ce of the" annular ring". According his examination, exine of Sequoia sem- pervirens, Glyptostrobus pensilis and Taxodium distichum becomes thin in
thickness in the annular ring as in Cryptomeria japonica.
The present specimens are identified withSequoiapollenites gracilis Kru- tzsch from the early and middle Tertiary of middle Europe.
Botancial affinity: Taxodiaceae, Sequoia.
Sequoiapollenites sp. a PI. 56, fig. 2.
Description: Small conifer pollen grain with a short ligula. Outline subcir- cular to oval in lateral view. Exine very thin, finely punctate to weakly granu- late, with secondary folds. The grain possesses no annular ring which is provided with a short ligula (2. 5 /l.m long), which is laevigate and tapers off swiftly.
Measurements: 31 X 26 /l.m in diameter.
Occurrence: Uge Member, south of Kanuka (C 31).
Remarks: A single specimen was found. This grain has no annular ring ar- ound a short ligula.
Botancial affinity: Taxodiaceae, Metasequioa.
Sequoiapollenites sp. b PI. 56, fig. 5.
Description: Small conifer pollen grain with a long ligula. Outline somewhat oval in lateral view. Exine very thin, chagrenate, with secondary folds. The grain has a clear annular ring which is smooth in exine and the long ligula8
p,m long and4 p,m wide at base, somewhat punctate near base.
Measurements: 25 X 14p,m in diameter.
Occurrence: Uge Member, south of Kanuka (C 31).
Remarks: Only one specimen was observed. This is similar toSequoiapolle- nites megaligulus Krutzsch (1971, p. 222, pI. 73, figs. 25-35)from the Oligo- Miocene of Germany, but differs by its form of the ligula and structure of the exme.
Botancial affinity: Taxodiaceae, Sequoia.
Sequoiapollenites sp. c PI. 59, fig. 8.
Description: SEM photomicrograph. The grain is splitting. Exine finely uneven. A small ligula is situated on the middle of grain probably with an annular ring in polar view.
Measurements: 17.6 J.lm in diameter.
Occurrence: Uge Member, south of Kanuka (C31).
Remarks: Only one specimen was observed. This is a small form with a lig-' ula.
Botancial affinity: Taxodiaceae.
?Sequoiapollenites sp.
PI. 55, figs. lla - b.
Description: Outline rounded-triangular in polar view. The grain has a very small ligula-like process (2 J.lm long) (see fig. llb). Exine very thin, less than 0.5 J.lm thick, laevigate. No annular ring.
Measurements: 15 X 15 J.lm in diameter.
Occurrence: Uge Member, north of Uge station (A)(C 33).
Remarks: Whether this specimen is the genusSequoiapollenites or not, is not clear.
Botancial affinity: Unknown.
GenusVitreisporitesLeschik 1955 emend. Jansonius 1962 emend.
de Jersey 1964.
Type species: Vitreisporites signatus Leschik 1955.
Vitreisporites pallidu8 (Reissinger) Nilsson PI. 60, figs. 5-6; pI. 61, figs. 4-5.
1938 Pityosporites pallidus Reissinger. Palaeontographica, B, 84, p. 14.
1950 Pityosporites pallidus CReissinger) Reissinger, Palaeontographica, 90, p. 109, pI.
15, figs. 1- 5.
1957 Pityosporites pallidus Reissinger. Balme. CSIRO, Coal Res. Sec., no. 25, p. 36- 37, pI. 10, figs. 112, 113.
1958 Caytonipollenites pallidus CReissinger) Couper, Palaeontographica, B. 103, Lfg.
4-6, p. 150, pI. 26, figs. 7-8.
1958 Vitreisporites pallidus CReissinger) Nilsson, Lund Univ. Arsskrift, N. F., 2, p.
78-79, pI. 7, figs. 12-13.
1962 Vitreisporites pallidus CReissinger) Nilsson, Pocock, Palaeontographica, B, 111,
Lfg. 1-3, p. 58-59, pI. 9, figs. 134, 135.
1963 Vitreisporites pallidusCReissinger) Nilsson1958, Balme, Palaeontology, 6, pt. 1, p. 24-25, pI. 6, figs. 7.
1963 Vitreisporites pallidusCReissinger) Brenner, NDGMWR Bull., 27, p. 74, pI. 25, figs. 1-2.
1964 Vitreisporites pallidusCReissinger) Nilsson1958, Singh, Reaseach Council Alberta Bull., 15, p. 102-103. pI. 14, fig. 1.
1966 Vitreisporites pallidusCReissinger1938) Nilsson 1958. Burger, Groen & Zoon, p.
256, pI. 27, fig. 3.
1969 Vitreisporites pallidusCReissinger) Brenner, Habib, Micropaleontology, 15, no. 1, pI. 2, fig. 11.
1970 Vitreisporites pallidusCReissinger) Nilsson, Kemp, Palaeontographica, B, 131, p.
121, pI. 26, figs. 11-14.
1972 Vitreisporites pallidusCReissinger) Nilsson, Miki, Jour, Fac. Sci., Hokkaido Univ., Ser. IV, GeoI. &Mineral., 15, nos. 3-4, pp. 560-561, pI. 8, figs. 21-22.
1972 Vitreisporites pallidusCReissinger) Nilsson, Miki, Jour. GeoI. Soc. Japan, 78, no.
5, p. 244, pI. 2, fig. 13.
1973 Vitreisporites pallidusCReissinger) Nilsson, Miki, Jour. Geol. Soc. Japan, p. 208.
1973 Vitreisporites pallidusCReissinger) Nilsson, 1958, B. D. Tschudy, GooI, Surv. Prof.
Paper 770, pp. 18-19, pl. 8, figs. 1-2; tabI. 1, no. 53.
1975 Vitreisporites pallidusCReissinger) Nilsson1958, Srivastava, PaleobioI. Contin., 6, no. 2, p. 82, pI. 38, fig. 6.
1976 Vitreisporites pallidusCReissinger) Nilsson1958, Norvick&Burger, BMR Bull., 151, p. 140, pI. 28, figs. 6-7.
1980 Vitreisporites pallidusCReissinger) Nilsson, Burger, BMR Bull., 189, p. 63, pI.
17, fig. 11.
1988 Vitreisporites pallidusCReissinger) Nilsson, Takahashi, Bull. Fac. Lebral Arts, Nagasaki Univ., Nat. Sci., 28, no. 2, pp. 120-121, pI. 12, fig. 7.
Description: See Reissinger (1938) and Nilsson (1958).
Measurements: 18-34 /..tm in length (Reissinger, 1950); size range 20-38 /..tm (Couper, 1958); size range25-30 /..tm (Nilsson,1958); length of body12-(17)-
21/..tm, breadth of bladders6-(11)-13 j.lm, total breadth of grain 21-(30)-
42 j.lm, length of bladders9-(15)-19 1Jro, ratio length to breadth of body1.2- 0.7) - 2.6 (Pocock, 1962); grain size22 - 29 j.lm central body 10 -15 j.lm, air sacs 10 J..tm in height and10 -13 j.lm in width (Burger, 1966); overall breadth
21-(25)-33 j.lm, corpus breadth 7-(9)-10 j.lm, corpus length 10-(13)-15
j.lm, saccus breadth11-(3)-17 j.lm, saccus length 10-(4)-18 /..tm (Kemp, 1970); 25-40 j.lm in overall length (Miki, 1972);total breadth of grain 25-38
j.lm, length of bladders11-21 j.lm, breadth of bladders 7.5-16 /..tm, length of central body 14 - 21 j.lm, bread th of central body 7 - 16 j.lm (B. D. Tschudy,
1973); total width of grain 20-40 /.lm, breadth of central body 6-18 /.lm,
length of central body 12-25 p.m, breadth of bladders 7-18 /.lm, length of bladders 12-25 /.lm (Srivastava, 1975); 45 p.m in size (Norvick & Burger, 1976);
overall breadth of grain25 /.lm, breadth of central body 11. 6 /.lm, length of central body16 /.lm, breadth of bladders 10.4 Jim, length of bladders 16 Jim
(Takahashi, 1988); the present specimens: overall breadth of grain 38 - 49
/.lm, overall length of grain17 - 30 /.lm, breadth of central body 8 - 15 /.lm,
length of central body16.5 - 27 Jim, breadth of bladders15 - 20 /.lm, length of bladders 23-30 Jim.
Occurrence: Uge Member, south of Kanuka (C31), Uge (C34), and cliff near private house, north of Uge station (C 6).
Previous records: Lower Jurassic (Lias) (Reissinger, 1938); Eocene (Reissin- ger, 1950); Mesozoic, western Australia (Balme,1957); Middle Jurassic, York- shire'(England); Aptian, Isle of Wight (England) (Couper,1958); Lower Juras- sic (Lias), Sweden (Nilsson, 1958); Early Cretaceous, western Canada plains (Pocock;1962);Early Triassic, western Australia (Balme, 1958); Early Creta- ceous, Maryland (U.S.A) (Brenner,1963); Albian, east-central Alberta (Can- ada) (Singh, 1964); Upper Jurassic-middle Valanginian, the eastern Nerther- lands (Burger, 1966); middle Cretaceous, deep sea core, northeast of Cat Island, the Bahamas, West Indies (Habib, 1969); Aptian-Albian, southern England (Kemp,1970); Santonian-lower Campanian, Kuji (Japan) (Miki, 1972);
Coniacian, Futaba (Japan) (Miki, 1972); Cenomanian-Turonian, Hokkaido (Japan) (Miki,1973). upper Campanian, north-central Montana (U.S.A) (B.
D.Tschudy, 1973); Albian, Oklahoma (U.S.A) (Srivastava, 1975); Cenomanian, Bathurst Island, Northern Territory (Australia) (Norvick& Burger, 1976);
Lower Cretaceous, Queensland (Australia) (Burger, 1980); Coniacian, Futaba (J apan) (Takahashi, 1988).
Remarks: Vitreisporites pallidus (Reissinger) Nilsson is similar to Vitreispo- rites signatus Leschik from the upper Triassic of Switzerland, but differs in having no Y-mark on the proximal pole.
BotanciaI affinity: Caytoniales, Caytonanthus.
Gymnosperm-angiosperm incertae sedis Genus Clauatipollenites Couper1958.
Type species: Clauatipollenites hughesii Couper 1958.
Clauatipollenites uariabilis n. sp.
PI. 80, figs. 1-8; pI. 81, figs. 1-11;
pI. 82, figs. 1- 5.
Description: Monosulcate pollen grains. Outline circular to sub-circular or oval in polar and lateral view. Sulcus broad 3 -12 p,m wide, extending full length of grain, gaping often in its central region and tapering slightly towards extremities; its margin ragged, fringed with clavate ornaments (2-3 j.Jm high
a
c
e
d
f
Text-fig. 4. Monosulcus-pattern of Clavatipollenites variabilis n. sp.
and 2- 3 j.tm wide); sulcus frequently indistinct and represented as an area with coarser clavae or as an indeterminate tear in the exine. Exine consist- ing of inner unsculptured endexine ca. 0.5 j.tm thick and ectexine formed of baculate or clavate projections 1-1.5 f..lm long, which either remain discrete or fuse at their tips to form a microreticulum or a tectate exine; lumina of reticulum irregularly polygonal, 0.5-2 j.tm in diameter. Grain often folded.
Measurements: 33-53 j.tm X 28-47 f..lm in diameter.
SEM: 24.7-28.2 f..lm X 22.9-24.1 J.tm in diameter.
Occurrence: Uge Member, south of Kanuka (C 31), cliff near private house, north of Uge station (C 6), and north of Uge station (C 15).
Holotype: PI. 80, figs. 3 a - b; 38 X 36 f..lmin diameter; clavate projections 1.5- 2 f..lm high in sulcus and 1-1.5 f..lm high in the exine; tectate exine; lumina of reticulum 0.5-2 f..lm in diameter; slide C31-b.
Name derivation: variabil- Cis) Oat.) =vary, -able.
Remarks: The present specimens which occurred abundantly from the Uge Member, are closely similar to Clauatipollenites tenellus Phillips&Felix 0972, p.466, pI. 15, figs. 19-21; Ward, 1986, pp.28-29, pI. 1, figs. 11-12) from the Albian of Louisiana and Kansas CU.S.A.), but differ by the larger size.
Botancial affinity: ? Chloranthaceae, (Ascarina).
Genus Clavainaperturites Van der Hammen& Wijmstra 1964.
Type species: Clavainaperturites clavatus Van der Hammen & Wijmstra 1964.
Clavainaperturites sp. a PI. 97, fig. 7.
Description: Inaperturate pollen grain with verrucate and clavate ornaments.
Outline originally circular(?). Exine with verrucate, clavate elements, 1-2.5
f..lm high and O.5 - 2 j.tm wide, secondary folded.
Measurements: 54X 42 /J.m in diameter.
Occurrence: Uge Member, south of Kanuka (C 31).
Remarks: A single specimen was observed. The authors cannot find a com- parable species.
Botancial affinity: Unknown.
Clauainaperturites sp. b PI. 97, fig. 8.
Description: Inaperturate pollen grain with clavate and verrucate sculptures.
Outline circular. Exine clavate and verrucate, 1-4 IJ,m high and O. 5 - 6 IJ,m
in diameter; projections distributed relatively densely.
Measurements: 49 X 46 IJ,m in diameter.
Occurrence: Uge Member, Uge (C34).
Remarks: Only one specimen was encountered.
Botancial affinity: Unknown.
Angiospermic pollen
Genus Arecipites Wodehouse1933.
Type species: Arecipites punctatus Wodehouse1933.
Arecipites pflugii(Takahashi) Krutzsch PI. 86, fig. 16.
1961 Monocolpopollenites pflugii Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, GeoI., 11, no. 3, p. 294, pI. 16, figs. 31-32.
1964 Monocolpopollenites pflugii Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, GeoI., 14, no. 3, p. 249, pI. 33, figs. 12-15.
1970 Arecipites pflugii (Takahashi 1961) Krutzsch, Atlas, Lfg. VII, p. 29.
1979 Arecipites pflugii (Takahashi) Krutzsch, Takahashi&Kim, Palaeontographica, B, 170, p. 35, pI. 8, figs. 23, 28, 29.
Description: See Takahashi (1961).
Measurements: 18.5-34.5 IJ,m in size, lumina of reticulum 1-3 fJID in diameter, colpus less than 2 IJ,m wide (Takahashi,1961); 18 - 27 IJ,m X11.8 -19.5 IJ,m in size, exine 0.7 IJ,m thick, lumina of reticulum until 1 IJ,m in diameter, muri un- til 1.2 IJ,m high (Takahashi& Kim, 1979); the present spesimen: 35 X191J,m in size, exineIlJ,m thick, lumina of reticulum1-3 IJ,m in diameter, width/length ra tio O.543.
Occurrence: Uge Member, south of Kanuka (C31).
Previous records: Palaeogene, West Japan (Takahashi, 196f); Campanian, Hokkaido (Japan) (Takahashi,1964); early Miocene, Changgi (Korea) (Taka- hashi&Kim, 1979).
Remarks: The only specimen detected reveals all the characteristic features of Arecipites pflugii(Takahashi) Krutzsch.
Botancial affinity: ?Palmae.
Arecipites sp. a PI. 86, fig. 14.
Description: Monocolpate pollen grain. Amb elliptical or prolate in polar view. Colpus narrow, undulate. Exine finely reticulate; lumina of reticulum 0.5 IJ,m + in diameter.
Measurements: 28 X 17 IJ,m in size.
Width/length ratio: 0.607.
Occurrence: Uge Member, south of Kanuka (C 31).
Remarks: Only one specimen was observed.
Botancial affinity: ? Palmae.
Arecipites sp. b PI. 86, fig. 15.
Description: Monocolpate pollen grain. Outline elliptical or prolate in polar view. Colpus very slender, straight. Exine thin, finely reticulate; lumina of reticulum 0.5-1 IJ,m in diameter.
Measurements: 32 X 22 IJ,m in size.
Width/length ratio: 0.687.
Occurrence: Uge Member, cliff near private house, north of Uge station (C 6).
Remarks: The only specimen observed differs from the formerly described species of Arecipites.
Botancial affinity: ? Palmae.
Arecipites sp. c PI. 87, fig. 1.
Description: Monocolpate pollen grain. Outline elliptical or prolate with more or less pointed corners in polar view. Colpus slender, slightly curved, reach- ing both corners. Exine thin, 1IJ,m thick, finely reticulate; lumina of reticulum less than 0.5 J.1-m in diameter.
Measurements: 26 X 13.5 IJ,ID in size.
Width/length ratio: 0.519.
Occurrence: Uge Member, south of Kanuka (C 31).
Remarks: Only one specimen was found.
Botancial affinity: ? Palmae.
Arecipites sp. d PI. 87, fig. 2.
Description: Monocolpate pollen grain. Amb elliptical or prolate with point- ed corners in polar view. Colpus frail, straight, reaching both corners. Ex- ine finely reticulate; lumina of reticulum 1-1.5 /lm in diameter; muri baculate, 1 /lm high.
Measurements: 35 X 23 /lm in size.
Width/length ratio: 0.657.
Occurrence: Uge Member, north of Uge station (A) (C 33).
Remarks: The only specimen observed is superficially similar to Arecipites longicolpatus Krutzsch 0970, p. 112, pI. 25, figs. 1-13), but differs in hav- ing broader form and weaker colpus.
Botancial affinity: ?Palmae.
Arecipites sp. e PI. 87, figs. 3-4.
Description: Monocolpate pollen grains. Outline broad-elliptical or prolate- subprolate in polar view. Colpus narrow, curved, almost reaching both cor- ners. Exine reticulate; lumina of reticulum 1- 4 /lm in diameter; muri bacu- late, partially tectate, 1.5-2 /lm high.
Measurements: 29-32 /lm X 20-27 /lm in size.
Width/length ratio: 0.689-0.843.
Occurrence: Uge Member, north of Uge station (A) (C 33).
Remarks: The grains resemble superficiallyArecipites oligocaenicusKrutzsch 0970, p. 108, pI. 23, figs. 1-10) from the Oligocene of Germany, but differ in having larger reticulum and muri.
Botancial affinity: ?Palmae or ?Liliaceae.
Genus Artiopollis Agasie 1969.
Type species: Artiopollis indivisus Agasie 1969.
Artiopollis cr. indivisus Agasie PI. 95, figs. 5 a-c.
1969 Artiopollis indivisus Agasie, Micropaleontology, 15, no. 1, p. 28, pI. 4, figs.
6-8.
Description: See Agasie (1969).
Measurements: Diameter of tetrad 27 - (35) -46 J.l.ffi, exine about 2.5 - 3 J1,m thick, lumina of reticulum about 0.5-1.5 /.lm in diameter, muriO.5-1 /.lm wide (Agasie, 1969); the present specimen: 40 X 35 J1,m in diameter of tetrad, lumina of reticulum 1-3 /.lID in diameter, muri baculate-tectate, 1 /.lm high, each grain of the tetrad 28 X 23 /.lID in diameter.
Occurrence: Uge Member, south of Kanuka (C 31).
Previous record: Early Late Cretaceous (Cenominian), Arizona (U.S.A.) (Agasie, 1969).
Remarks: A single specimen was observed. This is closely similar to Artio- pollis indivisus Agasie from the early Late Cretaceous (Cenomanian) Dakota Sandstone of Arizona, but differs somewhat from the latter in its minute comparison of reticulum and muri.
Botancial affinity: Dicotyledonae.
Genus Asteropollis Hedlund& Norris 1968.
Type species: Asteropollis asteroides Hedlund& Norris 1968.
Asteropollis clauatus (Phillips& Felix) Ward PI. 82, figs. 6-7.
1969 Clavatipollenites sp., Doyle, Jour. Arnold Arboretum, 50, no. 1, fig. 1-h.
1972 Porotrichotomosulcus clavatus Phillips&Felix, Pollen et spores, 13, no. 3, pp.
471-472, pI. 16, figs. 15-16.
1980 Asteropollis asteroides Hedlund & Norris, Burger, BMR Bull., pI. 19, figs. 15- 17, 19(p.p,).
1986 Asteropollis clavata (Phillips&Felix) Ward, Palaeontographica, B, 202, p. 26, pI. 7, figs. 9-10.
Description: See Phillips& Felix (1972) and Ward (1986).
Measurements: 24- (27) -30 /.lm in equatorial diameter (holotype25/.lm), cla- vate projections O. 5-0. 75 J1,m high (Phillips&Felix, 1972); 30 /.lm in diameter (Doyle, 1969); 25 /.lm in diameter (Ward, 1986); the present specimens: 27.7- 30.6 /.lm X 19-23 J1,ID in diameter, clavate-baculate projections in the triradi- ate zone 1.7-2.4 J1,m long.
Occurrence: Uge Member, south of Kanuka (C 31).
Previous records: Upper Albian-lower Cenomanian, Atlantic coastal plain (U.
S.A.) (Doyle, 1969); Albian, Louisiana (U.S.A.) (Phillips&Felix, 1972); Lower Cretaceous, Queensland (Australia) (Burger, 1980); Albian, Kansas (U.S.A.)
(Ward,1986).
Remarks: Ward (1986) stated that Asteropollis clavatus is treated as variants of A. asteroides and its oroidate sulcus ends of the triradiate zone are war- ranted as a specific distinction.
Botancial affinity: Unknown.
Genus Callistopollenites Srivastava 1969.
Type species: Callistopollenites radiatostriatusCMtchedlishvili 1961) Srivastava 1969.
Callistopollenites radiatostriatus(Mtchedlishvili) Srivastava PI. 88, figs. 27 a-b.
1961 Tricolporites radiatostriatus Mtchedlishvili, Trudy VNIGRI, no. 177, p. 249-250, pI. 81, figs. 1a-g, 2 a-b.
1965 Tricolporo-pollenites radiatostriatus (Mtchedlishvili) Bratzeva, Trudy Geol Inst., Acad. Sci. USSR, 129, pp. 26-27, pl.l0, figs. 4-8; pI. 11, figs. 1-7; pI. 12, figs. 1-6 (p.p,).
1969 Callistopollenites radiatostriatus (Mtchedlishvili) Srivastava, J. Sen Memorial Volume, p. 65, pI. 2, figs. 37-38.
1969 Tricolporopollenites radiatostriatus (N. Mtch.) Bratz., Bratzeva, Trudy GeoI. Inst.
Order Red Banner Sab., Acad. Sci. USSR, 207, pI. 31, figs. 3-5.
1969 Tricolporopollenites radiatostriatus(N.Mtchedlishvili) Bratzeva, Chlonova, Trudy Inst. GeoI. Geophys., Acad. Sci. USSA, Siberian Br., p. 60, pI. 13, fig. 11.
1982 Callistopollenites radiatostriatus (MtchedlishviW Srivastava, Takahashi&Shimono, Bull. Fac. Liberal Arts, Nagasaki Univ., Nat. Sci., 22, no. 2, pp. 44, pI. 9, fig.
20.
Description: See Mtchedlishvili in Samoilovitch&Mtchedlishvili (1961) and Sri- vastava (1969).
Measurements: 23.8-35.4 Ji,m in diameter CMtchedlishvili in Samoilovitch&
Mtchedlishvili, 1961); 25-47 fJ,m in equatorial diameter, 23-30 fJ,m in length, exine 2-2.5 Ji,m thick (Bratzeva, 1965); 3004-36.0 fJ,m in polar axis, 28.8- 36.8
Ji,m in equatorial axis (Srivastava, 1969); 24-39 fJ,m in diameter, exine 2 - 2.5
Ji,m thick (Chlonova, 1969); 38 X 33.5 fJ,m in equatorial diameter, muri 1.7
fJ,m high (Takahashi&Shimono, 1982); the present specimen: 36 X 32Ji,m in di- ameter, muri baculate, 1 Ji,m high.
Occurrence: Uge Member, south of Kanuka (C 31).
Previous records: Maastrichtian-Danian, western Siberian lowland (Mtchedli- shvili, 1961); Maastrichtian, Zeya-Bureya depression (Bratzeva, 1965, 1969;
Chlonova, 1969); Maastrichtian, Alberta (Canada) (Srivastava, 1969); Maast- richtian, Hida (Japan) (Takahashi& Shimono, 1982).
Remarks: Hitherto, this species has been reported from the Maastrichtian of Canada, the Far East, western Siberian lowland and Japan, and this time was found first from the Santonian of Taneichi (Japan).
Botancial affinity: Unknown.
Genus Cricotriporites Leidelmeyer 1966.
Type species: Cricotriporites guianensis Leidelmeyer 1966.
Cricotriporites sp.
PI. 95, fig. 8.
Description: Triporate pollen grain. Outline subcircular in polar view.
Exine two-layered, secondary foleded; ektexine finely punctate on the surface;
muri finely baculate in the optical section, 0.6 J,J.m high; endexine thin, 0.5 J,J.m
thick. Pores circular, 6-6.5 J,J.m in diameter and furnished with both an an- nulus and costae pori; thickness of annulus ca. 1-3 J.lm; one pore is placed subequatorially.
Measurements: 45 X 37 11m in equatorial diameter.
Occurrence: Uge Member, Uge harbor (A) (C 39).
Remarks: A single specimen was observed. This is much larger than Criotri- porites guianensisLeidelmeyer from the early Eocene of British Guyana (1966, p.54, pI. 4, fig. 4), C. anulatus Takahashi& Jux (1989, p.260, pI. 16, figs. 6- 7) and C. nigerianus Takahashi & Jux from the Middle Tertiary of Nigeria (1989, pp. 260-261, pI. 16, figs. 10-16; pI. 38, figs. 1-2).
Botanical affinity: Unknown.
Genus CupulijeroidaepollenitesPotonie, Thomson& Thiergart 1950 ex Potonie 1960.
Type species: Cupulijeroidaepollenites liblarensis (Thomson 1950) Potonie1960.
Cupulijeroidaepollenites ditis (Takahashi) Takahashi PI. 87, fig. 10, pI. 98, fig. 7.
1957 Tricolpopollenites ditis Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, GeoI., 5, no. 4, pp. 218-219, pI. 38, figs. 46-49; pI. 39, figs. 39-44.
1961 Tricolpopollenites ditis Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, GeoI.,
11, no. 3, p. 312, pl. 22, figs. 24-31.
1964 Tricolpopollenites cf. ditisTakahashi, Mem. Fac. Sci., Kyushu Univ. Ser. D, Geol., 14, no. 3, p. 240, pi. 39, fig. 2.
1979 Cupulijeroidaepollenites ditis (Takahashi) Takahashi, Takahashi& Kim, Palaeon- tographica, B, 170, p. 37, pl. 9, figs. 1-2.
1988 Cupulijeroidaepollenites ditis(Takahashi) Takahashi, Bull. Fac. Liberal Arts, Na- gasaki Univ., Nat. Sci., 28, no. 2, p. 129, pI. 19, fig. 18.
Description: See Takahashi (1957).
Measurements: 15 - 47 p,m in size (holotype 31. 1 X 19. 2 p,m in size), width /length ratio: 0.4-0.8 (Takahashi, 1957; 25-30.7 p,m X16-20.2 /.lm in size, exine thin (Takahashi&Kim, 1979); 20.3X14. 3 /lm in size, exine thin, O.7/.lrn in the pole area, width/length ratio: O. 7 (Takahashi, 1988); the present speci- men: 29 X16 p,m in size, exineO. 7 p,m in the pole area, width/ length ratio:
0.55.
Occurrence: Uge Member, south of Kanuka (C3l).
Previous records: Palaeogene and Miocene, Japan (Takahashi, 1957, 1961);
Maastrichtian and Eocene, Oyubari-Hokkaido (Japan) (Takahashi, 1964);
early and middle Miocene, Korea (Takahashi&Kim, 1979); Coniacian, Futa- ba (Japan) (Takahashi, 1988).
Remarks: This species appears abundantly in the Palaeogene of Japan. Here in the Uge Member, only one specimen was found.
Botanical affinity: Cupuliferae.
Cupuliferoidaepollenites fallax (Potonie) Potonie PI. 87, figs. 20-25; pI. 98, figs. 4, 8-10.
1934 Pollenites jallax Potonie, Arb. Inst. PaHiobot. Petrogr. Brennst., 4, pi. 70, fig.
10.
1951 Cupulijeroipollenites jallax Potonie, Palaeontographica, B, 91, pI. 20, fig. 66.
1953 Tricolpopollenites liblarensis(Thomson) Thomson&Pflugjallax(Potonie) Thomson
&Pflug, Palaeontographica, B, 94, p. 97, pi. 11, figs. 133-151.
1961 Tricolpopollenites liblarensis (Thomson) jallax (R. Potoni~) , Takahashi, Mem.
Fac. Sci., Kyushu Univ., Ser. D, Geol., 11, no. 3, p. 314, pi. 23, figs. 5-11.
1977 Tricolporopollenites jallax(R.Pot. 1934) W. Kr. 1960, Krutzsch&Vanhoorn, Pal- aeontographica, B, 163, p. 70, pi. 29, figs. 36-40.
1979 Cupulijeroidaepollenites jallax(Potonie) Takahashi, Takahashi&Kim, Palaeonto- graphica, B, 170, p. 38, pi. 9, figs. 2 (?), 22-23.
1980 Tricolpopollenites liblarensis (Thomson in R. Potonie, Thomson & Thiergart 1950) Thomson& Pflug 1953 ssp. jallax(R.Potonie1934) Thomson & Pflug 1953. Thiele-
Pfeiffer, Palaeontographica, B, 174, p. 144, pI. 11. fgigs. 13-15.
1982 Cupulijeroidaepollenites jallax (Potonie) Takahashi, Takahashi&Jux, Bull. Fac.
Liberal Arts, Nagasaki Univ., Nat. Sci., 23, no. 1, p. 42, pI. 4, figs. 26-30.
1984 Tricolpopollenites liblarensis (Thomson in R. Potonic:;, Thomosn&Thiergart 1950) Thomson&Pflug1953jallax (R.Potonie 1934) Thomson& Pflug 1953. Mohr, Pala- eontographica, B, 191, pp. 76-77, pI. 12. fig. 8.
1984 Tricolpopollenites liblarensis (Thomson in R. Potonic:;, Thomson&Thiergart 1950) Thomson&Pflug 1953 ssp. jallax (R.Potoni(j 1934) Thomson&Pflug 1953, Kirch- ner, Palaeontographica, B, 192, p. 118. pI. 6, figs. 15 a- b.
1985 Tricolpopollenites liblarensis(Thomson 1950) Th. et Pf. 1953 ssp jallax(R.Potoni(j 1934) Thomson&Pflug 1953. Nagy, GeoI. Hung., Fas. 47, p. 201. pI. CXV, figs.
5-6.
1986 Cupulijeroidaepollenites jallax (Potonie- ) Potoni~, Takahashi&Jux, Bull. Fac.
Liberal Arts, Nagasaki Univ., Nat. Sci., 26. no. 2. pp. 132-133. pI. 22, fig. 21;
pI. 27. figs. 15-16.
1988 Cupulijeroidaepollenites jallax (Potoni~) Potonic:;, Takahashi, Bull. Fac. Liberal Arts, Nagasaki Univ., Nat. Sci., 28. no. 2, pp. 127-128, pI. 18, figs. 6-11; pI.
19. figs. 10-11, 13-17 a.
1989 Cupulijeroidaepollenites jallax (Potonie- ) Potoni~, Takahashi&Jux, Bull. Fac.
Liberal Aats, Nagasaki Univ., Nat. Sci., 29. no. 2, pp. 406-407. pI. 11, fig. 9.
Description: See Potonie (1934) and Thomson & Pflug(1953).
Measurements: 12-14 J.lm in size (Potoni{l,1934); 13 jJIflin size (Potonie,1951);
10-18 J.lm in size (Thomson & Pflug, 1953); 12-18 J.lrn in size (Krutzsch&
Vanhoorn, 1977); 11-13.5 J.lrn X 6, 7-8. 2 J.lm'in size, exine thin, 0.3-0.6
J.lm thick (Takahashi & Kim, 1979): 7.4X 13. 3 J.lm, width/length ratio: ca.
0.6 (Thiele-Pfeiffer, 1980); 11.7-14.4 J.lrn X5. 0-7. 7 J.lrn in size, width/length ratio: 0.4-0.5(Takahashi & Jux, 1982); 14X 17 /.lIn in size (Mohr, 1984); 12 J.lrn in size (Kirchner, 1984); 10-17 J.lm in size (Nagy, 1985); 10-18 J.lm X5- 8 J.lm in size, width/length ratio: O. 4-0. 5 (Takahashi & Jux, 1986); 11. 4-18 J.lrn X6. 6-11 J.lrn in size, width/length ratio: 0.42 - O. 76 (Takahashi, 1988);
16 X10 J.lrn in size, width/length ratio: 0.625(Takahashi & Jux, 1989); the pre- sent specimens: 16.5 -19 J.lm X9-11 J.lm in size (SEM: 12.8-15 J.lm X 7.3-10
J.lm in size).
Occurrence: Uge Member, south of Kanuka (C31), cliff near private house, north of Uge station(C6), north of Uge station (A) (C33), and Uge harbor (C17).
Previous records: Eocene, Geiseltal (Germany) (Potonie, 1934, 1951); Palaeo- cene-Pliocene, W. - Germany (Thomson& Pflug, 1953); Palaeogene-Miocene, West Japan (Takahashi, 1961); upper Landenian, Epinois and Loksbergen
(Belgium) (Krutzsch& Vanhoorn, 1977); middle Miocene, Yonil (Korea) (Ta- kahasi& Kim, 1979); Miocene, Wackersdorf/Oberpfalz (W. -Germany) (Thiele- Pfeiffer, 1980); late Oligocene, Bergish land(W. -Germany) (Takahashi& Jux, 1982); late Miocene, Frechen and Fortuna Garsdorf (W. -Germany) (Mohr, 1984); late Oligocene, south Bavaria (W, -Germany) (Kirchner, 1984); Miocene, Hungary (Nagy, 1985); late Oligocene, St Augustin (W, -Germany) (Takaha-
shi& Jux, 1986); Coniacian-Santonian, Futaba (Japan) (Takahashi, 1988); late
Eocene to early Oligocene, Fayum Oasis (Egypt) (Takahashi& Jux, 1989).
Remarks: This species appears widely in the Tertiary of Europe and in the Palaeogene and Upper Cretaceous of Japan.
Botanical affinity: Regarding its botanical relationship, Leguminosae, Cupu- liferae and other herbs are mentioned (Potonie, 1934; Thomson& Pflug, 1953;
Nagy, 1985).
Cupulijeroidaepollenites lanceolatusn. sp.
PI. 87, figs. 26-28; pI. 98, fig. 11 (cf.).
Description: Tricolpatc pollen grains. Amb long-elliptical or perprolate in equatorial view. Three colpi narrow, running parallel from pole to pole.
Exine two-layered, thin, O.5-1 jJ,m thick, laevigate or weakly intrabaculate.
Measurements: 19-23 jJ,m X6-10.5 11m in size.
Width/length ratio: 0.315-0.5.
(SEM: 29.3 X 11 jJ,m in size; width/length ratio: 0.375).
Occurrence: Uge Member, south of Kanuka (C 31).
Holotype: PI. 87, fig. 27; 23 X 8.5 11m in size, exine 1 jJ,m thick, very weakly intrabaculate; width/length ratio: ca. 0.37; slide C 31- b.
Name derivation: lanceolatus (lat.)= lanceolate.
Remarks: The present specimens resemble Cupulijeroidaepollenites liblaren- sis (Thomson 1950) Potonie 1960 and Tricolpopollenites lanceolatusTakahashi
& Jux (1989, p.408, pI. 12, figs. 2-4), but differ from C. liblarensis in being
much narrow and longer form and from T. lanceolatus in h_aving laevigate or weakly Jntrabaculate exine.
Botanical affinity: Unknown.
Cupulijeroidaepollenites vulgaris (Takahashi) Takahashi PI. 87, figs. 11-14; pI. 88, figs. 16-18;
pI. 97, figs. 9-10.
1957 Tricolpopollenites vulgaris Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, GeoI.,5, no. 4, p. 218, pI. 38, figs 44-45; pI. 39, fig. 38.
1961 Tricolpopollenites vulgaris Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, GeoI., 11, no. 3, p. 312, pI. 22, figs. 32-37.
1964 Tricolpopollenites vulgaris Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, GeoI., 14, no. 3, p. 260, pI. 44, figs. 22, 40.
1979 Cupulijeroidaepollenites vulgaris (Takahashi) Takahashi, Takahashi&Kim, Palae- ontographica, B, 170, p. 37, pI. 9, figs. 7-8.
Description: See Takahashi (1957) and Takahashi & Kim (1979).
Measurements: 14-36 /lID in size, exine less than 1 /lm thick, width/length ratio: 0.4-0.8 (type 25 X 17.2 /lID in size, exineO. 8 /lm thick, width/length ratio: ca. O. 7) (Takahashi,1957); 22.7-23.6 /lm X 12. 2-12.6 /lIDin size, exine thin (Takahashi& Kim, 1979); the present specimens: 17-27 /lm X 10-17 /lm
in size (SE.M: 18. 8-21 /lID X 9. 4 /lm in size), exine O. 5- 1 /lm thick, more or less chagrenate, width/length ratio: 0.45-0. 765.
Occurrence: Uge Member, south of Kanuka (C 31), north of Uge station (A) (C 33), and north of Uge station (C 15).
Previous records: Palaeogene, north and west Japan (Takahashi, 1957, 1961, 1964); middle Miocene, Yonil (Korea) (Takahashi& Kim, 1979).
Remarks: Morphologically the specimens appear to be closely comparable to those of Cupuliferoidaepollenites vulgaris (Takahashi) Takahashi.
Botanical affinity: Cupuliferae.
Genus Cyrillaceaepollenites Miirriger& Pflug ex Potonie 1960.
Type species: Cyrillaceaepollenites megaexactus (Potonie 1931) Potonie 1960.
Cyrillaceaepollenites minor (Takahashi) Takahashi PI. 94, figs. 6-8.
1961 Tricolporopollenites minor Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, GeoI., 11, no. 3, pp. 320-321, pI. 24, figs. 18-31.
1962 Tricolporopollenites minor Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, Geo!., 12, no. 1, p. 11, pI. 4, figs. 34-36.
1963 Tricolporopollenites minor Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, Geol., 14, no. 2, p. 150, pI. 21, fig. 13.
1964 Tricolporopollenites minor Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, GeoI., 14, no. 3, pp. 244-245, pI. 39, figs. 16-18; pI. 41, figs. 22-25; pI. 42, fig. 10.
1979 Cyrillaceaepollenites minor (Takahashi) Takahashi, Takahashi&Kim, Palaeon to-
graphica, B, 170. Lfg.1-3, p. 46, pI. 13, figs. 32-36.
Description: See Takahashi(1961) and Takahashi& Kim (1979).
Measurements: 13.4- 24.5 pmin size, exine less than1/.lm thick, width/length ratio: 0.8-1. 0(holotype ca. 17 /.lm in size) (Takahashi, 1961); 11-17. 2 /.lm X 8-16 /.lm in size, exine0.5-1. 2 /.lm thick, width/length ratio: O. 75-0.88 (Ta- kahashi& Kim, 1979); the present specimens: 13-18 /.lm X 12 -17 /.lm in size, exine0.5 /.lm thick, width/length ratio: 0.888-0.923.
Occurrence: Uge Member, south of Kanuka (C 31) and north of Uge station (A) (C33).
Previous records: Palaeogene and Miocene, western Japan (Takahashi,1961);
Campanian to Maastrichtian and Eocene, Hokkaido (Japan) (Takahashi, 1964); middle Miocene, Yonil (Korea) (Takahashi& Kim, 1979).
Remarks: The minor form of Cyrillaceaepollenites is closely similar to Cyril- laceaepollenites megaexactus (Potonie, 1931) Potonie (1960) and C. exactus (Potonie, 1931)Potonie (1960) from the Tertiary of Middle Europe, but differs by its thinner exine.
Botanical affinity: Cyrillaceae, Cyrilla.
Genus Fibulapollis Chlonova1961 emend. Stanley1970.
Type species: Fibulapollis mirificus(Chlonova1957) Chlonova1961.
Fibulapollis enodatus(Chlonova) Takahashi PI. 85, figs. 8-9; pI. 86, figs. 1-5.
1961 Accuratipollis enodatus Chlonova, Trudy Inst. Geol. Geophys., Acad. Sci. USSR, Siberian Br., 7, p. 91, pI. 16, figs. 126, 126 a.
1972 Accuratipollis evanidus Chlonova, Miki, Jour. Fac. Sci. Hokkaido Univ., Ser. IV, GeoI.&Mineral.,15, nos. 3-4, pp. 574-575, pI. 11, figs.6a-b(p.p.).
1982 Fibulapollis enodats (Chlonova) Takahashi, Takahashi & Shimono. Bull. Fac. Lib- eral Arts, Nagasaki Univ., Nat. Sci., 22, no. 2, p. 80.
Description: See Chlonova (1961).
Measurements: 45-(57)-69 /.lm in size (Chlonova, 1961); the present speci- mens: 29-45 /.lm X 28-43 /.lm in diameter (37 - 52 /.lm in length of a side in polar view), exine striate or net-like, muri baculate-tectate, 1-2 /.lm high.
Occurrence: Uge Member, south of Kanuka (C31) and Uge (C34).
Remarks: The Uge specimens are more of less smaller than the Chlonova's specimens, but seem identical to Fibulapollis (aI. Accuratipollis) enodatus
(Chlonova) Takahashi 1982.
Botanical affinity: Santalaceae or Loranthaceae.
Fibulapollis evanidus (Chlonova) Takahashi PI. 85, figs. 1-7.
1961 Accuratipollis evanidus Chlonova, Trudy Inst. Gelo. Geophys., Acad. Sci. USSR, Siberian Br., 7, pp. 91-92, pI. 16, figs. 125, 125 a.
1972 Accuratipollis evanidus Chlonova, Miki, Jour. Fac. Sci., Hokkaido Univ., Ser. IV, Geol, &Mineral., 15, nos. 3-4, pp. 574-575, pI. 11, figs. 7-8 (p. p.).
1982 Fibulapollis evanidus (Chlonova, 1961) Takahashi, Takahashi&Shimono, Bull. Fac.
Liberal Arts, Nagasaki Univ., Nat. Sci., 22, no. 2, p. 80.
Description: See Chlonova (1961).
Measurements: 23-(27)-34 j.J,m in size (Chlonova, 1961); the present speci- mens: 32-40 j.J,m in overall length in lateral view, 22-31 j.J,m in height, 15-20
j.J,m in width of wings, 15- 22 j.J,m in length of wings, exine finely reticulate, lumina O. 5-3 IJ,m, muri baculate-tectate, 1-2 IJ,m high.
Occurrence: Uge Member, south of Kanuka (C 31) and north of Uge station (A) (C33).
Previous records: Late Upper Cretaceous, western Siberian lowland (Chlono- va, 1961); Santonian-early Campanian, Kuji (Japan) CMiki, 1972).
Remarks: Fibulapollis Cal. Accuratipollis) evanidus (Chlonova, 1961) Takaha- shi 1982 is smaller thanF. (aI. A.) enodatus CChlonova, 1961) Takahashi 1982.
Botanical affinity: Santalaceae or Loranthaceae.
Genus Foveotricolpites Pierce 1961.
Type spesies:Foveotricolpites sphaeroides Pierce 1961.
Foveotricolpites concinnus Singh PI. 91, fig. 16.
1971 Foveotricolpites concinnus Singh, Res. Coun. Alberta Bull., 28, 1, p. 195, pI. 29, figs. 10-11.
1986 Foveotricolpites concinnus Singh 1971, Ward, Palaeontographica, B, 202, Lfg. 1- 6, p. 40, pI. 8, figs. 21, 22.
Description: See Singh (1971).
Measurements: 25 - 33 j.J,m in polar axis, 14 - 25 IJ,m in equatorial diameter
(Singh, 1971); 35 X 18 Jim in size, lumina 0.3-1. 5Jim in diameter (Ward, 1986);
the present specimen: 37X 18 Jim in size, muri baculate-tectate, 1. 5 Jim high, lumina of reticulumO.5-1.5 Jim in diameter; width/length ratio: 0.486.
Occurrence: Uge Member, Uge (C 34).
Previous records: Middle to upper Albian, Alberta (Canada) (Singh, 1971);
middle Albian to Cenomanian, Kansas (U. S. A.) (Ward, 1986).
Remarks: Only one specimen was observed. This seems to be comparable to Foveotricolpites concinnusSingh (1971).
Botanical affinity: Unknown.
Foveotricolpites fastidiosus n. sp.
PI. 93, fig. 6; pI. 94, figs. 1 a-b.
Description: Tricolporate pollen grains. Figura ellipsoidal or prolate in lat- eral view and more or less triangular in oblique view. Three conspicuous margocolpi radially symmetrical, converging at the poles; margo 1. 7 - 2.0 Jim thick. Exine foveolate; lumina O.5-5 fJm in diameter or length, circular, elliptical, elongate in form, muri baculate-tectate, 1 Jim in width,1.5 J.IIflhigh on the sides and 2 Jim high on the poles.
Measurements: 61-69 Jim X44-48 Jim in size.
Width/length ratio: O.72.
Occurrence: Uge Member, Uge harbor (A) (C 39) and Uge harbor(B) (C 40).
Holotype: PI. 94, figs. 1 a - b; 61 X 44 Jim in size, colpus with margo (2 Jim in thickness); exine foveolate; foveolae circular, elliptical and elongate in form, 0.5-4.5 Jim in diameter or length; muri baculate-tectate, 1.5 (side)-2(pole)
Jim high; width/length ratio: 0.72; slide C 39-a.
Name derivation: fastidiosus Oat.)= feeling, loathing.
Remarks: This new species is a large form ofFoveotricolpites Pierce (1961).
The specimens differ from Foveotricolpites globosus n. sp. in form.
Botanical affinity: Unknown.
Foveotricolpites globosus n. sp.
PI. 92, figs. la-b, 5, 6a-b;pI. 93, figs. 4-5.
Description: Tricolpate pollen grains. Outline circular to subcircular or oval in lateral and polar views. Three conspicuous colpi radially symmetrical, con- verging at/near the poles. Exine foveolate; foveolae circular, elliptical, elon- gate etc. in form, O. 5-3 Jim in diameter or length developing largely in the
middle areas of the mesocolpia and finely along the colpi; ectexine in optical section intrabaculate or tectate on the sides, 1-1. 2 IJ,m high and baculate or tectate on the polar areas, 1.5-2 JlI!1 high.
Measurements: 40-(51) jJ.m X 34-49 jJ.m in size.
Width/length ratio: 0.833-0.93.
Occurrence: Uge Member, Uge harbor (C17), south of Kanuka (C31), and north of Uge station (A) (C 33).
Holotype: PI. 92, figs. 1a - b; 43X40 jJ.m in size; exine foveolate; foveolae cir- cular, elliptical, elongate (straight or curved) etc. in form, 0.5 - 2.5 jJ.m in diameter or length; muri baculate on the poles,!. 5 jJ.m high and intrabacula- te on the sides, IlJ,m thick; width/length ratio: 0.93; slide C33-a.
Name derivation: globosus (lat.) = spherical, globe-shaped.
Remarks: The specimens differ from Foveotricolpites sphaeroidesPierce (1961) with -a rounded shape in size of the grain.
Botanical affinity: Unknown.
Foveotricolpites sp. a PI. 89, fig. 6.
Description: Tricolpate pollen grain. Figura ellipsoidal or perprolateinlate- ral view. Three colpi narrow, converging at the poles. Exine foveolate; fov- eolae scattered, circular in form; ectexine intrarugulate, 0.5 IJ,m thick. Both the poles more or less pointed.
Measurements: 29X 14 pm in size.
Width/length ratio: 0.48.
Occurrence: Uge Member, south of Kanuka (C31).
Remarks: A single specimen was found.
Botanical affinity: Unknown.
Foveotricolpites sp. b PI. 100, fig. 11.
Description: Tricolpate pollen grain. Figura ellipsoidal or subprolate in equ- atorial view. Three colpi radially symmetrical, converging at the poles.
Exine foverolate; foveolae less thanO.5 jJ.rn in diameter.
Measurements: 17. 7 X 14 pm in size.
Width/length ratio: O.79.
Occurrence: Uge Member, south of Kanuka (C 31).
