3種トミヨの交雑実験
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(2) ^ 10 m, ^ 2 ^ (^^) m\\ 34 £)2 12 ^. 3 it h ^ 3 ©. 5C ^. ^ tt ^. ^mw^^mww. Hiromu KOBAYASI : Cross-experiments with three species of sticklebacks,. Pungitius pungitius (L.), Pimgitius tymensis (NIKOLSKY), and Pungitius sinensis (G-UICHENOT), with special reference to their systematic relationship. 1. Ikeda (1933) studied the geographical distribution of 3 species of sticklebacks in Japan, and reported that Pimgitws pnngitius (L.) distributes in the coast ragions of the Pacific Ocean and the Sea of Japan, Pungitms sinensis (GUICHENOT) in the coast regions of the Pacific Ocean, and the Sea of Okhotsk, and Pu.ngitins tymensis (NIKOLSKY) in the coast region of the Sea of Japan. The present author has found the mixedhabitat with P. pungitius and P. tymensis in the catchment-area of the river Ishikari, and that with P. ptingitius and P. sinensis in Akkeshi region. Several specimens intermediate between P. pungitws and P. tymensis were captured in Sapporo and Ebeotsu regions of the catchement-area. of the river Ishikari. In addition a few individuals showing an intermediate charactsr between P. pzcngitms and P. sinensis were obtained in Akkeshi region (Fig. 3); Tables (6-7). Some ecological observations revealed that P. pnngitms and P. tymensis are similar in both habitat and breeing season. 2. The results of reciprocal crosses carried out between three species of sticklebacks indicated that the development of eggs ane the time of their hatching are nearly similar to the normal case. In the coruse of development the eggs showed a mortality nearly similar to the control eggs (Fig. 2). The larvae were similar in several bodily characters to the maternal form just after hatching to 30 days. Then, an influence of the paternal form made its gradual appearance in the larva. Almost all larvae grew normally in both behavior and appearance : they grew into adult form about a year later. There were well-developed gonads well in both male and female specimens. 3. In every combination of crossing, the results of measurement were nearly intermediat in value between the parent species, and also almost identical in mean value to those of the reciprocal hybrids (Tables 8-11). The individual specimens possessed, however, the characters of the parental species, as. well as those intermediate between the parent in various proportions (Table 12). The results of measurement of bodily characters carried out in the intermediate forms found in nature were nearly similar to those of hybrids produced experimentally.. 4. Based on the results of the present investigation the following conclusion may be possible that the three species of sfcicklebacks here considered are in close relationship in their taxonomical situa-. tion, and that the intermediate forms of sticklebacks occurring in nature may be the hybrids produced in field.. ^-%^u-^ t- y^^wM^^m^w^mt^mm, :^w (1933, '35, 'ss, 'se, '41) ic:. ^tg^TjA$Bffl^;?;?, —m®^)^^?^^'^. .Ih^Mdx^b^^^^T^ 1y^ ^WM^^^^w^t ^mm^&k ^, ^ ^3M?^^, ^ y" i- ^ 3 PtmgUius iymensis (NIKOLSKY), 1- 5. 3 Pungitius siwnsis (GUICHENOT), A ^ 7 1- $.3 Pungitius pungi-. tius(L.) (D 3 @®^gt^> C ^^'^tJb^TM^. L^L, A^-Mtc^U-^ ^^b 3@®®^X^:SM <l:-^®^:e®19^^^^T^-^^^^^.tM^^^/@^?f/^^@-5a^, '/feS (1933,'35), /? —363—.
(3) ^. /^ ^. (1957) ©^{CcUzU;', ^Zb®^l, ^-y" I- ^3<1:^^7 ^ $ 3, it£^~{c:-f^7l-^3^1-$3®. rSR:f^T(^, ^l—j:t!i^K:NS®7l^m^b^, ^-^b®?g^J:fi^l^?c:^M^NS®^PB^c1: ®;?^0^/f@W^h't^»^^L-CJg^. t^W(1934){tj^<^®^g{c:^-r, -f ^7 {. ^ 3 ^^-y" 1. ^ 3 <^ 2%=42, n=21 ©^-fa-^^^^-^, ^®^^}C®&^T<fc%/^^^^®gi^. btZ^l^^^LTl^. y±®^iil^it^LT^SR:, —j^^g^^^^^T^^ ^-.jZb ^. ^ 3N®3@®-fe^f3s]{c:^i^T^, ^^^mwK^^^^w^^T^^m^m^n^.. JII-N-, W (1947), i^ (1950, '51, '52, '53, '57) iH?J'H^-Jte^XT^ ^-7 V ^ a !^®»il. it^?l\ ^^b7&it@rB^c:^U-^^3a?^t®^^^gfcl^-C^g^aL^^r@<9, /? (1957) (^A^M^F^a ^^l-t^X3@®ra]®x%^^^;fT^^, -e®^^B7i^^, ^^b®^®%f<!. '(^®^|^fT^OTjg6. CCDm^WS.^^WW^^^K&b'C, ^-@®g^J®^^~ j^b^-6^(c:^oT^. ^^\±^m, ^m^cj^t^^ ^^ } ^ 3, ^y"1~^3, i>^3®3 gr^, •^z-€:'tuE^»^^i\ ^-nb®%^a7®, ^^^s®m^^^T<l:i^f^^, ;&^. ^^%@t.®^s§n^^^{<:,?^fi]j^^^t.u:^Hj^/c. cn^®as®^t^^, @?^w@w ^7 i~ ^ s^^-y'^ ^ 3, ^^71- ^3^ 1- ^ 3®i^@®4]I^SI^^^^^'f@<4:<l:®J:b^^^^. i\ l< ^ 3^311®^ ^^©-^i^^iclfc^T^^^M^^. ^<:{<:—j^®^H:^^.^®^^^T 7s.. B^^a6^^^c^'^, M^®^f15, ^®/?@^%i:^^P^^^^A^M^:'^Ft3ls^^;fe fei7-. ^rJW^^b^HJT^. t^^/fTO®fM^Mll{c:m^T^^B-% ^0-iS-J'HN ?^MLT^^-{'c:^itT^.. mfc'c^CWS -CTL^I- ^3M®3@®^m^, ^Lfe Wll, yl^^, ^JII. ^Jll, glfiU, J?^, ^li, ^ ^1, ;?M®;&:W<^^a-<^/c^®-(?, ^%^^{c:.^mL/c<fc®^.(bU,rl7^^g?14^®Fs^c:. f^^-C, 1t@JIIW^6D^I-^<fc^»L^ 1- ^ 3^fL^T{T^^®^^Ji[7j<^J;^.f:^^-L/;-f^ 7t-^3, ^y'l'^^^.. ^l|iiaR:{CTL/c^®(t, ^LT-jCTJIKcW, ^^{c^l^T^B^ffl|^^^J-f@{cN L, ?^-®^T^®^^o/;. ^Kc^L, ffi^{i/f"r^^@®^?S^^^i^^lt-r^/f@{^f'c:^ ^10, (lflb®rBl^A:C?^|^T^-3^. C. Table 1. Numbers of cross-bred experi-. ® ^ -t 7s W\? ® i?^ ^, Mi^ ® B ffi ^. ments bewteen the 3 kinds of sticklebacks. n L <, srrau ^swm^w^^m^. in each year.. In the table abbreviations are used for. ^o/c^®-??, flX^1±^T^»:R^^tW{c:%. indicating the 3 kinds of fishes as follow-. WL^Wjis^/;. MWm^i/^- v. ing, Pungitius tymensis (NIKOLSKY)---P. (., Pungitizis pungitius (L.)'- •P. p., Pungitius. }c^, -XSI-t^^^cj^^fil-^L^.. sinens'is (GUICHENOT)'--P. s... ^ k c O^^^a^i^ LT{t, ^-B?X^- ^ ffl^. Cross. ^^^•lo^ir^m^^k. ^a^M^. 1954. 1955. 1956. Total. ^7j<-?®^Anit7X^St^?&^ , Eg^ 15 cm ® ^. P. (.? XP.p.S. 7 ^ i/ -^ - L^ {CAH, B (c 2, 3 [U^K^T^ ^. P. p. ? x P. (. S. 4. 1. 5. ^^ ^^]0 ?^5?^ L ^)/;. ?^ L^-ff^. P. s.? xP.p.S. 3. 2. 5. ^M.Tih7X4'^^WL, J^M^-^T^R::^. P.p.9 XP. s.S. 3. 2. 5. ®^t|}^L, WUc:{tig^33cm, ^^ 20cm. P.s.? XP. t.S. 4. 4. P. f:9 XP. S.S. 6. 6. ® ff 7 ^ 7X-ft[^ {c 2 WJ; ^) 4 /f@fr^A^> 6%-^ —364—. 2. 10. 8.
(4) 3@ h ? 3<D^. ^^M-t^L^. t"]WK:ffl^/;7j<MT7j<^, r"J^^LT^, 5.^^^, ^ t- ^ ^^ffllV;. ^^ ® ? (C ^ 10% (D^^-7 ^ y a^-'fM L, s^^X, ®, ^E^^© M? ^ ^ 4% ^ ^ .=- y g$7j<^. «M3S^7^^. ISS3JS3^^(Z)^M^^ • A^MtC^y-^ 1-^ 3^^®|®^^^;^{c:^^T(t, rfeB(1933), /J^(1952)^|®:^i.a5(.5' —®:L, ^m^mQ^m^t^t^y!j\\w, ^(Qwm'^j^^, ^^7 i- ^3^ i- $302 Ei^^r&^?^^MAT^MJfl^^^®M^%^^^, ^y" I- ^ 3^-f^7 t- $ 3®2@®, ^^-y^ms^M^A-t^^]\\w^(Qwm^^^, i- ^ s I@HI®^?BLT(^. L^L^fi:^*-^^'%-'/Q^%^J;^^^^^/;^^?^{c^^T, 1- ?. 3®^XT/5y^^^H L^-c. (IO^^^TC^, B^^'/&^^®^y" I- ^ 3 ^-f ^7 I- ^ 3®^XT^W®$tJ«5;tbtL -C^CB^, ^m^^^'V-^^^, 3@® F ^ 3^c®.?.^(c^,fT^y^%~j^^/)z^i5Tt^. -?^^^>^(c^^b^^. ^/=@^?^^^^T^, M;f|®^^bT, y&^t^^^^^^:7 ]-^ 3 ® ^feXT ^> y^g^^) b -Hr (/ ^.. ±tg^ 2;jsix® i. ^ 3^1g®^X^^^b/t'zn^^¥^r&^^i.^®J?^^ s ^W^mcQ ^CTJfl/J^cg^T, N@®^X^fil®Sli^^^^-^^^11, J?^?^®-^^7 1- $ 3^ ]• ^3, /CTJll7j^®^y" 1- ^3^^^^ 1- ^ 3®^(t, N^^^ ^^'^^yl^^;^^^ L, f^-7jc^® 7"IJfl?^T^, WW^(Q^^'^M\W^^m~^^Wt^^. <=-(D^. aEg^o^fyJtKc^ U-^"f^;7 ]^ 3^ 1. ?, SOyM^-fitc:, N@®^^6^^Sfr^B^)b^^^ L/C?felS(1941)®^<J:. (moT@^{c^otz^. ?7lM{cWS 2 @®t^$lg®W^LU^-NR:Xx-3T, m^m^-t^^, ^®^^%®@9^^ Fig. 1 (C^T^-9-^^^. ^®%g^^l955^:J;<9 2^^,. s^ g 100^ 0). ^. V 90 -. s. "80-1 d. a 70 H d 60 -1. ^. ^ 50--j. ^ 40 ^ 30 -j 43 20 -j at. h bO 10 -I. March April. I. T. May June. Collecting date. July. Fig. 1. Comparison of the breeding season between 2 kinds of sticklebacks, P. tymensis and P. pimgitms. ••••P. ti/mensis 0---P. pnngitms.. —365.
(5) ^ ^. 3A. ^fi^]o^n ^xsr^c, 20 ["iiwfToA^^o-it.F^^m^jii, ^m^db is ^w^ ]o. iW?^fu3®tiMjn, »:?^®wit^®w-^T^^®^, ^-^-as (ci^t^^^p. f^rffi^, i!.-fe^-faa®^®mM^{9, Ns®^ig»!®^:s§^-Niij?L^. N@®^m%^^^T ^, ^y~ 1-^ 3^-^A^^< 3^T^}^^ 5^^^g®^^^^®{.c:^L, /f^:7 ^ ^ 3{t4^T^T Ai9 6 n^^]^(Dm^&'0, NSO^^M^^RS^^y-'I-^ 3^'4^43/HJ, -f ^71-^3^5^%. •OT, ^(Dm^20 By±®Mlut^^, ^^si3Ma^^^^^/i;B^^^®tt6?®^^^^fit m^^^^^O, NS®^m%^—^^^^T^XLT(/^;b®^B^L^. t^i^c:5^±^J ®~MJH^-^TMt$®Bt^L^^y" 1- ^ sttoNIS^^ ^7 1- ^ soi^^-fajt^^" 1- ?. 3®. ta^^^^B, g%(/^^6D^®^%;S-»T-5WHj^, X^^y" 1- ^ 30^^-^-f ^7 1- ^ 3^-fa^-feX-l, i^l^l-^^7 t- ^ -a(D^^^-/} ^ s^^m^^^-'^p^f^^m ^—m^Wmir^'Vk Li^'L^'^o^.. ^®^}c?g^?(c^U-^N@f^®^X^:lM^'@^-rt§^T^»:^<t^, /;ch^N@®rB^^ ?PS^ ^^ ^ b LT ^, N@®rs3^^^^^-^^^^> ^ o -9 »^T L ^S^-r^»{tHd^^^ h^7^ b^Z^>.. ^M(D^t^^ F, (D^m.^ 1-^3, ^^71-^3, 3iy"i~ ^ 3®3@^®iE^^%(c:^^-r, ^nb^b^G^^^^-^y. ^HjX^^i'c^, ^@^^-<i:®^^^^i^T^a^^'M:^§^^^^»^^^> M?<1:J^T ^>^%/^z^ia^%^^^^-o^. ^-a^^-^w^^ <^, •^®a®^^^^(t%^<l:'^x^%^T »^<, ^^m, ^(DW, %^^-%{c^T^B^B€:^A^i3-MPSE<h-^L-r(/^. W^, ^. 2 Rt ^ ^ JC ^ ^ ^ < {; -}g L, 30 f^ rai tc ^ »^ ^ ^ ^ , Hatching^ Heart formation-j Eye balls appear 6 myotome stage Formation of embryonal body G-astrula stage-1 Blastula stage-| Morula stage-] 16 cell stage-i 2 cell stage-. 10 20 50 40 50 60 70 80 90 100 M:ortality of eg^s {%} Fig. 2. Difference in the mortality throuth the process of development of experimental eggs produced from the crossings between 3 kinds, P. pzmgitms, P. tymensis and P. sinensis, and those controls. 7\-- -the mortality from minimam to maximam of the controls. ^••-the mortality from minimam to maximam of the experiments. 366—.
(6) 3@h ? 3®2<^. OTsi3 Qm^^ww^^k- t^-%^a^4:'^^-r^^p®Bi%^t^t/c^-mE^{5^®:L, (=-^^(D&m± Fig. 2 (C^T. ^^b®^^p^{C^^-C^, {-&®M@, ^^|SN5<lW-. {•^M^tz^p^, ?(Q-W?^(D^WcMir^^^mW, -?^®M^^^^T. w^^n^-b^tL. y-c/c t- y'^^mm^^^^nkM^^t^^, ^^m^^^ W<, ^aE®^^bt;^-?X{c:^^T^-^{<:^^^^B^Fcl:^^^tL^^^/c. i|tU-7-^. ^m™c^tz^:fc|^K:?y)TCT^X?, ^SlK®ffi%^^^. ^ffi^3^^ ^»^L^ff^^/fnT^^@^TjE'g>(c,&l^t^^^ L, ff-^tga®^M(A^-ffi^ t)-^^^ 4.5 mm^^, MUcm^N^aX®^^^ , MB?Xfft.<1:^: < -®: LTlVc. gu{-T^imcW^pM?®HW»^{cS-^ U^'Uf%-?^%a^&^, ^®^^|t^^. ^b/cfftjc^-cu;, Wo^mw, ^-(Dm^WWWi^^niT, ^WCT^PM. U^HT^;-^. »ew^© 80% ^±^Bufft.^%j; o aff^^i^Bip^®^^^^^, fin^ ^ ^ ^ ^. mw^. 10 B ^B^ L ^ -fft. ^ aff-t. ^ ^ o, {"T^ ^ ^M 7 mm K± ^ ^ ^>. ^ ® R^-% ^ (^ ^m%/^^:K:%^tL, ^it^TOL, ^-^©^s^fli^^^^i:, it{c:m^(A^^^%;b. T^(c^. c??{c{f^,S^m, H$^ff^(t%/i.Z'^^L^. ^-^b%@®fttf, ^{-T. M®Mm-f5?;MCTW®M?^-®:L, ^(Df^W^WW^t^- MB 20 s^l?. Wtil, Wi-l'c^^^ l;, ij?a 30 Ei ^•a^T^i^'^M 12 mm, {^ 2 mm, -f^^ 2 mm. ^<9, ?J, m^^^J^L, t, ^^^^^!C,&-5'<, ie, )sM^^^^t^, !m^{c;i?L TffiBLT^^^^^'^^c^^^^^. »/f^40 B^aMT/tZ^', -^-M^ 19mm, ^^ 3.5 mm,. ? 2.3 mm ^ 4' ^, -ft.a ^ (^ ^^® ^-fa-fe^^^ ^, -^^ ^f^ {C^gB^ ^ ^o ^. t ^ fM, ^ ?i5i3-^L, BXM, !^^;^^^{c:-%ifL, ^^^M-ffi^i'c^^^: LT^%^:^^'bgi^ b/tz^^ (c^o/c. C(D^m^]0, --§-^®M^^^-ff-fe^a^(lc$<:^®^^^%^L^a6^. %^%!'c-A-3. /;ff^, ^ffi^^{lcJ;6?CT?jm^^{c:ML, $<:^®^^{c:J;(9, ^^m^®^^E{f -^.^ ^, ^(o^rn-^^^ LT^. -^^^, ^ y i- ^ 3 ^ i-?. s, ^V" ]- ^ 3^-<^7 1- ^ 3®N. M^o-^J; ^ ^l;^ff^®mft"X, ^-MX^{iii5{3;'Na@M^Efft.®^^;&^t^$P^^t^^ •fc '•t^. m^ 50 B^^miM, w^^^m^^]o, -^yi' ^ ax i- ^3, ^-y- ]- ^ 3 X -^ 7 1- 5. 3 ^©^-M^^^^ (9 ^ b ^-f@'^®^^-^M 25 mm, 'f^ 5 mm, -^g 2.7 mm, ]-. $ 3 X /f ^7 t- ^ 3 ® M^-^-^ c^ <9 ^ 1; ^-f?®^^ 27 mm, ^^ 5 mm, 'f^^ 2.7 mm b ^. 10, C(DW(D^^mm, ®^^^^^L, Wk^ff^bk- ^oTC®B^{c:^-r@€(c: ^-M^^ ^l^fft^, ^-^^®ffi^^-^ft^-®^^^^^^T^^ ^O^^^b^^). ^ ^?^-C{^ Table2 (C^L^$P<, ^ ®tNf%!ic^^®^^t.^'lB% LTJ^-S ^, UzM]t-h Table 2.. Average hatching retio ol each cross-bred egg, number of each hybrid at 3 steges of the growth, and their sex. Average. Cross. hatching ratio. J^L. Number of hatching. la rv a fishes.. Total length. Total length. ^mm). (mm). 10-20. Sex. 20-60. P. t. ? xP.p. s. 35. 300. 98. 78. (? 25,. S 17, ? 35. P. p. ? XP. (. S. 42. 150. 72. 42. ? 19,. S 6, ?. P. p. ? XP. s. S. 38. 130. 61. 37. ? 8,. $10, ? 19. P. s. ? xP.p. S. 40. 130. 65. 35. ? 8,. $11, ?. P. S. ? XP. t. 3. 38. 100. 68. 34. 9 9,. $12, ? 13. P. t. ? XP. S. s. 36. 150. 82. 65. 913,. S 27, ? 35. —367. 17 16.
(7) ^. ^. ©^Wc^;5<b®^, ^|l{Cjj(@;?T^®7m^ch^;f^n^. »/f^ 70-80 5^a;@1tztf^y" l ^ 3<l:-f ^ 71-^3, ^y'~^$3^^^ 3®NM^-^^-®^j^®^^^X{l34mmtu. a, ^^7 ^ $ 3<h^y" 1-^ 3®^?^®^^l^M(t36mmgfi^<i:^^, ^J;l^)^te,£l^tl ^^•tmKt£-^k. L^L, /?^-^®M®^^{t, ^©Rt^lt^lg^-^, -^S^^MX®!!^^^ ^®^tz<l:?^T^i{f, <i®^-»!®Ji?<^M®JtW, )^^®^tzJ;^^^X<%^tL^. ^® ^l?a®M®^lfd, dWm^&^^^&t^, 6%S®S^R:^, ^^i9B^^^{@^M-^%^ ti^^{c^oT^y-;. ff<H^4~5 ^^^B7iT^^^{9^^®<gT^?^^, d^LbM^®)^ '^-W]bt^mwm^i'^k. ^tzb®^^^ii^^^j<fc^s^®3^%?^j^x^iig^< l?®:i$-^4^L, J&T^X^TM-^L^), S^3^43^J^^^^N^®^^-^r?g^L/;B^ ^a^-^-^®<f@-?ch^EFi^®^, 3a^®6^, gp^$i?<ba^i^^^^^<1:^-^^. ^ ft b ®^^^^^W%3F1S^;ia l;%^ti^n$M/f@/N-^@ < -ffi^-?? ^ ^. XSx^l®^^—B^ {±s%t-^^%w^^]<l:B?(f^^$p^^®{t^/^z^^btL^^-3^y-nc1:', m^®5^^, ^ ^iit^si^%^T^®^?^^%^n^. cnbdtegi^Mo^'t^x^, —IIJBR®^. ^^, ^wmwmw^t^^(D, M^%^®c?ciii, g%i^A3gs^^-T^^®^^ Ji^bfl, -f^7 i- ^ 39 X I- ^3$^^)^, l'f@^®M:?X^S!i^~^^L^. L^U, ^tZb®^. MM?4^c:^ l~2^(D^m^t^^<o, ^TL^MS»®^iJKgl<l:^;f^b'?^. d;}Zb®id$MS, ^{A-TOH;, -fnI^^^^aiO BKrt{C^^T$gXL^. 3@h^3®CT(::ji[(,^r J.y'I-^3, -f^7}'^3, I-^ 3® 3@H§®^|ga®^lt®^^^^^-5tc:^^, ^^^. ®&;^tg;&%^^nT^^mL/;. L^L, %^?S^-^^nT^^^^b 3@® ^ ^ 3®. W^W^Kst^^^, ^(DWb L-CN^^M^, ^n^M(DmKWt^^^. {c:^T'L^3g^<l:®^/ti-5^®tf^^^(t^^-^^. Jc<i-^T, ^-%^^^^L^3g® l- ^3. ic^'o, -^nb®—l^??CT®im®SII|?^f?\ M{c^iiHc^|(cffl(/^20W4:®;§-@® 1-. $ 3?c®e^T, mwm^ww^n^^^^k.. Table 3. Comparison of each mesuring part of the bodys; body height, head length, trunk length, tail length, length of caudal peduncle, and height of caudal peduncl, among the 3 kinds of sticklebacks. And the mesuring parts are. shown by mean value ±tandard deviation of length of mesuring part/body length, in the 10 individuals of each sex in each kind. {°/o). P. s.. p. p.. p. t.. Body height. Head length. Trunk length. Tail length. M ± a. M ± ff. M±a. M ± a. Length of caudal peduncle. M±a. Height of caudal peduncle. M ± a. $. 18.0 ±0.71. 29.7 ±1.02. 30.0±T.20. 40.8±1.04. 11.4±0.85. 2.6 ±0.27. ?. 17.9±0.64. 28.8 ±0.75. 30.6±1.20. 41.0 ±0.98. 11.8 ±0.45. 2.6 ±0.22. s. 18.5 ±1.02. 30.0 ±1.13. 31.2±0.77. 39.0 ±1.10. 10.2 ±0.63. 2.9 ±0.24. ?. 18.7 ±0.54. 23.7 ±0.64. 31.4±1.06. 38.7 ±1.00. 10.6 ±0.96. 3.0±0.30. s. 20.7 ±0.72. 32.9 ±0.70. 34.0±1.10. 32.3±1.07. 5.9 ±0.60. 5.0 ±0.31. ?. 20.6 ±0.90. 31.8±1.12. 35.6 ±1.20. 32.9 ±0.83. 5.8 ±0.62. 5.2 ±0.25. —368—.
(8) 3SE h ? a ®3$| 1. -<A-7 h^ 3 Pungiiitis pungitius (L.): ^^(1—^^^M < , {-^'fa(A 3 ijmgt^H^^. ^{c:^, »^;tLT@i9, ^MiJo^^^^^^B^^-??^-^^^, ^M%!?^%^n^at'^^® M-fa^-Sl^m^^-^. -^-^©MW, )^.^^?^^B<M^L, f%^(^8,9,10^^ ^W^tobfl, mode ^9^^^^. Mffi^(i^X<, M,&§®N^^^^R^^^^T^-6.. I?®'M;?^^? ^sw\7]^(D^(D^im^^w^w^^m^^n,' s-j^,. i^^®^ © }c (A, tu^J; ^@ffi^®m'i:X^-^^'?M^^?XL G^m^^^%^T{@^^yM ^LTt/Vc. t^%^®^t^^®^{t^S'fe^-^L, Sg®^-^^^^®^^-^^-^^. Table 4. Comparison of each mesuring part of the body; length of central dorsal spine, pelvic spine, pelvic girdle (A) and (B), among the 3 kinds of stickleacks. And the mesuring parts are shown by mean. value ± standard deviation of length of mesuring/body length, in the 20 individuals of each kind. {%) Length of pelvic girdle. Length of pelvic girdle. Length of central. Length of pelvic spine. M ± a. M ± o. M ± tf. M±s. P. s.. 6.1 ±0.37. 11.9±0.87. 17.4±0.83. 12.1 ±0.90. p. p.. 5.8 ±0.42. 10.1 ±0.74. 15.4±0.69. 10.8 ±0.82. p. t.. 3.7 ±0.34. 5.8±0.85. dorsal spine. AAL. _IB)_. 8.9±0.64 .. 4.4 ±0.47. 2. x1/' 1s S. 3 Pungithis tymensis (NIKOLSKY): ^@®^fit{A 3 @®® I- ?. 3 43.^1fei. ^^, ^U^itLT^. fr^^^^N&^^o^', ^^^I§<, frfMiJ©^m^^^^t^^ ^o/c. L^L, ^m%!^%^^^?^fr'fe®li^^^Si)?aj^^^-^^. -^^-^©MX^)^-^® ^MW^m<, t%®-f/?c%^%;MVj:i^®-^, gm;(B)^®^'?t^,i:a®%/^af M^_)b^^^^^^®^%ji^^^. t%?3;ji@i^^<, 9~12^^®/f@/^^g^i6b^, mode {111^:^^-3^. m^M^<;fc<, MIII®J?^S^R^;^~T^6. ftS1;X®^,td 3@^.f^M<, JS^cjgL, ^^Mffi^P^I. -fe-fe^rt^, %/^H-fe^LT(^. Table 5.. I/H^^{cT^(^. t/c^tiO^ta^d.S^M. Comparison of each mesuring part of the body; number of dorsal spines, soft rays of dorsal fin, soft rays of anal fin and scutes, and types of scute-row, among the 3 kinds of sticklebacks. And the mesuring parts except the type of scute-row are shown by mean value ± standard deviation of the mesuring numbers, and mesuring numbers from minimum. to maximum and their mode, in the 20 individuals of each kind. Number of dorsal spines. M±a P.s.. 9.1±0.30|. p. p.. 8.8 ±0.56. p.t.. Mode ( ). Number of soft rays of dorsol fin M±ff. (9), 10] 11.1 ± 0.77|. Number of soft rays of anal fin. Mode ( ). M.±a. 10, (11), 12] 10.3 ± 0.91|. 8,(9 ),10| 10.9±0.57| 9,10, (11), 12|. 10.8 ±0.88| 9, 10, (11), 12] 10.6 ±0.58| 9,10, (11),. 9.7 ±0.88 9.6 ±0.66. 369. Mode 8, 9, (10), 11| 8, 9, (10), 11|. Number of scutes M±<7 33.9 ±0.30. (2.5 ±1.03). +(10 ±0.95). Mode. 33 (34) 0-(2)-5+. 7-(10)-12|. 9,(10),U|0+-(5.2±0.78) 0+4-(5)-7. Type of. scute-. row.
(9) ^ n. §A. 3. h S. 3 Pungitius sinensis (GUICHENOT) : ^@®^^(^ 3 ;g-%® ^ ^ 3 ifi ^ <b ^H^ «?L, ^fe^-f^7l~^3<l:^y"{-^3®^rB]^, ^t-fe^^^^-L, W>S;<fc?i^?BJ5at-r^-) /c. ^.mm ^ ^ u- ^ ? ^ ® 'f^-fe (A , ii is- ^ y" i- ^ 3 <h f^ ^ i.c ^ ^ ^ ^ < s ^ L /c. -^^-^ ® M x y-'. ^'?Si?^?;tL, tM?9~10;^, mode ^9^-Z?^^,. Mffi^d^^^ffl-M < , M(tS ®Ni?R^^<, M^^b-C^. ti^^S^TX<$l5tL/;A311-^, g^-gp^^Mffi^ ^R:34{W,?L, 1t(c:J?<:^T(A, ^^^®iFM{c:jS^-oTi/^. t ^^|^®^#Ma{AM'fa -fa^^^<, ^-fe^^LTl/^. il'J^li1: ±§G 3 \ ^ 3 ®W??®^11/& Tables 3, 4, 5 {c^-T. Table 3 {^@. m^m^iom^iw, ®-x, gis, mx, mffi^®¥^;&/N-^®^-^^i^b/c^®® 100^^^, Table 4(ttta;& 10 ®fr^^^L, ^tMM, f^^A^, gp-t5^^ i^:B^, gp^, ^^'tt^,t5M®¥-^^-?M®¥^^'!^L.^^®®100^^^^^, Table 5 {i ^ ^i??-& 10 'i?^ B^ L, t i. '^(D^^^, ^. n b ® mode ^^(t^^^jfiL/c <fc ®^^> -S. ^ H b ©MIJ^P^^H/C LT^-@iN®^fig6^/f:SS^. M^mZ^W^S^Cj^T, F^W^^^^^^-3^. ^©^JI^TableS ®^^ y \ ^3^ i> ^ Si.C^i/j-^-ft^, ®M, BIX®^^, Table5 ® 3 ii®^^W, -^fl^'tl95% ®%-^{C-^^T^-^®^^J^i6^-^^'j±',^y-;®^, —j^^^b. ®^^^^-@IXrB^®%,S^%^TN}^^/&^^^b^^>. MK;<itzb®??m^llHc:WT^ {c, %^zit^-r®^7t(<:^T, -<^7 t- ^ 3<1: l' ^ 3?i6T7&{OT^>/?;&^LTj^®? L, ^V" 1- ^ ^hA ^:7 \ \ s^y<~ i- ^ ^(Dm^^^^ ^, ^ts. ^^n^'N^'%^^^^^- ^ ^ L, "t^-l^^ir^i^ -c ®<^ (^ d ® ^-1^^^ ^ ^ ^, Figs. 6, 9 iC ^^^n^$D < , ^ y" I- ^ 3 b-\^:7 \ \ 3®?f;X{t7i^?S-^, •€®^^i'5i?%-f^LT@^, %/±K^m\^k \ ^ 3®l?^<h. (i^^oTl^. C<DWMQW^W^tt^, A ^7 \ \^b \ \ 3^-tW^rat-^ ^^t?i<N6Tfi^re<^^> <b®T?6fe^.. ^H'+'MWWS 2@%®l- ^ 30'/g^^Xitiic:, N@®43!M;^T/fW®;?T^?, ^^rlkiB (1933, '35) ic:J;D-f^7 1- ^^^ [ ^ 3i.c:5j^-r, J¥^WTa®CTa^iJ^cnc:@aBl^M>/ l-^l ©Mi^^^tzT^-5. /i?H ^ t /; 1954, '56^®2@®J¥^?^^'aiC^^-r, '/fei33<h^^ /f^7 1- ^ Sch 1- ^ 3®^^§1^^^-^^^^^1@ff^t||^L^. ^^^-:t(i-f ^7 1- ^ 3<h^y'~. 1- ^ 3rI^^X%^^^> B^^-'/&^®/6^Jll7J^^®^L^^?I^^®Ni:t!l^^ t)^a-L/c fc®®i=l--i ic^ ^7 ^ 3^-i-y" 1- ^ 3®<nm^ ^^iJ?L^^t/\ Ng®^rs]^]^^T/f@ft<^?^^li'^^sX ^ Hi^/c. d ^ b ©t?(A ^-T b fc, -^ ^ b ® M-^T ^—^® ^ fBl?Ji'J^%^ T <b ® ^ fc ^ < , <@{^ ^^^, -en-eni^u, '^©-^^^^^T^rBi^.^a^t'b®^^^^', °X^{A-^®^?N. m^^K^wnw L^c ^ ® ^ ^o/c. <i ©^^nm^s^n^ ^ ®®^(c ^ ^(D&^W ^T(t^-@®^?t^ b^/;. C^b®^^ ^iiti.C®^ L < N@®4'miy^^t<1:®^^^ ^® ^S^b, C^Cst^-^^W&^W^k- ^®^X^ Tables 6-7 {C^T.. Table 6 ^iSliW^-r.^^W L^im, g^Mt' L < ^kW^tW^^:^&^. ^^^t^(cr&ls®%j!iL^F)--ify3^@'?®^^^^^^-gciL^ fc®^, ^-^©-Xc^'fnIfL^, -?:. u b ^-?g^i^ u/; ^ ®® 100 ^^%;b L^. ^ ^ b® -5 ^i?tH» No. 1, 2 ©N'flg^d, -e®m^, t^X^B^^i^t- ^ 3^7&{^-t^<fc®^', /ftk®^slA l- ^ 3^-{•c:,& i^^'^o/c. L ^L, iW^WM^^^m^(Dt?^^>^, m-X, tMM^^L6-<^7 1- ^ 3?_l^^o^. —370—.
(10) Table 6. Mesuring results of each part of the body ; body height, head length, trunk length, tail length, length and height of caudal peduncle, length of central dorsal spine, pelvic spine and pelvic. girdle (A) and (B), number of dorsal spines and scutes, and types of scute-row, in 13 individuals of the intermediate form of P. sinensis and P. pnngiti'us. The length and height parfcs are shown by. % of length of mesuring part/body length, and the parts of number are shown by practical number. And in this table the mesuring results of Ikeda in addison to one of the present auther are entered.. No.. Body Head Frank Tail Sex 1 eight ength ength ength. Length of. Heigtn of. Length of. central caudal caudal dorsal peduncle peduncle spine. spine. Length of pelvic girdle "(A). Length of p el vie. Length Number of of pelvic dorsal girdle spines '(B). Number of scutes. Type of. scute-. row. 1. ?. 18.5. 29.5. 32.0. 38.5. 11.2. 3.02. 5.7. 11.7. 18.0. 10.1. 10. left 33 right 33. A A. 2. ?. 17.5. 30.0. 32.0. 38.0. 11.8. 2.9. 5.9. 11.3. 15.5. 11.5. 9. left 34 right 33. A. 3. ?. 16.4. 27.5. 33.2. 38.4. 10.5. 2.8. 6.4. 11.8. 17.4. 11.0. 10. left 44-2+16 right 4+14-16. F F. 4. 6. 17.7. 30.2. 30.8. 39.0. 11.5. 3.7. 5.2. 9.1. 15.4. 11.5. 10. left 34-15. G. 5. ?. 17.2. 29.5. 33.5. 37.0. 9.5. 2.6. 5.9. 11.4. 18.0. 9.2. 10. left 5+16 right 5+16. 6. s. 16.5. 29.8. 33.2. 27.2. 9.9. 2.8. 5.4. 10.3. 17.9. 10.4. 11. left 4+15 right 44-15. 7. s. 18.1. 30.2. 32.0. 37.8. 10.1. 3.0. 5.8. 10.7. 16.5. 11.3. 10. left 4+17 right 44-17. G G G G G G. right 4+1+15. E. F. Colleeted region. Investigator. Akkeshi. Kobayasi. Akkeshi. Kobayasi. Akkeshi. Kobayasi. Akkeshi. Kobayasi. Akkeshi. Kobayasi. Akkeshi. Kobayasi. Akkeshi. Kobayasi. 10. 5+9+5. D. Akkeshi. Ikeda. 10. 34. E. Akkeshi. Ikeda. 9. left 34. A F. Kurile Islands. Ikeda. right 1+18 left 34. right 7-4-20. D. A. Kurile Islands. Ikeda. 9. left 34 right 04-30. A B. Kurile Islands. Iked a. 9. left 3-1-16. F. Kurile Islands. Ikeda. 9. right ?. 0. ^.
(11) Table 7. Mesuring results of each part of the body, body height, head length, trunk length, tail length, length and height of caudal peduncle, length of central dorsal spine, pelvic spine and pelvic girdle (A) and (B), number of dorsal spines and scales, and types of scute-row, in 10 individuals of the intermediate form of P. tymensis and P. pungitius. The length and height parts are shown by % of length of mesuring part/body length, and the parts of number are shown by practical number.. No.. Body Head Trunk Tail Sex height length length length. Length Length Height of of of central caudal caudal dorsal peduncle pe dun de spine. Length of pelvic. Length of pelvic girdle. spine. '(A). Length Number Number Number of of of of pelvic Soft ray; Soft rays dorsal girdle ?f dorsa of anal spines '(B) fin fin. Number of. Type of. scutes. se.ute-row. 1. s. 21.7. 32.6. 30.6. 36.8. 9.7. 3.3. 4.7. 9.7. 11.3. 10.8. 8. 10. 9. left 1+7. right 2+7. I I. 2. s. 20.0. 31.1. 33.3. 35.6. 10.0. 3.4. 5.1. 8.2. 10.0. 10.0. 9. 11. 9. left 04-7 right 0+7. J J. 3. s. 19.1. 33.0. 30.9. 36.1. 10.0. 4.3. 4.9. 8.5. 12.5. 9.2. 9. 10. 9. 4. s. 19.1. 29.6. 32.4. 38.8. 11.0. 4.1. 4.7. 7.4. 13.0. 9.4. 9. 11. 10. 5. 3. 20.4. 32.9. 32.9. 34.0. 8.1. 4.4. 4.9. 9.6. 13.8. 9.8. 9. 11. 9. 6. $. 19.5. 32.3. 34.1. 34.1. 8.3. 3.6. 5.5. 8.8. 14.7. 10.2. 9. 10. 7. $. 18.2. 29.0. 30.3. 39.5. 10.1. 4.4. 3.85. 8.7. 9.6. 6.7. 11. 8. ?. 19.7. 32.5. 32.5. 35.0. 8.8. 3.5. 5.8. 8.2. 8.9. 5.7. 9. ?. 18.7. 31.2. 33.4. 35.5. 10.0. 4.0. 4.5. 7.8. 9.2. 10. ?. 19.6. 30.0. 35.7. 33.5. -9.0. 4.7. 4.8. 9.5. 9.8. right 04-7. J J. left 0+8 right 0+8. J I. left 2+8. I I. 9. left 0+7 right 1+6. J. 11. 9. left 04-7 right 0+-7. J J. 9. 10. 11. left 0+7. J. 4.5. 13. 12. 9. left 1+8 right 1+8. I. 4.8. 11. 10. 9. left 04-8 right 04-9. left 0-1-7. right 2+8. right 0+6. I. J I. J. J. ^.
(12) h?3 No. 3 ^ ^ No, 7 ^i, -fn?<fc-j^^:7 1- ^. 3^0'flI-N^ab-^^, No. 3 h No. 4 -f@?(l. b b ® ^ ^ {?: M ^ 1-2. -®®m;m^L-rj^. c(Dm^Mw^ M>[^y \ ^ 3{a»a6b^j:i^®'{to® ^/c. <:®/ffeCflb®5-f@-fr(t, t^XK^®^. ^{cjg^T^mffix, gM?, g^^p^®^^ ^, i- ^ a^oft^^m^^^^LT^^yi W^ b n/c. t k, Table 7 {j; ^A^Lfe zLKm^^tf^^w L/c ^ ®® 43 ct ]o '^ m^ ^ ^ ^m(D^m^m^ -f ^ ^ ©f L ^ 10 -f@. ft(c:^ ^^.^(Dm&^w ^ t^^', ^ ^®S^/f?!?X^L^<b®© 100^ ^^%^L/C. —B^-< ^7 1- ^ 3^^y' 1~ ^. 3 b SW^W -^ 7 ]- $ 3 <^ 1- $ 3 ®^-^^J-bL, f^§-^{cgfc^T?it^fflB?%??. ^©^ab^', Table? ^a-^W?^. T(^<^'b(fb^, -^(DW^^^.'O, Wm ^y'1'^3^, ^^(^-f ^7 I- ^ 3^^^LT. Fig. 3. Natural intermediate fishes of the both kinds of P. tymensis and P. piingitms.. \tL. dtLb©rtXg?®^®^?C®e^T. a. P. tymensis b. Intermediate form c. Intermediate form d. P. pungitws. A^^-f ^7 ^ ?. 3R:,fi;l^^;bn^>??W. ^ No. 1, 3, 7 {Cifc^T^^C, No.lo^^^^y" 1- ^ 3^, tMX, BMMX^^Fs^^r^ ^,. No.3®^, MW, S^X^^^^^^^^LT^'9, No.7®tMX^^'tM ^WB^b^R:^ 1- ^ 3S1^^^, ^'^^, ^-iS^^^M^^^^^^-^LT^ k. t^cfib^M^> gJ, ffi, M®;§-^^®?^<^?!^6<]j-y" 1- ^3{!c:,fi:i\ No.6, 10 {cjt^ T-fc No.6{c^U-;5U?, g^'^^^BJ3b^^-^ ^7 ^ ^ 3mi^, ^itj, mffi-X, @ffi%. ^N@®4]f»^L, No.lO{cX^T^, f-^^, J^ffiX. Mffi^, tMM, fiM%M(-tN@®4i FB^^LTI^. tt^, <^tLbW®No.2, 4, 5, 8, 9^^-fBT^^fro-^©MiJ^-N^%/^^N^. ©4^^, ^n^m^Wir^^Wt^^^W^^W^^^^- ^®fcibFlut5®$P<, -•& s®^f^®43rB^'f@fr^^^^^^ bn^>»^^ ^ R:M^T?T^. 3 OT(D$^^ Q^L^c ^ (D?i$lN(.l|£G^r 3@^© 1- ^ 3®]E^N^.^^^^^^l;^ffi}^ Table 2 ^^-i-$n< ^^^^^^^-3^^ <mb^m?ffl®f?S™NW, j^x^-^^y^^^TSsf^, ^^(C^-M-^^^^^)^^ (c:^^^IJ?g-^L^^^®^, M^CML^R:^ <> ^V" I- ?. 3° x/f^7 1. $ 3$}CT40 m, A ^y \ ^ 39 x ^ y" i- ^ 3 $ R:T 15 -fg-^, 1- ^ 3 ? x ^ 7 I- ^ 3 8 (CT 12 -fg-f^, ^ ^ 7l'^3?Xl-^3$ i.CT 14 /f@{^, 1- $ 3? X ^ y' 1- ^ 3 $ i'CT 14 /f@-^, ^- y' ]- ^ 3 ° X 1- ^3. $ }CT35W^-^c. ^£-^^W^-®^I&^^^S^'^, —^^^^{c^^^a^T^^^tl. /5> ??a7^^±»L^'»^^)T, -^^|S®HI?XO';'^lS?M?^fJ-^^. -C-fib®^!1 ^f^LT TablesS, 9, 10, 11 }C^T. d^b®^, Tables 8, 9, 10 i^gfi^ ^/;^?^> 'fnl^^. i?W®¥;^^{?W®X^^^ ^©^, {"Ttz^frX^I^L^ fc®^-100^^^-%^ 373—.
(13) ^ ^ Table 8.. 5^. Comparison of each mesuring part of the bodys; body height, head length, trunk length, tail length, among the 6 kinds of hybrids. And the mesuring parts are shown by mean value ± standard. deviatio of length of mesuring part/body length, in the mesurd individuals of the hybrid in each combination. (%) Measured. body length (cm). Cross. Measured number. P. t.9 xP.p.S. 50. P. p.? XP. t.S. 30. 3.0-5.7. P. 8.? X P. p. 6. 25. 2.8-5.1. P.p.9 xP.s.s. 25. 2.9-5.6. P. s.? XP.t.S. 30. P. t.9 XP.S.&. 40. Body height. Head length. Trunk length. Tail length. M±<r. M±a. M±tf. M±C7. 19.14:0.92. 31.5±1.14. 33.0±1.36. 35.5±1.16. 19.5 ±1.40. 31.1±1.31. 33.5 ±1.36. 35.4±1.23. 18.1 ±0.91. 30.5±1.17. 31.2±1.44. 38.2±0.92. 18.2 ±1.07. 29.7 ±1.15. 30.6 ±1.13. 40.1 ±0.98. 3.0-5.0. 18.8±1.01. 31.8 ±0.98. 32.6±1.33. 35.8 ±1.54. 3.0-5.8. 18.9±0.88. 32.5 ±1.25. 31.8±1.21. 35.7 ±1.22. 3.0-5.3. L/;. tk^r Table (Cffi^ ^^{g^^^'^-M^^-^ic:^ ^ ^^ ^W, ^-a^^-^(c:ck (.9, 7 fy^l^. ±/±^L^-f@-^^^^M^^o/;^7?^t), M(C^-Table {Cj^, MiJ^<@ft^(C^|&®^^) b^ ^<:d, M&^^^19, ^-f^®^:^^^—^K±{CU^i'J-/{Z(f^:b^^^/;^^^^. ^&^^:F,(DW, @S, BIS, mM®l?^M^>R:, TableS {C^T$H<, J-V" 1- ^3<1:. -f^7 1- ^ 3®lE^Nffi^^^^ ^) ^i {t{5^-^Ll^?^^L, NSIX®4]ilS^7&^?^^%^b ±. ^^y\^^^[^ 3®lE^Nffi^^-^^g£l^-t{l, N@tX®^P?'mLT^/c^ra •^^^^lmT^WBlt^-3^^, {•^39X^71-^3$®^ d^-^^ ^7 f- ^ 3. }^&<, '^^7l'^39x^36®F, ^M^®M^K^M1 1- ^ 3(c:^^-f^/;J:b^^%^t^® ~^&^. t/-; l- ^ 3^^-y" ]• ^ 3®:£^^I^^-t±(C®£l^T, NM-^^-^A^^^^ ^i (liiiS-^L ^'N^L, NIM®':t:lfES®W(c%;b^i/;. M(<:<i^^, ^^7 1- ^ 3<J:^-y" 1- ^ 3®JE^a^^ ^(D'^^bW-tM, -fW-fg^C^.g^WJ^btZ, -f^7 i- ^ 3<1: 1- ^ 3®^?;^lt^. F, ^^{cfc^flSLTt/^y^^-^^. MffiS, @ffi^, tMM. BM^M, .®'lf^^g£^T{A Table 9 (^T$n <, -^^7 F 5. a^^-y'l- ^ a, t- ^ 3 ^ j- y" i ^ 3 ©iE^Nffi^^-^®^- ^ ^ ^Mffi Table 9. Comparison of each mesuring part of the bodys; ]ength and height of caudal peduncle, length of central dorsal spine, pelvic spine,. pelvic girdle (A) and (B), among the 6 kinds of hybrids. And the mesuring parts are shown by mean value ± standard deviation of. length of mesuring part/body lenth. in the mesurd individuals of the hybridts in each combination. (?»). Cross. Measured Measured i body. number | length (cm). Heigth ^Length of central of. I Length of. caudal peduncle. caudal peduncle. dorsal spine. M±a. M±<r. M±a. Length of pelvic spine M±</. Length of|Length of pelvic | pelvic girdle | girdle. '(A) | -(B). M±a. M±cr. P. t.9 xP.p.S. 30. 4.3-5.3. 8.3±0.87|. 3.6 ± 0.39| 5.05 ± 0.47| 9.6 ±0.79| 14.8±1.52| 8.05 ±0.52. P. p.? XP. t.S. 25. 4.1-5.7. 7.3 ± 0.89|. 3.8 ± 0.41| 5.0 ±0.58| 8.7 ±0.92| 14.5±0.92| 7.7 ±0.78. P. s.? XP.p.S. 20. 3.9-5.1. 10.4 ± 0.59|. 3.0±0.22| 6.05±0.25|11.1 ±0.68| 16.9±0.81|11L.l ±0.76. P.p.9 XP. s.S. 20. 4.0-5.6. 10.6±0.51|. 3.1±0.35| 6.0 ±0.27|10.8 ±0.57| 16.4±0.80|11L.O ±0.63. P. s.? XP. t.S. 20. 3.9-5.0. 8.5 ± 0.62|. 3.4±0.23| 5.5 ±0.50]10.5 ±0.86| 15.2±1.06] 9.0 ±0.67. P. t.9 xP.s.S. 25. 4.2-5.8. 8.3 ±1.1. 3.5 ± 0.32| 5.3 ±0.56|10.05 ±0.78| 15.3±1.15] 8.8 ±0.83 374—.
(14) 3 gh ? 3®^t^^. ^®^tNiM®43^^L-C(/^^, A ^^ \ \ ^h I- ^ 3®iE^Nffi^3-^® FI (t, -f^7l-^3^, ^(^. (A-C-tzJ^^^fi^^oTl^. ^©MBMCT^ ® <^ ^. 7s ^ ^0, fsb 6^^^rtr"iw{<: ^^mi(Dfm^. IW^?X!M^^-^^i?^ ^ ^ ^ ^^. b ^. S. tMMWmXU^M-^^ ^ ^ b^Fi <h t),. {t^MA/(/i. ^U^Ld^. N?®43^^-?^R:%;MV;:. ^®»^^t^,. M?'f@'N^(c^-M^®)^M^®^^ l ^^f®^ja^S^ t^ttiT@/;:^7Wj:^^^b-tz^. wm^m. (AA^. W/&^t ^ ® <h LT, .£ y- 1- ^ 3 ^ -^ 7 1- ^ 3 ®IE^. NM^-Mtc:, tM®M^X^2@iH^ft-3W,K^ m^k- ^ ^ b ®%/^ Z^M%^iJ4] i!c:@M® 2,3 /t^? t^®^, MiM-f ^ 7 1- ^ 3^^^M(i^y i- $ 3^ -z?^o^. cn^&^^(D^ [ ^ 3^y" l- ?. 3®IE^N. Fig» 4. Dorsal spines possessed the characters of both parent kinds. a. b. P. tymensis X P. pitngit'lus c. P. tymensis x P. sinensis. a-^o^-® F, ^ }c ^m^) b n^^, %x®^'^.'/ 1- $ 3 ^ -< ^7 i-?. 3 Fi ®^:t^^ ^ -fc ©^ ^o/;. ii-c^ ^^ 1- ^ 3 9 x J. y" ]- ^ 3 $ ^, ^-y't- ^ 3° x/f^7 l- $ 3$ ®iE}f|tiga^^. ^^^ ^T^ ^ ©rs^c^i^T^, axM^ic^^^b, ^a^^-t^^i (ttuM^^^-^i ^ ^x^: ^U^:}^?X/ft(/^. MW^^-^^, W(A)^, ^^-^(.c^^'t^^^^^^^^ ®S^^, ^-a^o^^ ^^NaS®^^^^-^^^, ^^(B)^, gpiS^^tt^^^OM^^ ^F, b ^NI?®41rs^i9A>3^M^c^-3^. L^L, Fi (C^y-^c®^®^^^ Fig. 5 }c:^T. $p<@^®^c:%;?, .^W®^, NM^®^Si, ^^^^^^^-^(I, -^(A^N^§®43^ m^^t^^t^-t^^^^b^. t^-f^;7 I- ^ 3^ 1- ^ 3{C^!d-^)M^^®^^^,. IWfi^^^', c®^^|^N@<^-I-y" 1- $ 3^®a^^^^^^D^^ ®^{cM^<. Fig. 5. Pelvic girdle of some hybrids and the 3 kinds of sticklebaks. 2. a. P. tymensis. b. P. pzcngitius. -375—. c. P. sinensis d. e. f. Hybrids..
(15) ^ ft. 5^. B as.. D. E •. ^OO^OV$OOOSC50^$%E2X£&?3^^S. Fig. 6. Types of the scute-row of the 3 kinds of sticklebacks and their hybrids. A type-•-the type has a complete scute-row. B type-•• the type has a scute-row lacking a few scutes at the former thorax. C type-•-the type has a scute-row lacking some few scutes at the middle abdomen. D type-'-the type has a scute-row lacking some few scutes at both parts of the thorax and abdomen. E type-"the type has a scute-row of complete number constituted by the small scutes. F type-"the type has scute-rows at the former thorax and tail, and a few scutes scattered at the abdomen. G type-•-the type has scute-rows at the former thorax and tail. H type-•-the type has scute-rows at the former thorax and caudal peduncle, and a few scutes scattered at the abdomen. I type-'-the type has seute-rows at the former thorax and caudal peduncle. J type-•-the type has only a scute-row at the caudal peduncle. K type-•-the type has only a scute-row at the former thorax or a part of the abdomen. L type ••• the type lacks completely a scute-row.. — 376 -.
(16) 3 g t- ? 3 COWM Table 10. Comparison of each mesuring part of the body; number of dorsal spines, soft reys of dorsal fin and anal fin, among the 6 kinds of hybrids. And the mesuring parts are shown by mean value ± standerd deviation of the mesuring number, the mesuring number from minimum to maximum and their mode, in the mesurd individuals. of the hybrids in each combination. Number of soft reys of dorsal fin. Number of dorsal. Measured. Measured number. spines. body length. Mode ( ). M±a. (cm). Mode ( ). M±a. Number of soft reys of anal fin M±(T. Mode ( ). P. <.? xP.p.S. 50. 3.0-5.3. 10.05 ± 0.79| 9, (10), 11. 10.8±0.45|. 10, (11), 12. P. p. 9 XP. t.S. 38. 2.3-5.7. 9.8 ±0.63| 9, (10), 11. 11.1 ± 0.68|. 10, (11), 12 10.0±0.58|. 9, (10), 11. P. s.9 XP.p.S. 32. 2.5-5.1. 9.2 ±0.80; 8, (9), 10,11 11.2±0.46|. (11), 12 10.0±0.66|. 8, 9, (10), 11. P.p.9 XP. s.$. 34. 2.6-5.6. 8.8 ±0.64| 8, (9). 11.3±0.48|. (11), 12. 9.9 ±0.47]. 9, (10), 11. P. s.9 XP. t.S. 33. 2.3-5.0. 10.8±0.78|. 10, (11), 12. 9.8±0.31|. 9, (10), 11. P. t.9 xP. s.$. 50. 2.7-5.8. 10.6±0.86| 9,10, (11), 12. 9.7 ± 0.59|. 8, 9, (10), 11. 10.5 ±0.66] 9, (10), 11 9.7 ±0.72| 8, 9, (10), 11. 9.4 ± 0.60| 8, ( 9),10,11. %t^>»^^ Table 9 ^ ^ ^-^c. tM^(A Table 10 }c:g5L^n< , -f ^7 1- ?. 3 ^ ^/ 1- ^ 3 ©IE^NM^^-^J; 0 ^feD/c ^ © mode (^ 10 ^, -^ ^7 1- ^ 3 ^ t- ^ 3 ^ ^ ^, IE^^ ^ mode (t9^:, ]> ^ 3 a: ^y ]- ^ sj;^)^, IE^^ mode ^ 10 ^-C:^^, M^^ mode (^N^S®^ mm^ tS. L T@/;. t, ^ Table 10 {c-^gfi L /; ^', c^ b d 3 @%® W^ ch (t ^ ^. =?^^, ?Wc:^T{t^BJ^mL/;. ^M^^^cfc^^l;/; Fi ®rrfX®^fi§^tS^T^ m^^^. -€®^ Table 11 ^-§^^C^, ^®$tJ<^%L^. A §U • S^J^J; ^Mffi^(c;ig^ LTX<-ML^m^^T^tb®. B^: S^-^tt)€®^'f@®^®^^X< ^®. C^: m^&^m^^^^, ^j^®im®m^< ^®. DSI: ?^^x<$i^b^-^-f:x^^^^, i?Bu:^mr^^Br/j®Mi®im^< ^®. Esy: .i^L/c^x^^o^, ^-^xj4-®^f ;t^©. F^: fi^^f|fi^7^M-^?^K^i'c:^tL^ l~2{@®r^X^;i'^^^©. G^: S^^tff^ Table 11. Comparison of the types of scute-row and modes of number of scutes among the 6 kinds of hybrids. The types of scute-row are classified in 12 ranks from A to L, as shown in fig. 6, Typts of scute-row. Cross. A. P.t.? XP.p.S. left right. P. p.? XP.t.6. left right. P.s.? XP.t. S. left right. 4 2. P. p.? XP.s.S. left right. 1 2. P.s.? xP.t.S. left right. 5 6. left. 9. ?.(.? XP.s. S. right. B. G D E F G H. I. J. K L. 42. 7. 8. 1 1. 9 17 11. 1. 41. 1. 9. 1 3. 2. 2. 1 2. 5 3. 9. 8. 4. 1. 2. 3 6. 8 5. 2. 2. 7 8. 1 1. 2. 4. 1. 6. 1. 11. 11. 2. 6 5. 2 5. 21. 2. 4 5. 4 2. 1. 1. 1 2. 2 1. 6. 3 3. 3. 3. 6. 377—. 7. 1. 1. 2. symmetrical. AIeasured number. 3+8. 21. 50. 2+6 1+6. 10. 30. 54-16. 6+15. 6. 25. 5+15 5+1+14. 8. 25. 34 (E) 34 (E). 16. 25. 34 (A) 34 (A). 18. 50. 24-8. 3. 1. Number of. Mode. individual.
(17) ^. ^' -?. ^M^tcm-B^t-^^^®. 'H.m: ^^ftti^<l:m'R^^M^i~2<@®^tr^^-^t^^®. iiii: BWfT^mCTR:iw;&irT^®. ]m: Mffi^®^^m^^e^^^®. 'K^I: fi&3f^ir^g m, ^?mu^R:®^,m^^--5T^®. Lm: ^{c,W^^Dt'^®{^IJ^>. Aj;^L. ^(Di2mm^m^mb, ^-a^o-^R:, &^(Dwmc^^^, ^mmc^^i-^w^ g5AL/c. ,^^§IJ{C®fc^T/4M(C Fig.6 ^^l^T^B^IglML/c. t'/'c, mode (C^l^T, ^J^{f. 3+8 ^g5L^^®^, + j^m®i^»^®,ms^%;bb, + ^^^^©^^^ss^® im^:?L^®r, M^5+1+14®$D< + ©FaKc^to^^^^, g(<il^m^<J:@B^-{;M<J: ®r?;^T, ^^b^^tLT^T^,^4X^^^L^^®^^^. Iffi^itt^-r, ^-ffi^^^cfc^l^^U;, 'f"r^^N^S®^X^®iH®^-@®^Sl^^n-€:'^%LT@^. L^L-f^7 1- ^ 3 <^ y" ^ ^3 {c^i^-r^jE^iiigM^^-^^ ^ -f. ^y \ ^ 3K:?&l^^T^ ^®^< , A ^7 ^ ^ 3<^ 1- ^ 3{c^^T^iE^i^M^^-^<1: <b-^ ^y \ ^. 3 (C?^^,&l^^%^T/f@^^'^ < g^ b^/c. t ^•<^7l'$3^1-^3, ^-y"{-$3<l:P^3. ^^^>o^, mw^±<m^^^^\. ^Mffii$rrfx®rs^c^{i®.^^^x^^oi@-ft^^g^ -M;bfi/;. M^, Z®M^^-^^^ fc£^®l-?. ;6^^M-fN®M-m?f;X;&^ < -fc ® ^ 1-2 <@?% •hU, ^'/ t- ^ a 9 x t- ^ 3 $ ^ <9 (t, ^{i©^X^. Fig. 7. Gaudal peduncle lacking the scute-row in the hybrid from P.. ± < ^PT^) <b ©^ 1 -fW^fz/-;. c n b ®?^§u ^t/c, Nt?<:(t^< J^^b^^^^y-c^®-^^^.. tymensis ? XP. piwgitms 6.. y±®^|i:^^^^^M^^-ij-^^i'j-^^.^^®^.^{®{c:®£l^T^./c^®^:, ^tiC^M^3. •^^ ^ ^ I; ^<W^-f@^ icXx 0 ±i/j;', ^^^-®^d6^f-^^^M^ L-c^^^', c ^ b (ANtg§©^ ^©tmc^T, ®^T^-@®^5H^w^®7?, ^-g^^^^€'nN^@®4]Fia^i^^ir^® <fc<9, -^©tJl^tS^-c^X^-t-^ fe®{c:^J^^^®^}c^F^^lS^i])g^^oTjg^. ^a^/-b^ M^c^^n^i^^, 'f"T^^S^{c,&i^f3B^^^T^ ^®®$R<-^^^^, g?^^;7]^3 {cjibt-i^&^:}to!fc®(tg^bfi^-3^. @ftg®^Jtel^Ttb^-?.^;b^^Q^^ Pi. {i'Mn<bNiffi®r|:'fM^%;bt$p<^^~, mwc^wwm^^m^^-c^-my)^^. ^m^6 e> ?. .^-a^o^ ^ ^ ^ b/c Fi ®mt®^^ Table 2 {c^T$0 < ^o/c^, 1 ^^ ^(DW^^-c^^m^ ^mm^m!-t^^-^ < sn-^ ^), ^"igNa^ge^-c:gi-<o ^, ?/?7~8 ^^®-f@{^T^^lM^:^-^{c--pfJ^jL^^'f@{^(Af£^^^o/;. €/}zicRL, 1^^±^aTiLy-cWt.ic^o-at, ;f"f^®M^^^-f@{^^^^-rifcs^rX^F^'%,i^-^ urjg/c. /f ^;7 ^ $ 3 ? X ]- ^3 $ ^(3/±^/CW^C|3, d^l^^^^o^ l/f@^^^^a^-^-3^rBT?^-%HL^.. <:®'fW®m:tfi(A£;&®N-?J^ bti, ^tm^{c:^gji^<bj|^(/j-^^^^®^^^®^^Ha^' ?LTt^. ikW\W(D^^(D^m^^^^, ^y- t. $ 3 ? x ^7 I- ^3 $~^(^, ,'i^ 19 -f?, 8 -f?, /f ^7 1- ?. 3 9 x. ^-yi-^3$^(t, M-fa2-f?, M^fe4'fi H., l-^39X-f-^;7l'$3$ -T?(t, MSfe. 7 -f@-Ns J7<-fa 3 -f@{^, ^ ^ 7 1- ^ 3 9 x 1- $ Kg. 8. Shortening form of the hybrid from P. 3 $ ^' ^^^ 2 W- ^^ 9 ®-^' ^ ^ tymensis 9 X P. sinensis S. 3 9 X^-V" 1- S. 3 ^"^{A, .il-fa 2 'fjgf^, ^U —378.
(18) 3g h ? 3 CQ^.^ 3 '[?, ^-fa 7 t?, ^y- 1. ^39x ]- ^3. (^^^^-WH^^ntc. MW4^c:^b/cB$M^ LTtt, -i-y'1-^ 3° x-f^7 l-^ 3$, 1'^ 39x^7 ]< $ 3 $ , ^ y- l. ^ 3 9 X ]• ^ 3 $ HJ; ^) ^ I;/; F, 1^ R:^- 1-2 -f@^, Fig. 8 {c:^^$tJif, •?M®S^- L < ^!@ b^W^m ^ b n /c. N^H'^ y' 1- ?. 3 0 X 1. $ 3 $ ®?»^ No. 526 -S-JIL^ (d, t^lj;£t, -^-M 41.5 mm, 'f^i^ 9.5 mm, ggM 15 mm, KM 13.5 mm, US 13 mm, t|?2.2 mm, SMM-X 4.7 mm, 1.8 mm, 1.5 mm, W (A) ^ 7.9 mm, )?(B)^. 5mm, JWmK^., ^tlb®te, -N^, mM^(c:? Lm^^y'gB^^S^ L < ^t^ tlT^S ^-, c®B®^^df^^®&^^^cl:^L^?ft^{JB^r^^. <i®^W?M®,LH%{c®fc^-c ^, -^(tij/J^^^O^t^t/^^^^^^^^O^d^^/O^if^b^^.. F, m^mw^r ^^ai^^^BML^^d, f"m®a^3^R:^T^, {5{5-^^^^o^. L^L, ,? ^%^®ST^/;^, /J^^^NW®^^{tB^b^^^^^', i?^@®^^%7?^^> 5^, 6^ (c:^ km^^m^t^wwM bW, 7-8 ^ ^^jorj;^ ^ < M-^ii^®j^i±i®.w: ^%^/}z/;. L^L, c©l^-%!{t^@^®®^,i^i-%<i: LTd^M^, 7j<^(A'tRt20°Cy±^^. ]o, ^-^t-f"m^7cx^^\ g^M^ij.gt^-fwu^^'M^v^o^. t/c^^a2^. ^^ L/ct^, S^WWfeflU^h L?g^, 6 n ^^^m^^mwwwm^nic.. ^- tl b ® F^, ^ V" t- ^ 3 ° X^^^ 1- $ 3 S ^QS' 1- ^ 3? X.^y* (- ^ 3 $ J;^ ^ D /; Fi tt^AX. M?LcV), W-^^^m^ ®r^AI»^T^^T^. L^LN^^-^^), ^^^(c^S W^'^<, ^^l^?-^feT^®{^ <, M-<T%%^SIJ^{C:H^L/C. c®]^B{c i^Td, ^?c%;btz^-jt'f5?%a-^t^<, %WR^-®W^-®^j^^, %^%rB3®^7k^.(<:j; 7sb^U^^W^. tk-^ ^^ t- ^ 3 9 X-^y- 1. ^ 3 $M^^-^© 1^^^^7j<^t^^fPia^. ^w^tz^m^-M^L-A:. wmm^ 7 j^^^^^n, ^®7j^?c{tf,?mi^;b-^®4/ti -?^ ts) 1f ^ ^T@ ^. ^P%^ -MM b /c R^ (1 %/^ Z ® ^ ^'X^P <1: ^ o-C@ ^ ^ C ® ]^ {<: 5-6 ^ © ^ -M^-^ML/^. ^^b®^-7-^-€©^i.c%^?g^j%i.c^^K^j^i6T^^. L^L, ^^b®. K^^^®±^-^^^»^T^^^jb-rLT^^L^. t^i^^,ML/;, ^-a^-^tt.® ^^I^L^^, •€ ^b(^ ^^{c:^,t L, ^£Wc:H;^a®^;Wb^T@^»^fl-^. ®^te®{;^T^%^®TIIX®^tlHcjiit ^^, l.W^^^i-M, Xm®ifc ^ ^^^. ^T^B;W^tz^b, ^RF, abS^^®^^"It^M(^^;bn^.. ]• $ ^JI,^^^^^W^^?W^.WWft^^, N!ES ('33, '35, '41) (^-f^7 i- ^3^ i- ^ 3®^^M<l:L-c, ^Hm®^t?jii, ^?n,a®j?^, -T-B®^^B^'J^^'?c:Bi. f^BX^^y l~^®?^^^y'T@^, d^b®rt, J?^, ^^B^iJ^^'S^gX^4?y l-^® m^^^r, NscDX^^ram^jiL/;. L^L, Nii:9^^^/%y;'^%a^^ji:fi^®'f@ ^-fnTfl<^^:7 I- ^ 3^^(A, 1-^ 30^-M^^^^^^^®^^^^ ^^IrraLT^^. ^^^l. ^i'<:yfeEg^ai:9t^®a'^JII^^^r, ^^7 I- ^3^: ]> ^ a^g-^X^cl^SH^L^Jl:, N^ ©^(c^^^i^^^^^Ifr^^^b^^^^^Lri^. MR:rfeBU;;?€®El;W?%?^ ^7 ]- ^ ^b-^-f \ \ 3®7i^^x?®^^T^»:^j:^fLTi^. ^^\^(Dm:s^^^mw ^^mcff.j^m^^y^^, -^^^ i- ^^^} ^ 3®^ram<l:©^z^m^%Mt-^<J:fi?if?^. ^:j[|7j<^®M2WX^%^^T, ^7 ^ ^ 3^-^y" 1- ^ ^o^WW^^W^m^ 379—.
(19) ^. Fig. 9. Photographs of the 3 kinds of sticklebacks and their hybrids. 1. P. sinensis 4. P. sinensis ? X P. tymensis S 7. P. tymensis 9 x P. sinensis S 2. P. pnngitius 5. P. pungitms ? ><P. sinensis 3 8. P. s'inensis ? xP. pungitius $ 3. P. tymensis 6. P. tymensis 9 x P. pzmgitius S 9. P. pz<,ngiti.us9 xP. tymensisS (the upper part-'-female: the lower part-'-male).
(20) h ? 3 CO^. HL/;. ttc^^dtL^om^W^, 2@^^®^,g^^®||^^^^^Q^. ^©^H^J; MW^^ k 2@%(t^< yg^X^J: L, ^?JU{c:^^Tjg,i6b^^$0^, ^^^^^-^IrT b^^W^^o/;. ^R:j^B®»^L;^, ^Uf^^^^^y-S^^^ 1- ^3cl:^y' ]- $ 3®f^[gB^®&^{t, ^-©.g^^tc-^-^Pi^^^^^b^-S^, ^<^^-3^Rt%^^^^ ^^> ^®^(t^ < ^Ti^@®rB^c{j;M:?MT^—B^%!^^t^y^s^6(l:^-^y;:. t/-c^^^ 1- ^3 ^t-^3, A ^v \ \ 3<J:^y i- $ 3®NM^^-^^^^T, A:C^t}c:J^lEM^I;&;f?3. L^I^K^^^, ^^(D^^^-m^^^mw^^w^c^^, ^©fft.^., uzb. ;&jMJc:^ij;t-^L^WHj^;:. cn^'sy^^t^^, §CT£j?c:^T, NSFsKc: 5i^^fr^^^^"Tt^?^^^^B%^U-^^®®$n<^^b^S. L^L^^^, ^@t.®®ffc ^(D^m, ^^a®?tM;fTm^ ^tt^^?C^J^,g®^ST{l®i6T^S^ <^®<l: ^^Tjg ^, g^ ±rr®2M;TO-^, (t/cLTC®^^1;Ma^^^^'?^^$^^.^^®^^^^^^{r®e^T, ^^WW^.^k. L^L, <m^H%-^{c:MT-S^, --^^^^^®^{c:^^^y^?g y-, w^^y^^ms. L^, NM^^-^^ ^ ^ ^^ Fi 'f@-?^ x^gi^^^ifr^ ®^^jAk^. ®®^i"u^^;t^T^/c. ^?wmW(D^m^^^^w^t^Q<^&^, <^b. ^^^Mt^^^n^w^, A ^^ \ \^h \ $3, -<^7 i-^ 3<l:^y l-^ 3®NM^^ •^^ ]o ^ i: ^ F, \m-^ ^ <0 S^tW Fable 12 ®$p < -^ ^. ^T -f ^ 7 1- ^ 3 ^ I- ^ 3 ©B^. ^{cgt^T^{c, /N^-^®J:W ^f^t^ <fc^X^^^^^@^^7&{^L, ^(cN@ffl^J®fii g^c]:^^mx^®e^-r{Afg^Tipao^<fe^»^^M&bfL^. i£^Na^-^ ^i ^M^^< ^'^ bti^?fa(t Gm^, c(Qm^Mw^Wt-Wj^&(Q-/t ^^ } ^ 3®^^^^^g^i6 b n^. i^^m^^f^w-^^^'h^ B, c, D,.E, F^®$p^^-@®,r^x^^^i6b^^.. ^WWm^M^^Jf.^, TB^BWJ, ^'}c:@^BX^°>/ l'^^@-?®M4X^f"T^ ^±gg F, ?^?^ L, ;-ft:i'crfeffl^'m3p! b L/^TB§aBX^^° ^ I- ^® ^^^ 1- ^. 3{C^i/J-^>, f^-^7j<43-^ ^^®^^^^<fc®^b^^^^^®}c:M^^fB^®^^^-^^^^ ^®^s^ii<l:M^LT^^^Rt-, -^®M-^;m-b/n^p<m^^. ^(c:, ^^71-^3^. ^ y i- $ a ® r^{? o i ^ T ^, Table 12 (C ^^-$p < , C ^ b ^i -f@fr(t @i6 T X ? ^ ^ Wt^ ^ ?&<, ^cF.W^ci,, X^^4]^3u'f@^®^(c^^yj:?9®^(c^M^^^^bn^. K±®^. mww^^'o^M, ^©a^-^^T^, ^?/m®;M®CT?^~a^ i^g^® ^IR: ^ ^;t ^ W^ ®^?;b ti ^.. ±m®X-M43!M^&Bm®^l^li^tb^, ^Hb 3 SIX® ]• ^ ^(D^mw^^^^. T^-S. 3@^®jE7l^^<l:.fc, ^t^%^B7i, ?<h^, fft.O^W^ll^^/^^M^E^^^ bT, ^SMm<s^^^^^k^ < t^ b/)z^>»g£?i®,t^.^^^®ltX /J^SS^-ft^ ^ ffi& ?®ai%^m'^ < w b^T, J^^L/C^^X^'^M^^-^^^® ^i ^f4b^^^> --&a^^. ^^ ]o^k F, (D^m^, N»®^®4:'F?^r, mw^m^^wm^wt^ ck^sf^LT, c^b 3@®|j'f^{^@^^{ct^^^^ < [?g-rb^y-c^®^(t^ <, ^l@i^®M. §^^t-€;)ZJ^ ^ItoS^iH^^ ^©^^^(/^^^^b^^. ^i'c:3@^®rB^®,£^|§^{c ^-c^tc, ^Wm, W^o^WBM, ^-^©j^^^H^^^, ^^b3@%ra] ^m^^i^^m^ ^ »^ ffi^ ^ ©-^ & ^ . ^OT <1 ^ b ©^-1 ^tlTl^ ^©^6 9 ^^^b^^>.. m^mw^^^ff^M^. C^b®rRl®(tM^^^, ^t. \ ^ 3®7l^<"?<^ <0, ^ iim®»^SLT, ^.mt^. ^i®^®-^, F, y^®^^^a®a^^^ ^^@6t)i^^®^@^^ ^ tir^WJST^ I^JIL ^^^^®^^^TC^. 381.
(21) Table 12. Comparison of each mesuring part of the body; body height, head length, trunk length, length and height of caudal peduncle, length of central dorsal spine, pelvic spine and pelvic girdle (A) and (B), number of dorsal spines and scutes, and types of scute-row, among the 16 individuals in both reciprocal hybrids of P. ti/mensis and P. pungitws, and P. sinensis and P. pi(,ngit,ws. The lenth and. height parts are shown by % of length of mesuring part/body length, and the parts of number are shown by practical number.. Measured body No.. Gross. Body I Head |Trunk| Tail Sex iheight!length|length|llength!. Length of caudal. Height of. caudal. peduncel|reduncle. Length of. central. dorsal spine. Length of pelvic spine. Length of pelvic girdle '(A). 18 ; P. (.9 X P. p. i. 19.2. 31.0. 34.0 . 35.0. 9.4. 3.8. 4.4. 8.5. 13.0. 25 ; P. t. ? X P. p.. 18.8. 31.0 ;. 35.0. 34.0. 8.2. 3.6. 4.3. 9.4. 13.2. 46. 20.5. P. (.? xP.p.;. 67. 18.5 \ 29.0. P. (.? xP.p.. 116. 33.0. 19.0. P. p.? XP. t.S. 32.5. 32.5. 34.5. 32.0. 39.0. 33.0. 34.5. 8.7 8.2 7.1. 4.2 3.5 3.9. 5.8 5.7 4.4. 10.0 8.8 8.6. 14.2 12.5 14.8. Length of pelvic girdle '(B). Number of dorsal spines. 10 7.2 10.3 7.7. 10. Number of scutes. left 04-8. right 2+7. left 2+8. right 4+8. 11. left 24-8 right 04-8. 10. left 3+7 right 3+7. 7.6. left 1+8 right 04-8. 121 ; p. p. 9 xP. t.S. 19.8. 30.0. 34.0 i 36.0. 6.2. 3.5. 4.8. 8.5. 14.0. 7.1. 10. left 0+6 right 0+-6. 125 ' p. p.? ><P. t.S. 18.5. 28.0. 36.0. 36,0. 9.2. 3.2. 5.4. 9.3. 15.3. 8.1. 10. left 0+6 right 0+6. 132 IP. p.? xP. t.S. 20.8. 32.8. 31.2 | 36.0. 9.0. 4.2. 5.0. 9.0. 15.6. 8.8. 10. left 0+6 right 1+7. 202 |p. s.? xP.p.S. 17.8. 29.5. 31.0. 39.5. 9.5. 3.0. 5.9. 11.6. 17.5. 11.7. 209 !P.s.?xP.p.3. 16.8. 29.0. 31.0. 40.0. no. 3.1. 6.3. 12.4. 17.3. 11.0. 210 p. .9. ? x P. p. 3. 18.3. 30.0. 32.0. 38.0. 8.5. 2.8. 6.1. 12.5. 18.0. 11.8. I. 217 ip. s.? xP.p.S. 18.0. 31.0. 31.0 ] 38.0. 9.8. 2.9. 5.8. 11.2. 16.5. 10.3. left 3+1+17. right 3+1+15. 10. left 5+15. right 6+15. legt 11+20. right 11+1+201. left 04-31 right 34 left 33 right 8+24-20. 307 . p. p.? xP. s.$. ?. 19.8. 29.5. 30.5 ; 40.0. 9.9. 3.1. 6.1. 11.3. 16.5. 9.8. 312 I P. p.? xP. s.S. 3. 17.4. 28.5. 31.5 i 40.0. 10.2. 3.0. 6.4. 11.6. 17.2. 11.4. 9.5. 2.8. 6.0. 10.8. 17.3. 10.0. left 5+15 right 64-15. 10.2. 2.6. 6.3. 10.5. 17.0. 11.0. left 64-14 right 64-14. I. 320 P. p.? xP. s.S. 9. 18.8. 29.5. i 31.0 i 39.5. 323 I P. p.? XP. s.S. I$. 16.8. 28.5. 30.5 ! 41.0. left 8-4-15. right 6+1+13. Type of scute-. row. J I I I I. J I. I. I. J J J J J J. I F F. G G c E. B A E. D G H G G G G.
(22) 3 g h ? 3 03$^. l) A^iKc^.i- ^3, ^y i- ^ 3, ^7 ^ $ 3®3@®© 1- ^ 3M^%^'^XT^>. C^. b®^, ^y I- ^ 3 <J: -f ^7 ^ ?. 3®rg^BJ:fi;W^:JIimR:, I- ^3^-f^^ ^ ^ 3®7l-^^B.. mwf^m^^^^^m^n^. ^-n^^m^^M^^^, ^-'/ i- $ a^^ i- ^ 3®^ fW^iW^^KK-^, I- ?. ^hA ^y \ \ -3(D^mmwj^m^^^^^m^^t^ en ^®^^^'f®-NS:®:&^®MiJ^Mll{<:®£^TjlS{c, ^-Zb®/fW®^^2@®^Si(!:N§®. TO^-^-efi^-^T^T^c. ^c^m^.wm^^t^im^^^^, ^y'l- ^3 bA^:7 \ \ 3?X^:fiM^%£U^S)T, N@%^^l—®^n®4'^^-^^yg^^X;£-T^(f^^. -^<, W^fW—W^^-C^tc. 2) ±sfi3 @ii® I- ?. 3 ®^^^^-^-3/;^j!i, -^©aM^m^®™^ < ^i-^E ©W^lHlb^o/;. ^LT, ^^^®^^ffl%^^i5{3-M^E^?HL^^-3^. ^^L/-;ff^ ^, ^ ® ^ 30 g tu^^ ^ s-^Mi® E ® ff-t <^ WtSi WS: t£ b -c i ^ ^, 30 5 J^^ ^ ^ ^ ^ ?^/^ CT^®M<^^, 5<:^®^^<fcBS%L^l;^/c. ^©^{cLTff^^-Mtz^X^F^^S^^L irl^l^^-o^. t/;ctLb®t.^?M^^t^ii)%^'^^^LT^^. 3) ^a^^^^^^d:}^/-; ^ ©-^©^^^^N^L^. ^-a^o^w^Kim, NM® ^CT^btl, t/c.TE^Na^-^^® Pi ®^.^{t^%/^Z^|i^J^<6b^^^o^. L^L, ^ n^^W^^T^^^, ^^b^NtlOSO'-it^^^tZb®^^.®^^^^®^®^^-^^.® ^^-It^T^. t^X^4^m^®i?^f^, ^/tib Fi ®M^^^<1:?^LT^^> ^, ^^wwiw-^M^^i F, ®M^^^^®-f"TtL^®/fn'?®aij^^il^H{3'—^:T^ <^® ^^^^^^-3^.. 4) ^±®^^^-a^LT^^^^, C/)Z®3@^@^T.fi:^H^{C^^»^B^^^^. X ^^®43^^'f®'^®ffi%®iKS^®&^T^^-^^, 3@®^®^;n^®i;ja%®^R:j^9^T^ b-t^^]0^, \ ^ 3^ ^7 1- 5. 3W, J-y'^ 5. 3<l:'f^7 1-$ 3®^®g,tX^^&@T^. ^/^ ^c j^ (Sj(/^j)^^ < ^'i^a^.. Hubbs, C., 1959 : Population analysis of a hybrid swarm between Gambztsia affinis and G. heterochir. Evolution 13 : 236-246.. MiEB^P, 1933 : h f~ ^ ^ 0^-^^ €®^M. Ifi^, 45 : 141-173. -, 1935: :TWJ^OU^tfiO^. B^Mi^^, 5: 213-232. -, 1935: ^M®^@3^)S?®». TO, 3: 377-386. , 1936: ^®M?!im. Sl^t, 48: 179. -, 1941: ^jnw^Ko-^^o^. mm, 53: 120. Jordan, D. S. and E. C. Starks, 1902 : A review of the herd branchi ate fishes of Japan. Prsc. U. S. Nat. Mus. 26 : 57-73.. jn-N-mip • 1947: K ^ a ^^W^K^ <5W®CT%. M, 2 : 149-152. -, 1947 : K y a ^?4^®!G^^ SW®W% II. SS^t, 57: 63-64.. ? ^, 1956 : h V ^ ^ ^M o $IMM ^ ^Sl ^ ^ ^ ^@ o w^m. W, 26 : 258. — , 1957 : mioM^I§& U'^r 2, 3 O^fy^r.gt'^-C. jfc^^ffi^, ^2% 8: 44-51. - -—, 1957: ^mWV'^3 ^^^^O^^r^l-^. M, 6: 160-169. Kobayashi, J., 1932 : General observation of Japanese Gasterosteud fishes. Jour. Sci. Hiroshima Univ., Series B, Div. 1, 1 : 145-154..
(23) ^ ^ ^ Makino, S., 1934 : The chromosomes of the sticklebacks, Pn.ngitius tymens'is (NIKOLSKY) and P. pun-. giti-ns (LINNAEUS). Cytoloia 5 : 155-168.. MsTO, 1955: i^mwm^m'M i.. r^H^, 1950 : K -^ 3 ^ ch ^i> -^ -7 K '-? s r> ^ OIE^N^H, f^tclttt^^r^L^. 18^®^^-^%, 1: 7 -13.. -, 1951: -^-7 Ki> a ^W^Mb^-M. ^^, 1: 216-225. Minamori S., 1950 : Isolating mechnism in striated spinous loach, Cobitis taenia striata. Jour Sci. Hiroshima Univ. Ser. B, Div. 1, 11: 55-59.. , 1951: The lethal phenomena in the second generation of the spinous loach hybrid. Ibid. 12: 57-66.. , 1952: Pysiological isolation in Cobitidae. 1. Two races of the striated spinous loach with special reference to the difference in their embryonic respiration. Ibid. 13 : 199-212. , 1953 : Pysiological isolation in Cobitidae. 2. Inviability of hybrids between the mud loach and some local races of spinous loaches. Ibid. 14 : 125-149.. , 1957: Pysiological isolation in Gobitidae. 5. Sterility of the hybrids between the mud loach and six races of spinous loaches. Ibid. 17 : 55-64.. Okada Y. and H. Ikeda, 1938 : Contribution to the study of the freshwater fish fauna of Hokkaido, Japan. Se. Rap. T. B. D. Sect. B 3 : 133-162. Otaki C. T., 1907 : The stickl-backes of Japan. Trans. Sapporo Nat. Hist. Soc. 2 : 111-115.. Sadoglu P., 1958 : Mendelian inheritance in the hybrids between the mexican bling cave fishes and their overground ancestor. Zool. Anz. 21: 432-439. Swarup H., 1958 : Stages in the development of the stickleback Gasterosten,s aczileatzis (L.). J. Embryol. exp. Morph. 6: 373-383.. 384.
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