A TAXONOMIC STUDY ON THE
CHRYSOMEuDAE
押NSE:CTA−COLE:OPTE:RAIFROM FORMOSA
PARTIT SUBFAMILY DONACHNAE
BJT
MicIlio CHOJ∂
(En紬mol噌iea王bboratory)
Subfamily DONACIINAE
Ponac;adaeKiTby,F飢1na触.・−AmeIican乱,ⅠⅤ,p.222(1▼837),
Donac芸ae RedtenbacheIこ,Gatt・I)eutschen Kafer・Fauna,P・113(1・845)・
Dd癒¢;des(Donacidae)Lacordairel,Monogr・Phytq)h・J(Mem.Soc・Roy■Sci・Liege,
Ⅰ、II),PP・91−92(Z845)・−Chapvis,inl−aCOrdaire,Hist・Nat・Insl・:Gen・Col・,Ⅹ,P・55
(ユ874)・・
わohaeはae C.Gl・Thomson,Skandinaviens Col”,VIII,P・105(1866).・J&COby et
Clavarぐau,in Wytsman,Gen・Insい,XXr,Col・Phytoph・‥DotLaCiゐe,P・1(1904)・
D⑳nae価Redtenbacher,Fauna Austriaca,ed・3,ⅠⅠ.,P・439(1874).}−Wickham,Canadian
Ent..,ⅩⅩⅤⅠ軋p.69(1紬6/.Donaci;gl盲Seidlitz,Fauna Baltica,.P・136(1875)・−Weise,Natur’g二Ins.Deutschlands,
VI,P.7(1882)・・・Everts,Col・Neerlandica,lT,PP。390&391(1903).・Heine et
Taschenberg,Exot・Kaf・,PP・・245&246(1908)・−・R・eit七er,FaunaGermanica,ⅠⅤ,PP.73&74(1912)・・Kuhnt,I11ustr・Best・一Tab・Kafl・Deutschlands,PP.801.(1913).∼
Schaufuss,inCalwer,Kえferbuch,ed.6,II,PP・888−・889,ユ907,ユ916(1913).−Reitter,
WienerEnt.Zeit”,ⅩⅩⅩⅤⅠⅠⅠ,1−3(Beもt.・−Tabl・eurOPえischen Col・,IJXXXVIII),P.21(1920)・1qScheerpeltz et Winkler,in Brohmer,、etC・,Tierwelt Mitteleurop寧S,Ⅴ,ii,2,
p・ⅩⅠⅠ−・220(1930)・
Doha¢i;na FowleI・,Col・Br■itisbIsIs・,ⅠⅤ,pp・265&266⊥267(1890)・
DonaciinaeJacoby,Fauna Brit..India,incl.Ceylon and Burm乳,一Col.ⅠⅠ:Chrysoml,
P.9(1908)・1−ChGj∂,TIanS・Nat・Hist・Socl・Formosa,ⅩⅩIV,135,PP.519 &‥521
(1934).1−Chen,Sinensia,ⅩIT,f−・6,PP。11−17(1941).−Gressitt,工ノingnanSci.Joum.,
ⅩⅩ,2・−4,p.275(1942).
E:upodinaeJoy,Pract・Handb.British Beetles,Ⅰ,PP・388&389(1932)(ParS)・
Genera璽e始ra¢康博(円gs..ト7,TO&柑):Bodyelongateor e王0刀g郎e−Oblong;bead,
antennae and scutellum finely pubescent,PrOnOtum andelytraglabrousinmany cases,
butinsome cases moreorless pubesoent and rarely pronotum fi71ely pul)eSCent hut
exp6sed,generallydeclined,distinctly
orrathやrindistinctlYCOnStrictedbehind theeye芦;neCk−reglOn distinQt;inter− ocularspace、vithastrongcehtrallon“ f■or・eleg・叫 ・・一一prOnOtu血ノ gitudinalgroove(丘pollta11ongitudinal humefuS・・..grDOVe)andinDonactaFabriciusthere
is another’weak gr’00Venear eaハheye
(OCular groove);the area surrounding
__9Cutellum middleleg・− scu亡q王Iar rて)Ⅵ of ︰り㌧︰り‖Jり︰r︰ − −−一■■−−−− 一l■■
pu読turus eachantennal socket more orless
elytral
SutuTe
raised,Clypeusgenerallyobpentagonal,
ヒe stria With加ntmargln Straightoremargl− nate ar)ddorspmmorIe OrIless convex.
...▼.....▲−−・−㍉㌧りシ〓
−■−−−●−−−▼1一■▼−−−−▲
elyとTOn Mouth−PartS’’(’Figs.3−6)placedatthe
anteriorendofhead;labrumtransvや葺Se, With f士IOnt COrnerS rOunded and front
marglnentireornotchedatthemiddle;
mandibles robust,incur・ved,POinted
atthe呼eX Whichis splitin some
CaSeS,genera11y armedwith?neOr
more.teethonthelowerside;makillae mediumlOr Sma11;maXillary palpi4・
・イernlir・ わin(=密・−・・・ apex of .・一 ely奴■On Fiヱ・1…肋花邪ね(ざ‖れ)㈹攣沼地icids,♀,f工Om Getmany;do工Sa■1aspe〇t.
Segmented,withlst segment
ⅥSu乱11y much sma11er th乱n any
One Ofthe three hllowlngSand
t:helatters rather vafiable in
Shapeandsize;1igulastlhquadrate、,
notstronglychitinised,Oftcnsemi・ membr’an?uS,‘flatorrぬerconve女, Withftontmarglnentireorgently emarginate;1abialpalpi3・Segme−nted,Withlstsegmentminute乳nd
thetwofo1lowingsrathervariablc
in shqpe and size;・mentum
trIanSVeise,With sidc占founded
Or angulated and fr・ont margln
emarginate・Eyes(Figs・1&3)
ratherlarge,rOundc申,COnVeXand
inelifac&ted・Ante竺nae(Figl1) 11・S、egmented,filifbrm,PrOPOrtiona・ t色1ylong,inserted’in fr・ont of theeyesandratberapproximat占 Fig・2・かの劇血¢・威r、1つ8和郎画FabIicius,♀,f工・OmGe∫mapy; VentZalaspec[,Withhead andleftlegs工emOVed.to each other at basp..Pr・onotum 伴igトS・l&きト甲乱dr乱teOt・neaT・1yso,
moreorIlessIon拷er,thanbroador the
r冶VerSe,Withou亡1aterIal,rmaTIg1nS,With dnteriQrPOrtion、15WOlleムortuber・Culate atoi,n呼・′伽∴二もnかs、inma叩CaSeSand fr?que山1yl桓がtud、in描アgrOOVed
onthe middle of dorsllh小 Scutellum
r顆・さ1)obtr・iaヮgul鱒or n甲巾sd・ El仰臥r硝・1)elbh亘ate,uSuallym血 br・0よdeT・atbaseth8nくpr・OnOtum、para11eト Sidedorna産ow9dposteriorly witha
gentlecurva抽re,rOlユndedor tr11neated
bremaT・か孔ted瓦t the8peX;dorsuh
more orless convex,butthe median
areae generIally flat,in many cases
regt)larlY PunCtate−StrIiate,butrareIy
COnnlSedly pⅥnCt111ate;in Hbemonia
I.atre;11ethelaterol−OSterior end of
each elytron strongly splnOSe Or
dentate and tlle Slユt11r818reamOre Or
7ess raised..Prostcrnum(Fig.2)
narrow orJ=・atl】Crindistinct be†wf>Cn
the 鈷re_COX乱e.blIも more のrless
widened at its posterior end snd
enterIlnghetween the apice$Of pro−
eplmera;aCetahulae clbsed bJ>bind.
Abdomen(Fign2)withls†苧ternite
mlユChlongerIthananyoneof†hefour
n)1lowingSlⅠ.eg’S(Figs・1&2)sIpnder,
Fig.3.刀血畑血(さ.8れ)¢7・が∂卸β8FabIicius,♀,frOm Germany=f王ead:doISalaspec[,with antennae re・mOVed・
Fig.4.刀伽畑血(β.れ)eノ,離さ如Fab工icius,♀,fIOm
Germany;Head‥Ventralaspect◆Withleftmandiblc
and maxillaI・emOVed.
gener・a11ylong;fbre・・COXae Cylindricalor conico−Cylindricaland
COntlguOuS Or CloselY aPPrOaChing to each other,middle・−COXae
moder・ately and hind−COXae Widely separated to one another;
femora somewhat thickened,hind1−femoralonger and thicker
thanthetwoanter・iorcorrespondingsegmentsandfrequentlvarmed
wjth one or more denticles on each lower side beyond the
middle;tibiaeslender,fore−andmiddle−tibiaearmedwithaspur
at theapexof eachlowerside,hind一−tibiae unarmedthespur,
hut in many cases the apex of each lower side sharply or
Obtuselydenticulated;tarSirathernarrow,f18t,with3rd segment
de甲1y bilobed and chw−・Segment Club1−Shaped and somewhat
mg.5.かow(ぬ(さ.紛.) 仰がβ卸郎 Fab工iciu$,♀, fⅠ・Om Ge工many;Left
CurVed;Clawssimple,long and curved;
in〟demク′乙∠■α Latでeilie the 臨r・SiveI・y Slender,C.yliムdrical,With3rdsegment
VerymuChshorterti泊n any dne(一f the
two p亨eceding$and not bilohed,and
Claw・Segment muChlonger than the
PreCCding se卯CntS COmhined・
Male genitaHa(Fi畠s.,10&13):
Medianlohelarge,tubular,CurVed,
With distalhalfstrongly chitinised,but
basaihalfnotstronglysoandoftensemi−
membranous;median orince situated
牢the apex,median foram?nlarfre and
occupying the ventral face of bssal
halflTegmen丘)rminga Welichitinised
F享g・6・pO脚申(s・声こク1・)¢冊S車齢Fab∫icius,宰・.FrOm Ge工many;Leどt maxi11a:Vent工alaspect.
rlng−Piece atthemiddle
and medianpleCe PaSS叫r
lng 血rou紳 thisI1ng;
dist盆Ipiecefor・ming an
elongateChitinouslobe,
notfbImingthe“1atera1 lobe〉,, With median
portionoftensemi・mem・
brano11S叩d8peX enliIe
Or Slightly em8ⅠSlnate
and luubescent or
Fig・71・Donac孝a(slSl7・)ora8SipesFabriiauヲ・♀,fromGermany;Left・Wing‥ doISalaspec亡.glabrous;hasalpiecc(tegminalstr’ut)semi−membranous and払rming a flat keel
Wings(Fig‖7):C?∠一simple,isolate;2Ashort,endin昌nearthe post−medianportion
OflAB and not connectlng tO thelatteli
Rang9=Palae甲Ctic,Or・ientala11dNearcticRegions;Senegal(OneSPeties ofDonacia tibrici呵,.Madagascar(DonaciastaFair・maire−incl.<)nlyonespecics;OneSPeCieso
Fablicius),AustTIalia(励rodonacta BluckbumiinclいOnlyonesp?Cies;OneSPeCics、of
Donacia Fabr・icius)and Mt,Ⅹico(One SPeCiesofDonacia Fabricius)・
Thepresent subf瓦milyis dividedin tvvo Tribes as払1lows:
Key to紬e Tr地es
A.Antennae taperlng tOWards the apex;tarSislenderand cylindrical,Sparingly pube〇Centbeneath,with the3rdsegmententireandtheclaw−3egmentuSual1ylongelthantheprecedingstogether;pubeocence of the underside oF body much modified,Often varnish・like HaeIⅥOnijn蚤
A,・一Antennaenot tapeElng tOWardstheapex;tarSinormal,dilated and thickly tomentosebeneath,with the3rd segment bilobed and theclaw−3egment Shorre工thantheprecedingstogether;pubescenceof
1n the above Tribcs,HdelTWnlinihas no relation tothc fauna of For・mOSa at preSent.
Tribe Donacibdis dividedinto somegenera,butonly−one genusinthem⊥−Donacia FねⅠicius一−・・is r・epresentinourfaunisticregionbythe pr・eSenCeOftwospecies・
GerluS Dona¢冨a FABRICI甲
DorIaCiaF?bricius,Sys亡.Ent.,p.195(1775)・−Latreille,Hist・Nat・CrustnInsい,Ⅹ†,P・淵(180り・・
Stephens,Illu出.Briti§hEnt・,MandibL・ⅠⅤ,Pp・267・望68(18:Sl)・・・Curtis,BritisヽEnt・,Ⅹl,Nol・ま9もユ823・ 1840(1834)・・IJaCOrdaire・Monogr・Phytoph・IO46m小Soc・Roy・Sci・Li6ge,IrI)・P?・92・・9S(1845)・1−Schilling,Uebers.Arb.岳chles.Ges..Breslau,p.88(18i6).−FaiImair・e,Gen:col・d,Europe,ⅠⅤ,P・206
(1856)..・C.G.Thom占ori,SkandinaviensCol..,VrII,p。105(1866).一Chapuis,inLacoIdairIe,Hist・Nat・ Ins.‥Gen.Col..,Ⅹ,P.5了(ユ874).−Weise,Naturg.・Ins・Deutschlands,Vr,Pl・14(1882)・−Fowler,Col・ BritishTsIs”ⅠⅤ,p・267(189o)小一Leng,Trans・AmericanEnt”Soc・,ⅩⅤ叩,p・159(1891)・・Seidlit去,FaunaBaltica,ed.乳p.728(1891.);払uilaT王anSSylvanica,p;819(1891).・Jacobson:HoraeSoc・Ent・
R占ssic左e,ⅩⅩVr,P・415(189月)・・Bedel,F?uneCol・BユS3inSeinc,Ⅴ,p・109(1889・・19Ol)・1−IEverrs,Col・
Neer・1andica,ⅠⅠ,ppサ f;91&398”399(1903).・JacobyetClavareau,iriWytsman,Gen・Insい,ⅩⅩⅠ,Col■ Phytoph:Do7もαCidae,p.3(1904)・・Jacoby,FaunaBIit・Tndia,incl.CeylonandBurma,Col・ⅠⅠ:ChIySOm・1,p・10(190声)・TPorta,Riv・Col・Italiana,ⅠⅩ,p・・150(ユ911)l・・Reitter,FaunaGermanica,lV,pp”7年&
7ら(19152).−Kuhnt,Illustr.Be3t・Tab.Kaf.Deutschlands,Ppl・801&810(’19ユ3)・,SQhaufuss,inCalwer・ Kaferbuch,ed.6,Ir,pp.8S9&89ユ,1907・1916(1913)l・・Re恒er,WienerEnt”Zeiし・,kxxvIIT,pp・1・3 (BesいTab.c11・OPAischenCol・,LXXXVIII),ppl潤&′26・・如(1920)・・Seheerpeltz6t、やinjく王er,呈ムetc.,TierweltMitteleuropas,Ⅴ,ii,男,Pu X汀−220{(一930)・−Joy,Pract・fIandb・BritishBeetlcs,r,pp・3由
&390(ユ932).・C旭j6,Trans.Nat.Hist.Soc・・Formo3a,ⅩⅩⅠⅤ,1籍5,pp”522&523・524r(19朗)・一Chen, Sわensia,Ⅹ‡Ⅰ,1・6,pり7(19隻1)・ Arun戚na叫SVoe亡,qat・Syst・Col・・,lI,p・31(ユ776)・ Genotype:DolulCiaora8SipesFabricius,Syst・Enし,p・195(1775),ftomEurope〔−Fixed byJ・Curtisin Bri亡isb Ent.,Ⅹl,No.494,ユS2:‡・・1840(18髄)〕・Range:PalaeaICtic,OrientalandNearc亡icRcgions;aSinglespecie3reSpeCtively f工Om Senegal,Madagascarク
AustTalia and Mexico.
Ger,ePa]chapaCtePS:Bodyelongateorelongate−Oblong;dorIsummetallicincoloration,
head,antennaC and scutellum finely pubescent,in血any cases pronotum and elytra
glabrous,butin somecases finely pubescent,and rar’elypronotumfinelypubescent as
we11as headbutelytraglabrous;underside andlegsvery thicklypubescent小Headwell
exposed,SOmeWhat declined,mOre Orless distinctly constr・icted behind the eyes;neCk−
reglOndistinct,With centralandlater・a170ngitudinalgroovesbetween the eyes;Clypeus obpentagonal,withfr・OntmarglnStraigIhtorgentlyemarg・inateanddorsummoreorless
convex・Mouth・Part・S:1abrumtranSVer・se・rOundedatthe ft・ontCOrnerS,g・entlyroundedor
straight or ratheremarginate and finely pubescent atthe front margln,mOre Orless
convexandsparselypubescentonthedors辻m;J㌻mandiblcsrobust,incurved,/boncaveAt
theinncrface,Sharplyansulatedandsplitat・theapex,Sharply and strongly unidentate
on thelower side;maXillae r・ather shoIt and robust;maXillary palpiwithlst segment
verymuch smallerthan anyoneofthethreefo1lowings,2ndand3rd nearly cylindrical
or obconicaland4thovalor ek)ngate−COnical;1igula subquadrate,flatorrather convex・
Withfront marglnrOunded orstraightorgentlyemarginate;1abialpalpiwithlst scgment
0r Str・Ongly roundedoneach sideandemar’ginate atLthefr?ntmargin:Eyes rather−1q・rge,
rounded,COnVeXandfinelyfacetted.Antenr3ae Sユender,filiform,With1stseg甲e叫9trOtngly
thickened and 寧Om¢What c里VCd,2nd tf} 4th rather slenderan一Ivariableinlcng叫,
generally the4thlonger thanany one ofthe two precedingS but sometimes ofsimi1ar
length,5th tollth subequalto one anotherI?r gradually dimir)ishing theirlength
terminallyandllthpointedattheapex.Prもnotumqd;dr・arteornearlyso,mOre()rless longer th左n、hrIOad orthe reverse,imぬarginAted坤e盆chside,With anterior portio71
swollenoItdherculateat or鱒rthe anglesinmanycases,but notso fnsomec争軍eS・
S9叫ellumobtriangular.ornearly so..ElytrIaelongate−triangularorI・tOngue・Shqp雪d or oblpng,
mu・Cと1br・Oad■er紙base thanp‡onO甑m,mundedor亡mnCatモdo=atherema‡glnated乳ttb・e
ap由;dorsum convcx,With血dian畠reagenerally widely鮎t,iri、many′CaSeS regularly
p血Ctat占−Striatc,butr益rely c16sely and confusedly punctulate or pube!C′ent・IPunCtulate,
intersticesofth?punCtate−Stri?emOreOrlesstransvelSelywri成1edor stIigosein誓any
qases,but not soin some cases.Abdom6n Withlst segment、nearly equalin:1engtIl/tO,
Or muChlorlger than,thc 錠山r succeedings together・Pygidium exf)OSCd bcyond、血the
apexofノelytr鋏OrhiddenunderthelatterィⅠノegSlong and slender;fbmorIa someWhat
thickened,hindqfbinor・a mu6hlongeraPd thickerthan theforeandmiddleonesand
often?rmedwith or]e,Or mOr9denticles on eachlower・side heyond themiddle;tihia
slender,f柁quently curvedor binuate,for¢・−・andmiddlel・七ihiae armedwith乳SPur atthe
apex o・f eachleweT Side,hindq・tibiae unarmed the spur’,butinmanycasesmoreotless
denticulatcd;tarSirlarrOW,rathcr short,Withlst and2nd segments trigonate br nbarly
so,3rd deeplybilob¢dandclaw−・SegmentClub1−Shap6dandsomewhatcurved;Cla示slong,
Simple and curved.
♂:Body narrトOWer,antenT)aC andlegslonger,1ast abdomin乳1$temite withhind maTgin
roundedandPygidiumwithhind margm gcntly roundedor・Strai紳tlytrdncated
♀:Ⅰ・aSt abdominalstemitβWithhipdmragin sharply angulateすor denticulat?d or
spino戸eandp沖diumwithhindmaTginsharplyorrIafherobtuselyangulated・
Thi$genuSisdividedinto four subgener・a aS follows‥
Key紬t如e Sゆ望enera
a・Do工SumOfbodyentirelyfine!y pube苧Cent aSWellaGtheunderside;elytraverynar工OWly elongate,
finelyand confusedlypuncctilateandintezspacesofthosepuncturcsmicroscopicallyfinelysculptu工ed
ヱ)0花α¢≠8gね Reitte工
a,.−Dorsum}Of body not entiIely pilbescent,∫elytraiegularly punctate−Striate b..Head and p工OnOtum finely pube〇Clent,eiytIa glabrous
b,.Head finely pubescent,PrOnO亡tlmand elytIa glabrous
助0ゐ托¢¢グαReitte工
c・Pronoちum ne恥1y o享、COrr)pletelyimpunctured,SmOOth・microscopical1y f’inely sculptured orvery finelytransverselyw王inkled;舎:1stabdominalsternitewithrap去irofsmalltubercleon the middle
q卯㊥胱e工・Goeeke
c,.Pronotum distinctly punctu工ed,Wrinkled,nOt SmOOth;$‥1st abdomまnalsternite without such・a
.かoMC≠αβ.政一.
tuberC・1e asin above
Subgenus Cyphogaster′GoEC王(E
Cyph喝a$!er(a:ia neW$ubgenus ofDonaciαFabricius)Goecke,Kol・Rundschau,ⅩⅩ,6,pp.琵15・幻7
(1934);Archo f”Hydrobiol・,Suppl・・Bd・Ⅹrv(Tropische BinnengやWa3SeI・,Bd,Ⅵ),pp.286・291
(i935);Ent”Bla:tい,ⅩⅩⅩIr,5,Pp.220・・色男ヨ(1936).・Chen,Sinensia,ⅩTI,1・・6,p.8(1941〕.
$ubgenoモype:Do)3aCi:aprovosli−i’Fairmaire,Ann・Soc・Ent・France,6,V,Bull.pp.LXIV−LXV(18$う), fromChina〔Designated by S.H.Chen,Sinensia,ⅩlI,1−6,p.8(ユ941)〕.
RarTge‡OrientalRegion;Manchurianand AuざtrO・Malayan Subregions.
Gene帽篭C旭goacteps:Body comparatively small(5.0一9・Omm.in1ength),叩indle・
Shaped,generallyyellow−browntoblackishbrownorblack,Witha$1ightmetallicdlimm13r・
Pronotum subquadrate,nOt distinctly punctured,rather smooth,Very finely transversely
Wrhlkled oralutaceous,錘hro鱒,With a distinct centrallongitud玩algroove.Scutelhlm
Obtrigonate・Elytra rather broad,but very muchlonger than broad,flat,SmOOthorvery
finely wr・inkled,glabrous,StrOngly andI・egularly punctate・・StIiate;1elytralepiple11r&nOt
dcfined.pubescenceofpropleuradistribu七edoverthewhoIcsurfacee草CePtn&rrOWarCa這
atthe side and apex.Mesosternum transverse”Pygidium narrowedand roundcdat the
apex“Hind・−femora armed with a distinct denticle on alittle bcfore the &PCX Of each
lower side and also a denticulated keelbetween theformer denticle and apex,in some
CaSes the male aImed with another−distinQt denticle alittle heforethefor皿ergeneral
denticle.
♂:Abdomen armed with a pair’of transversely settled tubercle or carina on the
Ⅲiddle oflst鈍eIni始
TheIe aretWO SPeCies helonglngtO the present suhgenusin Formosa andtheyare
disting−uisbed as f01lows:
Key to the Spe¢ies
l小針dantennalsegmentli・−ユitimesaslongasthe2ndone,SuPra・・antennalareaestr。nglydeve・
loped;合:hind−femoraa叩edwithonlyageneraldenticleoneachlowersidebeforethea匹ⅩWuCh
isusua11y followed by one to three minute teethor ser工ula亡ions・・・・・I・l・”‥・・”・・・∴provosliiFairmaire
l,.3rdantennalsegmentonlyslightlylonger than,OIabou亡aSlongas,the餌d one,Supra−antennal
areaenotstronglydevel瞭ed;舎:1血d・femoraarmedwithanotherdistふctdenticlea批tlebefore
thegeneIa7preapical、one払ndmoreoverwithsomefineserTulation$betweenthem・・・:ZbwiSch6nfeldtl.Dona¢富a(Cypね¢pa$ter)ppovo$蛸 FAIRMA王RE
DonaciaProvo8tiiFairmaire,Ann.Soc.Ent.France,(6),Ⅴ,Bull・P.LXIV・LXV(ユ885)(China:Peking). DonaciaProvostiJacobsohn,HoraeS6c.Ent・Rossicae,ⅩⅩⅤⅠ,p・420(1S92)(China=Peking&Yang一 tse−・kiang).・・JacQbyet Clavareau,in\Vyt$man,Gen・Ins・,ⅩⅩⅠ,Col・Phytoph.=Donaeitb,P.7(1904) (Peking).−Clavareau,inJunket Schenkling,CoIo Cat.,u,Chrysom・1・4,p・24(1913)(Peking).小Winkler,Cat.Col.Reg・・Palaearcticae,Ir,10,P”12B7(1929〕(N・China)・
Donaciap7OUO?liChqi6,Transl・Nat・fIis亡・Soc・Formosa,ⅩⅩIV,ユ3さ,pp・52さ&528・r529(1934)(Japan
proper…NagasakiIHiogoITowadaIWakayama〉 Mt・Kirisima・yama&Mt・・Hanaoka−yama;Formo
Lake Zitugetu・tan;China:Hangchowin Chekiang)・→Ho,in Wu,Cat・Ins・Sinesium,m,p・778
(19即)(China=Peiping).・Chtijb Trans”Nah His仁・Soc.Formosa,ⅩⅩⅩ,204,p・350(194q)(Korea: Nan,y6riinSui壷en−Gun,Keiki一工場);Trans・Shikok:1Ent・So9・,ⅠⅠ,f;,p・31(1951)(J叩ah=Matsuyama
Donacia(CTyE)hqgaste)・)P)OWStiGoecke,Kol・Rundschau,ⅩⅩ,6,pp.:望17‘&318−22l,ざpho:03”(19f;ま) (E・Siberia:Nikolaiewsk;JapanpIOPer:Tokio,Kiot?&Hiogo;Korea‥S6ul&Butshitshan;ManchuIia享 China=Peking,JentschoufuinSchantung&Ts云hin−kiangalongJang−t2;e・Ikiang,Kwei−Tschou;Indo−China Hanoi&L去n立−Son;Fo=mOsa);Arch・f・Hydrobiol・・Suppl・二Bd・ⅩlV(Tropische BinnengeやSer・ Bd・ⅤⅠ),P”印面5)(Nicolaiewsk,Tokio,Kioto?Hiogo・S妬Butshitshan・Jentschoufu・Tschin−Kian dlongJang・tZel−Kian虚・Kwei・・Tschou,Hanoi,Lang・Son&Formo3ユ);Ent.Bla:1t.,ⅩⅩⅩⅠⅠ,5;pp.221& 2芳8(1936);1・・C仙,ⅩⅩⅩVT,2,p・(1940)(FoImO3a). Donacia((軸血鮮血)provosliChen,Sinensia,ⅩII,1・16,pp.1詫&1か15(1941)〔China). Do}Wia bri液戒巌∵Weise,Arch・i’・NaturI女・,LXrV,i,2,p.・177(1S9日)(Amur:Nicola3efsk).・Jacoby etClavar甲u,inWytpman,Gen・Ins・,ⅩXI,Col・・Phytoph・‥Donacidde,p・7(190ま)(Amour)・−CIavarea早I
inJunketScl−enkling,Col.Cat・,LI,Chrysom.1−4,p・18(ユ913)(Amur)・−Winkle工Cat.Col.Reg.
Palaea工∝icチe,ⅠⅠ,ユ0,p・・ユ227(’享929)(AmuI). Donaoianilidicollis Welse,Arch・f.Natu工g.,LXTV,i,2,PP.177−I178(Ⅰ898)(China:Ch;nkiang)..− Jacc・byetClavat・eau,inWyt8man,Gen・Ins・,ⅩⅩⅠ,Col・Phytoph.:Donacidae,p..7(1904)(China:Chang ・Yang)・・・Clav?reau,inJunke己Schenkling,C(〉l・Cat・,LI,Chrysom.1・4,p.23(1913)(China:Chang1− Yang)叶・Winkler,Cat・Col・Reg・Palaearcticae,IT,10,pU1227(ユ929)(China).・fIo,inWu,Cat.Ins. Sinensium,IrI,P小777(1937)(Chiムa:Changyang)ルBodynarIqWly elongate,Slightly rounded on each$ide,narrOWed at boththe anterior
一ウ Fig・9.・〟舶用茄(q炒 タブ・0糊J定一壱■Fai工maiIe,♀・ Fig・8・・加′雄払(q膵ち伊如㌢)ダ1淵助成 Fairmaire,■舎・ andposteriorcnds,ratherflatonthe dor凱1m;headalldundersideofhody,tOgetherwith
legs,Verythickly cover・edwithfinesilverywhite pubescence一・Head bronzyblack,With ■aslidltCuPreOuSShimmer,pOSt・−OCParareawithtwotrIanSVer9elysettledyellowishわrown
of m?n4ibles piceous tob享ack・)and undcrside of head yt31lowish hr・0wn Or reddish
hrown,theselight colorations having a strongtendency todevelope andin$Ome CaSeS(
theYareOCCuPylngノnearlywholesurfaceofheadwiththeexceptlO110finter−andpost・qOCular
areae;dnten鱒e yellowish brown or reddish broⅥ1in general,and sometitnes they aIe
entirely so,hutin many cases with doTSalsurface oflst
Segment anddorso1−apicalparts
OfthefollowlngSCuPreOuSbro一
nzyorblackishhl・oWntO hlack,
an(l these daIk coIorations
having a strong tendency to
develope and in some cases
they are QCCuPylng nearly
tbe wllOle surface or th8
Wbole(−f the doISalsllrf8C?,
with the exception of each
basal extr’emity, yenOWisb
brown。PronotumIっ1ackor dark
hr’OWn,With a dcep golden
gr.een Or CupreCuS gI■eetlOr
CuPrIeOuS Or Violaceo−CuPrCOuS
Sheen.ScutellBm hmnzy or
greepishbronzy.勒tmye1lowisb
brown,With a slight cupreous
shimmer andsuturIalandlater&l
arcaemoreorless tingedwith
metallic針een,SOmetimecntir−
Fig・10ヱわ柁αCまα((触ろ尊朗抽つp7り0糊gき■Fai‡maiIe; Aedeagus ANAedeagus(hteralaspect):ml・−mdianlobe,mOqmedian Oliiice,mf−median foramen,dp・・distalpiece,rp・rlng piece,dp▲basalpiece. T−Tegmen(dorsalaspect,with basalpie9eIemOVed); dp−distalpiece,rP・・Ilngpiece・ MpApicalpart of medianlobe(ventralaspect).ely metallic green with a cupreous shimmer・Underside yellowish brOWn tO dar・k br・OWn
Orblackish and generallythe median arld posterior areaelighter;1egs yellowish brown,
dorsalsurfaces of femora morc orless stained with bronz.y or cupreousl)IOnZy,and
dorsalsurfaces of tibiae andtarsimoIe OrlesstirlgedWith blackishbrown・
Head wellexposed,finely and thickly punctured;interocular spaccwithastong Centrallougitudinalgroove,aPairof well−markedlateralgroove and some veIylarge
setigeIOuS PunCtureS On eaCh side;SuPra−antennalaTeaC distincily raised;front maIgin
Of clypeus gently emarginate;1abrum tr’ansverse,rOundcd at thc front corners,donvex
and fumishedwitha transverse row of setlgerOuS PunCturcSOnthe dorsum;genae Very
thickly pubescent,thepubescence differentin characterto those on thedorsum of head
hutcompletely same to those on the tlndeISideofbody;POSトgenaland gular aIeae
glahrousandsmooth Antennaefiliform,longerthanhalfthelength ofbodyinbothsexes,
finely pubescent;1st segment str・Onglythickened,CylindIicd and slightly curved,2nd
】
shortest,3rd aboutl⊥−Ilニtimesaslongasthe2nd,4thlongest,5thlonger・than6th, コ
thelatterandthefollowlngSnearlyequalinlengthtoeachotherinsomecases,hutinother
infh)nt,rath占rbroaderthanlon悠,frontandbよsalmarginsdistinctlyorratherindistinttly
borderedよnd′gently sinuate,anthe cor・nerSSbmewhattuheICulateand fumishedwith a
Setlg叩OtlS PunCture On CaCh,1ateralmarglnS nOt bordered・;dorsum gently tr・ansverselY
Cbnvek;品tnearlyflatonthemiddle,finelytransver由1ywrinkledandpunctulate,thewri−
nklesso嵐ewhatvolticatedoneachsidebe払rethemiddle,distinctlygrOOVedonthemiddle
fromalitflebehindthefrontmargintonearthehasalmarg1nandtheg・0{)Vedeepened
乱tthepoit飢ibrhalfthantheハanterior・halfい Scutellumobtriang由r,finely punctulate
andpubescent∴Elytra stron納r elongate,muChbIOaderatbasethanpronotum,gently
narrowed pds食eriorly with a−slight curvature,t‡unCated at the apex,thickly pubescent
On由chb瓦、5alcomer(neaTLlyinvisibleinthedorsalview);dorsum rather flat,Slightly
depr・eさSed・hehindthe sc一鵡11um,rIegular・1y deeply punctatein elevenlongitudinalrow$
(including a、畠hort scutellalandan external marginalrow)on each elytron,interstices
Ofthose rotvs very fi(1ely but notlthickly punctulate and transvcISely wrinkled on each
Side;hume吏i\raised and free from the distinct punctures.Underside very thickly
Clothed∴扉ithlSjlvery white pubcsccnce and spaISelymiⅩedwithmuchlongerPubcscence
Of the same colbur;1st ahdhminalsternite、distinctlylongerthan the 払ur follow!ng$
togtther;Pygidium well飢pOSed beyond>the ape=Ofelytra;hind−・fernora armedwith
a distihdt dbnticle alittlc before the apex ofeachlower$ide andinT11any CaSeS Orle
OrmO[efi’ne;dehticles bpyondtheformer gcneraldcnticle,andhind−tibiae gentlybut
distinctly sintiate.
♂:Comparedwiththefemalehodynarrowerandflatter,andlateralcurvatureofelytra
feebユer;antennae Veryml−Chlon汚erthan halfthelength ofbody;1st abdominalsternite
armedwith apalrOftr・anSVerSely$ettledelongatetLlbercle(1nthemiddle,hindrnargln
oflastabdominalsternite widely hutshortlyproducedand depressedonthesurface;
hindmarglnOfpygidiumwidぞ1ytruncated;hind・femorareachingtotheapexwhentumed
them ba(:k
♀:An†ennaealittlelongerthanhalfthelengthofbody;1stahdominalstemite without
SuChatubercleasinthemaIe;hind鱒rglnOflast abdomindlsternitegentlyroundcd
andratherangulatedatitsmjddle;最ndmargin ofpygidium producedangularly;hind・
femoranotrcachingtOthe apexof elylraWhen turnedthem hack
Lenがh…合5小q−7・・5mm・,・♀6・・0−9・Omm・
F00dqpIant:01・yJaSa偽a Linn.(Japanese name;hw)and Brasenia及h21ebe)・iGmel・(Jap・name:JuTa’
β心). Loe.D‡str.−Lake Zitugetu−tan,TaityB・Sy怠(1杏、2.X.1929,R.Takahasllileて・);Taihoku−Si, Taihoku−Sya(1舎,23。Ⅴ..19:;9,K.End61egい). Lake Ziとuge犯一ねn(af七er入4一Ch卑j6,.1934)・ Gen.Disb・.−E.Siberia,Manc−ュuria、China,Tndo・Chi:1a,Korea,Japan(HQユSさ1蔑,Shikoku&Kyむya) and FoImO3a.
むOna¢冨a(Cy紳○卵S息針)寝鼎扇 SoH6NFELDT
D。,WiaL。nZiSch8nfeld亡,Ent.Nacl−r.,ⅩtV,3,Pp..=;3・:!4(1$S8)(Japan:Hiogo);Ca仁.t(一1.Tapan,
7Nachtr.(Jahb・Nass…Ver・・NatりⅩLI),P・49(188S)(Japan:Hiogo).・Jacobsohn,HoraeSoc・・Ent・ Rossicae,ⅩⅩⅤⅠ,p.419(1892)(Japan:Hiogo);ユ.c巾,ⅩⅩⅤIf†,p・2紹(1894)(Japan)・−Tacoby et clavareau,in Wytsman,Gen.Tns・.XXて,Col.Phytoph・:DoTWidae,p・7,pl・i,f.1(1904)(Japan:Hiogo).・Clavareau,inJuqketSchqnkling,Col…Cat・,u,Chrysom・l・4,p・52月(ユ913)りapan:Hiogo)・−
1準inkler,Cat.Col.Reg.PalaearCticae,Ⅰ‡,10,p巾i21打(1929)(Jal)an)・ DonαCialend L,wis,Entomolog。,XXV‡,360,・pl1開(1893)(Japan=Kob6,Kioto&Osaka)”・Dc>i, D8butugakuZassi,ⅩⅩⅩⅠⅩ,466,PP・3芳4&3fi5(1鍋7)(Korea)・−C叫6,Trans“Nat‖Hist・Soc・Forrnosa・ⅩⅩTV,135,pp”525&5野・528(1934)▲(Japan:Iked虫&Hanaokayama)・→Ho・inWu,Ca主・王ns・Sinensium,
1II,p.777(1937)忙hina:Soocbow)・−・Ch坤,T王an3いShikokuEnt・Soc・,ⅠⅠ,3,P・31(1951)(Japan:
Muya・−Ch6,inTokusbima・ken,Shikらku)・伽ふα〔勒錘卿如うん壷Goecke,Kol・・Rtlnds琉au,ⅩⅩ,6,pp・幻8&225・胡7,5pho伽・(19舶)
(Jaba。:Tokio&Hiogo;Korea:、S611;N・−E・China‥Chin・kianq alongJa疲毒e・kiang);Archl・flHy血・ObioしSuppl.−Bd岬ⅩfV(TI0がsche Binnengew互−SeI,Bd・ⅤⅠ),pP・鯛0&291(ユ9舎各)(Tokio,
Hiogo,Sる11&Tschin・Kiang alonglang−tZe−kiang);Ent・B如t・,ⅩⅩⅩII,5,pp・男宏望&228(19fS6)
(NorthernpaTtfrom250inE・Asia);1仙Q・,ⅩⅩⅩⅤⅠ,2,p・(1940)(Philippines)・DoIWia(qyphqgaRter)Len弓iChen,Sinensia,Ⅹtr,ト6,pp・】3&:ほ・16(1941)(China)・
Donacihcwarit8(necBaly,1865)Baly〉Trans・Ent.Soc・London,p・69(1873)(Japan:Nag左saki&
Hiogo).−Lewis,Cat.Col・・J叩aneSe ATC埴elago、P.27(18791(丁−pan)…−J:aC)t,YこPrOC・Zool・Soc・
もondo叫p..75琶り885)(Japa−け・−Kolbe,如Ch・f÷ルNatuIg・,LTt,p.226(i鋸汚)(KoI・ea‥ S61ユi)・・ Sch6rlfeld†,Cat.Col..TaDan,P.142(18S7)(’Japan‥Nag軌ki&注iogo)・−Lewis,Entomoiog、,ⅩⅩVI, 鞘0,p・153(1893)(†0ヱ)0花αei■αゐ那言Sch6nfせnldt). Donaoia mepkeniWei亡一e,Arch・f.Naturg”,LXtV,i,2,pP・・178・179(189S)(China:Chinkiang)・−Jacoby ctc1年Vareau,inWytsman,Genl・Ins・,ⅩⅩT,Col・Phytoph”:Donaeidae,p・7(1904)(China:Chang・IYang)・ 一Clavareau,inJunket Scヽenklinq,Col.Cit.,LT,CllrySOm・1・・4,P・28(1913)(China:ChaTlg・Yang; Philippines:Luzon).−Schultze,PhilippineJoIZrnl・Sqi・D,Xt,r)・83(1916)(Philippines:Luzon&Tarlac)・ ∴廿■inkler,Cat.Col・・R甲・Pala占arcticae,lT,10,P・1227(1929)(China)・−Ho,inWu,Cat”‡ns・Sinensium, J汀,p.778(1937)(China:Changya昭)・ 刀0棚つぬ.卸側別個Ch鋸6,TIans・Naヒ.Hist・・SocひFoTmOSa,ⅩⅩTV,1箔,Pp・524&5鯛・5飢 り934) (FoImOSa:LakeZgetひ榊)・・・Goecke,Ent・B臨t・,ⅩⅩⅩVT,2,p・(刀㈹郁払・β′−椚09α)(1940)(蝕↓mOSa)Bodynarrowly6longate,narTOWed atboth anteriorandposteriorends)ratherflat on
the dor・sum;he五d and underside ofbody,tOgetherwithlegs,Very thiぐkly cover・edwich
fineβilverywhitepubescence小Headbronzyhlack;POSt1−・OCulararIeawithtwotransversely
settledreddishhrownmarkingswhichar℃frIequentlyhsed≦natransverseband,inter・OCular
spacewithametal1icviolaceousshimmeroneachsideofthecentrallongitudinalgr00Ve
inmanycases;mOuth−PartS dark yellow娩hr・own,1abrum tingedwitb bror)ZyOn the
basp,labium(ex坤tedthepalpi)andmargina†and apiぐalareaeof mandibles piceous
tobleck,POSトgenae and卯1a pICeOuStOblackorIbronzyhlack;antermaeinonecase
yellowishbrownand巴Cnerallytheapicalareaofeach oneofsomehasalsegments more
orlessinfuscated,hutin another casethelstsegment nearly enti;elybronzy&ndall
oftheremainingps moreorless tinged with bronzyorhlackish brown on each叩ical
乳reaいPronotumandelvtra metallic greenish−Orhluish・−OrPurPlish−aeneOuS・・Scutellum
bronzy or greenishbronzy・Uuderside上rOwnishtohlackish aTldalmostusuallythe thorax
yellowishbIOWn,dorsal aI二eae Of f61hora
(exceptedtheirbases)tingedwithbronzy、Or
Violaceous bronzy and apices.of tibiae and
tarsi more or lessinfuscated:
f‡eadw占11exposed,Vetyfinelysilverywhite Puhescent,thicklypunctured,Sparselyfovcolate− PunPtatealongtheinnermarglnOfeacheyeand
thosepuncturesincludingalong setaineach
Centre;interotularspacewithadistinctcentra1
longitudinal groove and a psir of well-marked lateralgIり0Ve;Sup工か・antennalaleae叩tVery
distinetlyraisedandcle姦rly separated t)eaCh
Otherbythe centrallongitudinalgI00Ve;front
ma曙inofclypeusgentlyeh−arginate;1abnlm
transverseJrOunded at both anterior corners,
COnVeXand clothedwithlonger pubescence
On the dorsum・Antennae filifbrm)mOreOr
lesslongerthan half thelength of body,
finely pubescent;1st seglT]ent StrOngly
thickened,CylindIicaland slightly curセed,
Fig・11・刀0朋¢£α(q伊ろqグα幽う如2言
Sch6n托1dt,合.
2nd shoIteSt,3rd very slightユylonger than the
2nd,4thratherlongerchanl÷timesofthe3rd,
5thlongest and about twice aslong as the3rd,
thesixfollowingssubequalir)1engthtooneanother,
each oneof them slightlylonger than,Or nearlv ノ
equi−・1engthto,the4th,andllthbluntlypointed
atthe apex・Pronotum neaIly quadrate,SOmeWhat
widenedinfront,Slightly bIOad¢r thanlong;fron七
and basalmarglnS VeIy gently sinuate,allthe
coIner・S SOmeWhat tubercula・ted and fuI瓜ished with
a・long seta on each tip,1ateIalmarglnS nOt
bordered;doISum tranSVerSely’COnVCX,but neaIly
flat on themiddle,iinely transversely wrinkled
and punctulate,the wIinklc去 much thickly and
COnfusedly settledon eachsideand the punc七ur’eS
On these aIeae nOt SO dis■tinct,distinctly gIOOVed
on the middle from a little hehind the front
margin to near thebasalmargln and the groov
StrOnger at the poster’iorPOrtion than at the
anter’ior portion“Scutellum obtriangular,董inFly
elongate,muChhroaderatbase
thanpropotum,g・eTltlynarrowed
POSteriorly witha slight curv姦−
ture,finely silvery pubescent
On eaCh anter’ior corner(nearly
invi岳iblefldm’
body),七rdnぐaねd乱ttbeapexand ▲
冒tもet血Cationstr軸hoIVeT・ア
Sliglltlyr’ounded;dorsum rather
flat,With suturalarea narr.ow王y
r’aised,StrOn岳lすregulよdyp11n−
Ctateine]evenlongitudinalrows
(inchding軋Sbortさcutellar8nd an exterpalm乱Tgind oれe)on
F卓g・13・Do7Wia(OyphogasfeI・)leIidSch6nfeld[;Aedcagus (The explanation of markings:See Fig・1O).
eachelytron・intersticesofthoserowsveryfinely punctulatら andtranSV9rSely wrinkled
OneaChside or near1y whole suTface;lmmer’iraisedand fr・eeftomthedistinctpunctures.
Undersideverythickly covcredwithsilver・y white p11bescenceandspar$ely miⅩCd with
muchlongerrpubes占ence6fthe sameiolour;lst abdominalstbrnite distinctlylong・er
than thefour fol】ow】ngS COmbined;Pygidium notor hardlァexposed belyondtheapc Ofelytrain−he male,butlⅣellexposedinthe‡emde;hindイemora armedwi†h along
denticleon alit−1ebeforethe叩eXOfeaehlower side andalsoa few minutedenticles
between the fbrmerlon竿denticle and apex.
♂:Body rather narTOWer tban ofthe female;antennae muChlongerIthan half the
length ofbody;1st abdominalsterIni亡C armed withapairoftr・ansVerSely settled elongate
tubef・cle befbre the middle;1ast abdominalsternite r・ectangularly depressedalongthe
hind margln Whichis widely straightlv tr・uncated;Pygidium not oT・hardly exposed beyondtheapex ofelvtra,Withhind margln Widely str・aightlytruncated;hindl−femora
reached tothe apexofelytra whenttlrnedthem back and arImed with another distinct
denticle whichis sit・17ated at a certain distancc below the middle and slightly more
irlternalthan the generalpre−IaPicalone
♀:Antennae alittlelonger than halfthelength ofbodv;1st abdom行は1stemite
without sueh a tubercle asinthemale;1ast abdominalsternite subtriangl,1arwithlateral
maT・?1nSgentlyrIOundedandnotdepr・eSぎedonthe apicalareaasinthemale;Pygidium
distinctlyexposcdheyond thcapexofelytra,narrOWly roundedandblllntly angulated at
the apex;hind−fbmora not r・c寧Chingtothe・aPCXLOfelytra whenturned them backand
unarmcdbefbre tb?genCralpTe−apicaldenficle・
Lenがh:合6い0−7一・U mm..,♀6・0・−8・O mm・ Fooc;−PJartt:Limnathe,numi’ndibum(Linn.)Grie3b・(Japane一ename:aqgaち血);Brose7乙ir‘Schクe‡eJiGmel・ (Jap.namc:Jbn㌔托i)and勒mpちαe旬Spl・(Jap・name:助ire7}nOト3yu)・ Loc.Di$れ−・Taihoku−Si,Taihoku−SyG(】舎&1♀,15.ⅠⅩ.ユ936,M・Cha361eg′)・ LakeZituge仁u■・tan(af把M・C旭i∂,193隻)・ Ge【て.Distr.−JapaLl(Honsyti,Shikoku&KyB3ya),Kor・ea,FoImOsa,PhilippinesandN・:El・Chiqa・摘 要 基湾産金花轟科(尾嵐網−・鞠菱塑目)の形態分所学的研究 ⅠⅠ 根喰金花轟詣科