᯱɻ ࢹɲȦɭȦȭʳȵ⏋᧨Шᇕਸɹែဣȴʰʛʳɭکᰍɹȡʳဤɫȡʳ⏏ఔ༑ɐʳݭغɻؾ ΥອᏙԒɫ᧨Шᇕ႕ਐȵᅵɲʰɲȦᮃˁ᧧ʉɄɭȵਖឹɫȡʳ⏏ ǵ߃းອᅋᣂ྇ɹཝࢹɴ֓ȭ⏋ຂϟɹୟ߂ɴЫȩຂɹಔؖɴʭɤɩ⏋షჿˁᇫɴᇵʳ ᭬ɴ⏋ΥיɫɠʬȩɰȦȦ߂ȷɇɹ⏋Ȧʼʪʳ̘ͻ̸̶͡ˑ̒˹͖ɹʛˁᩂែɐʳʭȩɲѝ ғ៎ȵʚʱ⏋ࢸμ݀ᰖɹᮛឹȵཝࢹɐʳɭɐʶɽ⏋ჼܧʼʶɩȦʳྫ֝༎֪ʦᅊȲ ʭʃᰩయɴʢ߂ȷɲ৯ᯱˁήȭʳɞʸȩ⏏ʚɛ⏋ຂɴਃϟɐʳϺɭɗȩɫɲȦܩݖз ɹᬚɴ҅ϺଘፈɹဣɴȲȦɩᐁཕᇕൔेȵঃȵʳخᔨਸʢȡʳ⏏Ё⏋߃းອᅋᣂ྇ɹ ɴ֓ȭɩຂϟɹ֪كɴʢઝˁكȿɩʪȼਖឹȵȡʳɞʸȩ⏏ɇʰɴ⏋͍˱ͲໜອᏙɴᅊ ਖ਼ɌɩȦʳᰖɹᅊ༎ر⏋ᅊɹៗ౩ˁਸ਼ɴȦ⏋ࢰഀɹອᅋᣂ྇ɹᣒᇕᩄᇕɴএ ɴࢪਝɫȷʳݜኄᇕݬˁҺȭʳɄɭȵʚʶʳ⏏ ৫ȁৃ ǵЁ܌ɹድɴѤࡶˁɻȴɤɩȦɛɞȦɛ͞ˠ˽ܕጓ߂കᨆ˽˪˱Ͳ́Ͳ ˵˓̠ͤͻ̜ᨆᬏ⏋̜Ͳ ͠ ˵͓̍͑̒˪ᦅᨆտᨆᬏ⏋LARReC ɹ˵˓̺̚͞ͻ̜ǵ̢͖͇̍͞ͻଘᬏ⏋ˡ̲ͅͲ ̶˝˗̊̓Ͳ ລ⏋˹Ͳ̊˗˝Ͳ ʹˑ͞ͅͲລ⏋ʚɛᠭീɴؾɇʶɛ͞ˠ˽ܕጓ߂ɹ˽̒˪̅ͲͅͲ ̺͠˪˵ͻລ⏋ ˵ʹˎ̜̒ລ⏋͞ˠ͌ˏລ⏋̊ͻ̠Ͳລ⏋LARReC ɹ̅Ͳ̸Ͳລ⏋̊ͻʹ˝Ͳລɴཉᡚɐʳ⏏ʚɛ⏋ᠭീ ลೳʦฌೣɹѳ፬ፈɴਉתȦɛɞȦɛ LARReC ɹ˵˓̺̚͞ͻ̜ǵ̢͖͇̍͞ͻଘᬏ⏋ˡ̲ͅͲ ̶˝˗
̊̓Ͳລɴઢᡚɹઝˁᝠɐʳ⏏ʚɛ⏋ჼܩᠭീɹ᭬ɇʚɊʚɲݬˁரњɇʶɛϺʁɭʱʁɭʱɴਉ ᅕɌΫɁʳ⏏ ǵೣድɹลЦˁήȭɩȦɛɞȷ⏋ԃᯆᇕɴᠭീˁఌ୫Ɍ⏋ɇʚɊʚɲᯆɫתȦɛɞȼɭɭʢɴᢵᩂɲ ݬˁήȭɩȼɞɇɤɛϴᨊ߂߂ˏ˻ˏˏ̶͠ˡܩݖድድዀɹإाχᮁؿ᠕ఠஐ⏋ቄᅒᏛᨀ մఠஐ⏋ጭᅒಅϓ֖ఠஐ⏋ࢸܮঔψລ⏋ϴᨊ߂ആˏ˻ˏድ́Ͳ̊ͻɹᚹٮᮁ֖ఠஐ⏋ᛚ౩੯ᅊລ⏋ ኜକ߂ࢸകЭڈ֖ଝ⏋մϴᨊ߂ᦅᨆɹകᩁᗔລɴȼਉᅕɌΫɁʳ⏏ ǵೣድɻ̜̊͜ᢧ 1999 ॷድ֖ଁ B ֖ଁᅱش 99B3-103ȋ͞ˠ˽ Bang hiang ा༑ݖзɹᅊɴȲ ȿʳᅊᣂ྇գᅎɴᬠɐʳድŅШᐏᇕయɹѳࡔᐙମɭᄩޅѳԃɹែဣȴʰŅȍ⏋̜̊͜ᢧ 2001 ॷ ድ֖ଁ B ֖ଁᅱش D01B3-130ȋ͞ˠ˽ Bang hiang ा༑ݖзɹᅊɴȲȿʳᅊᣂ྇գᅎɴᬠɐʳ ድŅܧܩɹሾਜ਼ɹ៨᪼ɭಔؖŅȍȲʭʃ⏋ዀድᢹݜᇻድ (A)(2) ᠩᰍᅱش 10041071ȋᅵᅊᏙஞ៘ɴ ᬠʼʳϺؤው֪ɭᣂ྇գᅎ˹˽̘͌ɹᢟȍ⏃ድБᝠᓔ⏧إाχᮁ⏆⏋ዀድᢹជ֖ᩆμᇕድဣ ০ଁ̺ͦ˫͌͞ᠩᰍᅱش 10CE2001ȋˏ˻ˏˏ̶͠ˡɴȲȿʳܩݖᑂଁ⏧݀ࢅ᥊ଁȍ⏋ॶଁ 14 ॷ ϴᨊ߂ఠᔒድ୷ᖂᢧፀΥشϟ༠༐֖ଁ⏃ቁೝ⏆ɹ֖ଁᩆɹΥᨆˁ؝ȿɩʼʶɛʢɹɫȡ ʳ⏏ ֨ȁဥȁȁࡃ ȏృᓬȑ2001 ॷ 5 14 ⏃ϴᨊႆ⏆ȋ˓Ͳ̾ͻ̋ͻ֪ඨ႒⏧100 ˁ୭ࡴ⏋ᡰٚ⏧ܕ᭬ѳᡸลᬠȍ Baird, I. G. 1999. Fishing for Sustainability in the Mekong Basin, Watershed 4(3): 54-56.
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ᅣǵ౩ᨀ⏏2000⏏ȋᦽʞ᩿ᬖᇍɭᑵʱᦐɇʶʳ᩿ࢀȍȏˏ˻ˏᄩޅᇐೆ 2000/01ȑആᐁཕృݬ⏋37-62⏏ ጩǵૠϠ⏏1985⏏ȋᦅອᤉɹᰖѳᡸɴɦȦɩȍȏཁອȑ11: 64-72⏏
