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(2) S.. CAM(ラ. し.. iJ. Ⅰ 1 mm トーーーー-. ＼琶. A. ち. ∫. T'へ-. -.I.￣'-確. /. ＼. ＼. i:'T;is Hl. 藩. ン. 4/. / /. i. ㌔ l. Fig,. l･. Bathycuma. cephalothorax pace. with. 〟. ,: ,<:. ”. rotundiiectorum 5.9mm.. about. 丘ve. free. of antennule,. portion Ⅰ,丘rst maxilliped;. thoracic enlarged;. ∫,second. sp. A,. anterior. segments, E,. antenna;. maxilliped.. holotype. nov･,. portion dorsal F,. immature. of body, view;. mandible;. C,. length. male,. lateral. antennule;. G, maxillule;. B,. view; D,. of cara-. distal. H, maxilla. ;.
(3) New. DescrlptlOn.. The. twice. long. as. the. from. the. tlp. by. phasized. concave,. acute. The slightly. more. The丘rst. aesthetascs.. second,. hairs. and. flagellum. main. the. combined;. lash,. incisor. The. maxillule. The. maxilla. The丘rst The. outer. provided. with in on. second. basis. long. to. the丘rst. as. two. side.. 丘rst. the. bears. two. long. as. as. sparse. and. The. third.. 31. and. segment,. the. of. B) is. each. the. distal. the. 1/3. A,. On. marglll. peduncular. about. is very. two. segments. second.. The. segment. of main. ac-. minute. short. and very 12 and beset with. undeveloped. boat-shaped. normally. on. setae. outer. a. row. distal. as. long. produced,. and. a. the. in. Fig.. palp.. splne than. mar琴In･. the. splneS. little. a On. the The. 1,. I and. projection). as. ten. of about. little shorter distal. shown. are. outer. times. on. 1, H.. Fig.. (including. is much hairs a. twofi1aments. maxillipeds. has. plumose. shape,. in. 2 2/3. nearly. the. on. palr. distal. than. as. segment. G) has. 1,. shown. distal angle. cylindrical splnules. as. of basis. The. together.. 1/2. (Fig. 1, E) is. antenna. (Fig.. and. is. distal. longer. shortest,. about. third. about. lobe.. first. the. carinae. is subequal. the. as. a. process.. is. length. (Fig. 2, A) Its. minute, the. or. the. on. segment. long. is the. (Fig. 1, F) is. mandible. right. 3/4. second. as. segment. and 2-segmented; The且agellum of. ocular. cylindrical, and 1 1/2 times is about. splnules. is slightly. is. the. for. rudimentary. segment. segment. third. 且agellum. The. the. nearly. the丘rst. segmented;. cessory. is. the. distance. are. prominent.. except. ridges. 丘rst. lobes. (Fig. 1,. are. plates. narrow,. a. segments. missing. lateral. as. em-. carapace. is. and Fig. 2, H). C and D) is slender,. A. margln;. side. and. for. peduncular a few with. provided. lateral. the. on. 1,. (Fig. 1, A,. antennule. damaged. except. (Fig.. The. longitudinal. two. misslng. segment. abdominal The. carapace. are. are. pleopods. length.. long. as. thoracic. lobe. for. lobe. and. Of. pseudorostral. ocular. 1/3 times. free. the. all. (Fig. 1, A) is. abdomen There. segments.. the. the. of. of the. 1. about. and. length. in front. antennal. pointed. ocular. The. elements.. The. margln. The. portion.. visual. meet. length. than. anterior. its dorsal. of. sharply. lower. or. anterolateral. without and. carapace. combined. The. its. on. downward,. 1/10 of the. The. angle. As. backward. runs. teeth･. or. is. cephalo-. dorsomedian. three-quarters. ridges. about. deptb･. The. Which. without. of. the. as. vaulted.. about. lateral. no. the length. long. splneS,. its lateral. and. splne.. small. occuplS. have. of carapace. deeply. an. little directed. the. and is measured. of. as. is entirely. of. and. 2/3. than. 1 2/3 times. and. row. lobe. ocular. small serrations in shape triangular and. long. damaged. male,. less. outline. double. a. bears. The. 3. Japan. from. immature. an. width,. dorsal. the. the. sides. is rather. notch. Crustaceans. is much. B). the. with. Of. The. margln.. 1, A,. as. side,. is provided. carina. is. holotype. (Fig.. carapace. from. seen. The. Cumacean. The length of abdomen･ of cephalothorax portion integument The is moderately calci丘ed.. posterior 5.9mm.. thorax,. Deep-Sea. the. of. the. merus. maxilliped. distal segments. inner the. exceeding distal margln.. merus,. third. remaining On. ∫.. distal of. end. The. and. ischium. and. furnished. is. little shorter. a. margln.. merus,. with. is. a. few. than. the.
(4) 生. S. CAM(ラ. carpus,. bears. and. than shorter Thefirst. two. the. small. carpus,. and. its. On. splneS. outer. longer. slightly. angle.. than. the. ′rhe propodus dactylus. its distal. is. damaged (Fig. 2, B) peraeopod and The basis is misslng. slender, cylindrical, moderately There is a row that of the third maxilliped. of short the. middle. is. a. little. are. segments. curved, and shorter than On and long stout splneS. portion.. ,=. r. I.. ∫-FT=. /,( √､/i:. E. .1ユ ＼. i ヽ.. )､r. し七. ?.ド. (/ ･//. フ:. 'れ. I. ー＼二･一:I. ll ′つ′. LAへ -メ/ ㌔/バ′ ^. ミ或. 1. ′′ノ. 〟. /A/. ノ. C. 】/4/ ′. ′1'/. ′. ′. ′/. I''/. ヽ､. I. ^. /ノ. ∧ Jl. A. I. ･＼ .1iへ.＼l. ､ヽ. 〟. /1. /∧. P. ^. 【 Il 〟. 岬1. ,. /TJ I AJ′い. /∼ ′､. I ./I;i. A. ′1′Lイ. ･1/. ∼ % % ∫ ^LI 1. 〟. 〟. A// ',/ ド,. B. ｢/ー. (. JT/. '4J4. l｢. /. T｢ /?. ナ｢. イィ. //. I-^. f′′. 'ク '7. /I. ′一. /. rp. 117. -. /. // 'r. /. /. ､′′′/】. ^. /･. づ. ′. 〟. /,､. /.. /;′′. 7′ノ. (ノ･7･ク. ′_′｡′. I. イィ. /チ //. I,メ. /1. ′.,1/. r. ′. /ち /7. イ. /I. ･7. 7fj Llソr;. 二/プ^/･. 1. -:/,)f･三-. イ/. /”. /1/〃プ. -7-7 ._丁. 7′￣. ￣. -I-} I.丁. J7-1. ･￣プ. ｢. ′_ウ. ･4/タJ≦. JJ7 .-･サ. -7 ?7 ′･ヽ. pJl. 一￣7. 一一■■ワ. 一っ. 一っ. ｢. ￣′. G. ｢((:I J< ｢工｢I JL,/丁￣/ 'J(i-く∠フ i,I. 宿 Fig.. 2.. Bathycuma. maxilliped; E-F, left mentary丘rst. B,. (E). sp. rotunditectorum basis offirst peraeopod; and (F) of fourth right. pleopod.. nov.,. C,. H. holotype second. peraeopods;. immature. peraeopod; G,丘ftb. male.. D,. A,. third. third peraeopod;. peraeopod;. H,. rudi-.
(5) New. ′rhe second. basis. the. remainlng. than. shorter. setose. rather long. long. as. the. as. splneS. ′rhe. third. and. carpus. long. a. and. 1 1/2. more. than. together.. The. fourth. one. the. basal. The. that. the. to. teristic B.. the. of. from. 1975), Sagami 1227m. Japan,. lndian. 4040m. Ocean,. The. segments･. described. two. are. as. dactylus. is. lateral. short. with. the. the basis. second;. distal on. segments. side, the. either. only. the right with in Fig･. and is. 2, G･. shown. as. exopods･. segments. abdominal. but it is referable two. is closely. to. abdominal. and. uropods. Bathycuma. the genus. have. segments. charac-. Fage, in. (Day, 1975), new. species. by. species. having. is easily. that. the. the. on. carinae from. distinguished. 1913;. Day,. Mediterranean?. Jones, 1969. magnum. lateral. the. dorsal. the. B･. Calman･. (Stebbi咽,. 1967), and. (Gama,. 1951), and. longicaudatum. deep. 1174-1218m. lOOOm. about. B･. to. related. California,. 1915;. (Stephensen,. is about. misslng･. the丘rst. nov･. Diego,. Bay,. are. segments. The. remaining damaged. developed. posterior. genus,. and. sp.. o庁San. a. much. genus.. rotunditectorum. 1912. tO. asslgn. the. peraeopod. are. the. than. remalnlng,. well. uropods of. cephalothoracic. features. have. and. absence. it difBcult. makes. by. The. segments. and. merus. provided. shorter as. The丘托h. peraepods. segment. telsonic. Remarks.. four. shape,. distal. むst. of. splneS,. (Fig. 2, E, F). basis･. of Of. palrS. much long. as. peraeopod. portion. four. The丘rst. times. the proximal. only. with. one. D) is. in. propodus･. and. aplCal. plumose. 2,. the. as. basis. the. The. short.. combined,. short. (Fig.. peraeopod. long. as. propodus. three. and. is slightly left. is 3 times. than. is cylindrical The together.. is very. ischium. Japan. longer. much. peraeopod. The. which. carpus,. is. distal segments. splnOSe.. and. the. as. second. of. from. Crustaceans. (Fig. 2, C). peraeopod. The. peraeopod.. Cumacean. Deep-Sea. the. from. the. abdominal. above-mentioned. pro丘1e. of. the. carapace. to. the. fact. that. is. entirely. vaulted.. The 丘1e of. specific the. carapace. urotunditectorumn. name. is very. like. a. vaulted. Family Makrokylindrus. refers. the. dorsal. prod. roof･. Diastylidae. (Coalescuma). jubatus. sp･. nov･. (Figs. 3-6) Makrokylindrus as. M.. (Coalescuma). sp.. Material Allotype oostegites;. abdomen;. depth. 1650m,. 254,fig.. 1, D-F;. subadult. female. developlng with 2 females with. Day,. 1980:. 244. (in key,. GAMO.. Holotype. examined.. subadult 2 manca. terior. 1971:. Gama,. sp.,. male; larvae. St. T-13 far off north. 2 females. marsuplum;. with. of. developlng. damaged,. 1 adult male, poswithout 125oOl･0′E, 125oOO.8′E126oO3.8′N,. (females) and (KH-68-2),. OOSteglteS;. 26oO3.8′N,. Miyako-jima. i., Ry缶ky5. Is･ ; 25. May,. 1968･. Gear:. Beam-trawl. DescrlptlOn･. Holotype. is. a. subadult. female. with. developlng. OOSteglteS,. length.
(6) S.. CAM(ラ. ( tl.i(. J4^イ. 〟. .IJィ. pT<. >. J1-A一ニ ',1. ･}=' -■】. .∫ >. > >. 1 ′. ∴;,I-:書 >. -A..',? -･･'.i ′ こJLー. ,ツヽ. L. i--､J. 〟一一.Vl▲. -. >＋､. I.J:㌔表. f<. 『＼. ンノ. l. ＼. ざミ?. ∼. メ ＼ ニユー. I/. ∼vレ.. v. ー+聖ー. A.ち. C. D Y V V V V. V. I. ′. ′. Fig･. 3･. Makrokylindrus. (Coalescuma). anterior portlOn E, antenna; F, segments 1ateral. with view.. Of. right telson. holotype jubatus sp･ nov･･ (including telson)･. ll･7mm about body, dorsal view;. developing?ostegites,. C-D,. G, maxillule; mandible; left uropod and ; ∫,last. antennule. H, two. and. maxilla; abdominal. fen?le. subadult. A, its. lateral distal. Ⅰ, last segments. two. with B,. γlew-;. portion. (D). abdominal with. telson,. ;.
(7) New. ll.7mm. about covered. 1. about. 1/2. runnlng. do. dorsal dorsal. antennal izontally. distance. Of. about. 1/2 of. fused. together.. The of the. is subequal. The. the. segment,. the. vided. a. ten. The. palrS. The. a. is. ment. a. their. to. is nearly segment. second. 1/3. 2/3. than. 1/3. or. A,. (Fig. 3,. about. in length;. subequal. fourth,. the. as. 1/2. than. more. is. B). dorsally. are. much. long. as. large. which. long. as. the. is. as. the. aslthe longest,. the. second･. 2/3. and. are. third. the. long. as. segments of. The. main. lash,. about. 3 times. long. seg-. third. second,. is 5-. 1､he. as. long is. which. subequal･. minute･. consists as. and. second,. the. the. segment･. and. in. mainflagellum two beset with. as. very. side;. furnished. cylindrical the. half;. pro-. splneS･. and than. distal. peduncle,. about. two. 1/3. is about. is. as. aplCal. slender. the. On. and. portion,. little longer. a. splneS as. is. distal. the. is. Of. long,. seg-. each. on. splnules. long. abdominal. abdominal itself･ and. telson. pre-anal. palr. rather. long. long. as. twenty. a. two. the. as. the. as. and. D) is. as. long. the sixth. as. last. of the. as. long. as. long. as. of about. splnules. segment. length. 1/4. rows. of several. third;. the. combined. 1um. of. lateral. the丘rst. aesthetascs; subequal. about. 3/4. segment. nearly. segmented,. is about. peduncular. less. length. combined. is. row. little. the. C,. Brst. with. provided. segments. 2 2/3 times. (Fig. 3, A,. antennule. shape.. meet. and. anterolateral. segments. and are. 1 1/3 times. about. 1ongitudinal. portion. about. fourth. and. cylindrical,. four. about. portion Of. palr. post-anal. with. to. pre-anal. with. hor-. almost. lobe. The. The. B･. and. are. the. splneS,. sixth.. subequal. and. ments;. arst. (Fig. 3, Ⅰ,∫)is. telson. third. spinose,. longest,. is the. to. The. length･. segment. the丘fth. 1, A. segments. thoracic. The. length･. (Fig. 3, A) is. body. total. large. ocular. length･. midOn. splneS･. lobes. the. low. a. several. eyeless. round. free. the. all. of. carapace. abdomen. by. Fig･. in. pseudorostral. carapace. total. are. ridges but. carapace,. stout. with. there shown. the. of. the. as. wide. as. transverse. of. pair. set. closely. as. than. interrupted. are. of carapace,. front. the. a. encircling. and. are. a. bodylength,. total. more. is SplnOSe･. length. the. line. position The sinuated･. carapace. ′rhe combined. and. calcified. the. as. little. are. almost. and. ridges. in. is. there. regular. 1/4 of. about. margln. side. portion. a. forward. projected. lower. the. anterior in. lobe,. mid-dorsal. is shallowly. notch. a. the. arranged. are. moderately. long. as. which. width,. each. encircling. of the. surface. which. for. The. groove･. the. 1/3. nearly. frontal. the. in. meet. not. B) is. as. the. from. upward. ridges. long. as. Just behind. depth.. the. (Fig. 3, A,. times. is. integument. The. ■/. Japan. splnules.. minute. carapace. from. Crustaceans. Cumacean. (including telson)･. with. The. Deep-Sea. and. accessory且ageト. of. 3. the. first. segments; or. the. the third. One.. The. antenna. The. mandible. incisor The. in. Fig.. (Fig. 3, F) is. normal. shown. 3, E. type. and. 14. bears. setae. on. process.. maxillule. The丘rst. and. and second. maxilla. are. maxillipeds. as. in Fig･. shown are. shown. 3, G. in Fig･. and H･ 4, A and. B･. the. right.
(8) S. GAM6. llJ',. ∫. ㌔. D. Q't. ㌧､. rJI. Jt^^ 1. 1. 忌. :こ･. 1鞍三｢ 4･..1,::,;,, ll.･1,. I. ･:1.. I. ■. l. g 0. ノ/. > I. /. ノ. コ. E A ∧. . ♂♂ L4 `イ. ▼. 11イ. 4. 4. ♂. ● .114',. イ. ノ1. d d く丁. イー// ...;. C. 1 -I. 3,1.>ア1r 'モー■. Fig･. 4･. Makrokylindrus. 丘rst ma∑illiped; B, E, second PeraeOPOd;. (Coalescuma) second. jubatus. maxilliped; FIG, peraeopod;. sp.. C,. ･毛｢. nov.,. third. thir･d (F). to. holotype. maxilliped; fifth (H). female.. subadult D,. basis. peraeopods.. of. A,. 丘rst.
(9) New. together,. and. tion reaches. provided the. the. as. propodus,. long. as. The. basis. curved,. and. are. segments. Only. basis. an. long. splneS. that. the. Brst. the. carpus,. is. basal. the. as. long. that. The. long. splneS. the. lateral. providedwith long. the. The is about. lateral. the. On. with. a. than. as. 6/7. few. The. long. the. as. is. long. as. the. is about. the. than. more. peduncle. provided with is 3-segmented, twice. 1/4. inner. marglr).. with. spinules. little less. a. as. long. as. furnished. 2/3. of. 1/2. than. as. as. long. 1/2 as. splneS the. as. the. long. distal. short, long. long. as. a. and. as. the. On. small two. the丘rst. of. On. splneS. splneS. small. portion. bears. and. 1 1/3 times. with. stout. the carpus,. as. stout. Of. palr. about. basal. On. two. lateral. the. splne Of. palrS. and The丘fth the. and. of the. furnished. and. The. margln.. is. about. fourth. 3/4. as. (Fig. 4, H) peraeopod basis is slightly more. distal. segments.. long. as. the basis. second;. its basis. remaining. the. the. together,. segments. third,. half. than. shorter. I) is about. I,. distal as. a. and. fourth,. of the. the. 1/3. as. of exopods. much distal. the. (Fig. 3, On. than. together.. as. length. very. about. of splnules. is. segments. uropod. about. than. long. splneS. eight. F). is. iscbium. the. near. rows. of small. distal. 2/3 of the combined. than. spines. slender, furnished. and. remaining. rows. remaining. and. peduncle. 4,. (Fig.. longitudinal. the. 1/3. merus On. The. segments.. 2/3. The. ll.5. about. than. is. ischium. stout. margln. splnules. longitudinal. peraeopod as. few. (Fig. 4, G) is shorter. little. The. less. a. third. peraeopod long. a. and distal margln.. has. some. distal. propodus, and The dactylus. margin.. length. more. margin.. two. as. inner. inner. the. on. as. argln. peraeopods. dista】. remaining. marsupium,. The. The and distal margln. is cylindrical, slightly more. merus. inner. the. as. is 1 1/3. (Fig. 4, E) is spinose, rather slender, peraeopod distal segments together, remaining and provided with. the. as. basalm. on. 2/3. about. propodus. The. maxilliped.. complete with female, and. together.. provided with distal end. The is long carpus dactylus propodus and combined,. outer. several. moderately cylindrical, It is measured margln.. lateral. with. holotype. 4/5. splneS. the. bears. of second. The. spines.. the. on. propodus.. the. On. spinose,. third. peraeopods. of the. of the. portion. basis. as. The. margln.. spines. and is cylindrical,. margin.. the. specimen. and. about. the. as. of. paratype. to. similar. carpus. on. is. B), has. the. as. ′rhe. and. stout. On. with. carpus,. carpus. (Fig. 4, D) is. distal segments remaining bears a and stout spine. merus,. 2/3. female. adult. is very. the. as. as. the. distal projecinner. the. provided. segments. misslng.. (Fig. 6, A,. mm. with long. as. than. inner. the. on. its outer near. ex-. slightly. distal. remaining. and. The. curved,. splnules. merus,. 9. Japan. margln;. with. the. margin.. of first peraeopod furnished. 1 1/7 times. about. distal. inner. little shorter. a. the. as. the. as. from. C) is moderately. beset. and. and spinulose dactylus.. the. as. is. and. On. long. as. merus. inner. the. on. 3/4. The. margln･. times. as. end. is about. ′rhe ischium. spines. of ischium,. 4, long. as. splneS. with. Crustaceans. (Fig.. of third maxilliped distally, about 1 3/4 times. panded. long. Cumacean. basis. The. inner. Deep-Sea. inner. the. as. The. margln.. nrst. peduncle;. the. segments. together,. segment. and. telson,. and. endopod. is. furnished. a. little with.
(10) 10. S. CAM(ラ. ∼: i-.:. A. B. ? √ヽ. ll I ) ( /. E. 9 1J. H. F. I. 笥. ー. Ⅰ. G. 71ク1 0 〇. D. ←.⊥空』 TA;B. C. 芋t;iLJ^i .こフ. つolo9; (■ノ. ∫. K. L. Fig.. 5.. Makrokylindrus. about. ll.3mm. dorsal. view;. C, telson,. portion. of. H,丘rst. peraeopod;. (Coalescuma) jubatus sp. (including telson). A, lateral last. two. abdominal. dorsal. view;. I, second. segments. E, antennule;. peraeopod;. nov.,. view;. with F,. JIL, third. allotype B, telson. young. anterior and. antenna;. (J) to丘fth. portion uropods;. G,. length. male,. third. of D,. body,. distal. maxilliped;. (L) peraeopods..
(11) New. Deep-Sea. Cumacean. Crustaceans. from. ll. Japan. 1. G. mm. G. r去BA･L;-:.:_. lby. ー. く. ○. ･G. -i. , AqJ( -. F' L. ′′+ ∠ ′▼. 七′･一山L. A′一. ▲. E. F. @. .:/二￣ユ. I.Ip.i. A. i. mrn. l■---. A′C.D. B. d 〟. でさ. I. ′L.. ′ ′. .クJe′. ■′-( ′. /-1L /. L(･. ￣ノ∼′.′′. C. 4J. ′二_二二:二. ▲1.■. ㌔. Fig.. 6.. Makrokylindrus. male. with. with. marsupium,. portion pleopods.. of. length length. body, G,. manca. jubatus. (Coalescuma). marsupium,. carapace larva. about. about length. (A) D-F,. 13.5mm. about. (female),. nov.,. sp.. ll.5mm 4.3mm about. paratype ;. B,丘rst. subadult. specimens. peraeopod. male. (D)‥E-F,first. 6.3mm.. without. (E). and. AIB,. C,. fe-. female. posterior. second. (F).
(12) 冨12. s. GAM6. SplneS. On. third,. and. SeVenteen. than. the. bas is. three lateral. the. four. young. holotype. female. antennule. is. are. shown. Bay,. has. an. later. Banda. no. is. exopod. setose. on. the. (Calman,. by. a. 1913:. Day,. stout. with. anterior. has. two. encircling. the. of. of stout. new. Vemakylindrus. that. the. carapace. carapace. surface The. and. the. new. species. is also. of Bali,. 780m. deep.. How-. stout. with is derived. a. of stout. absence. the. ridges. like. from. is most species Sulawesi, 1158m. new. but. splneS,. species. splneS. but. (C.) jubatus, much the. 1980) in. carapace. are. surface and G,. it is distinguished. of Bone,. Gulf. mid-dorsal groove･ 1969 from off south. name. cluster. The. from. a. Jones,. species. speci丘c. the. utinomii) and. by. 1905). set. closely. M,. in Fig. 6,. variations. Gama, 1968. which in. long. as. surface･. (Stebbing,. On. cinctus. has. carapace. D.. carapace. 2789m. splneS. (C･) The. carapace.. anterior. ridge. later. in. utinomii. palrS. However,. male.. sexual. and. from. aspects,. braod. four. shown. allotype. growth Diastylis. as. as. of. pleopods. The丘rst. are. the. segment. rudimentary. anterior carapace B; Fig･ 6, A, C, D. to. SOme. of. and. 5, A,. like. 1n. long. as. enterrupted. M･. peduncular. pair. to. similar. the. On. related. very. deep,. cingulatus. stout. not. to. ever,. segment. and. second. female. Fig･. (about 2/3. Sea,. encircling. allied. The. is very. specimens. spines. B;. be is. broad. (C.). stout. may. splnulose. M･. has. ridge. apex･. (Fig. 5). segments･. paratype. holotype. 3, A,. lOOOm. as. the. deei), and. of. Fig･. carapace. 2/3. to. related. third. of endopod,. A. abdominal. other. the. species. about. its broader. SplneS. the. This. than. to. splnations. Remarks･. On. in. two. the. similar distributions. different. less. segment. the. at. little shorter. a. the. margln;. splneS. ll.3mm. about. the丘rst. of. also. as. observed. by. second. length. male,. each. different. Sagami. the. two. in. on. they. these. and. of. end. have exopods. peraeopods In general appearance,. some. margln. is. segment. inner. the. On. The appearance･ general little dilated distally (Fig. 5, E).. a. present. and. splneS. second. marglnS.. Allotype. are. the. the. margln;. inner. the. On. splneS. bears. little exceeding. a. inner. the. of. encircling. Oll. splneS. from. the. fact. the. the that. mane.. sp.. grandidentatus. nov.. (Figs.7,8) Makrokylindrus. (in. key,. (B), Gama, 1971)).. (Vemakylindrus) A sp. (Gamo,. 197l:. sp.. V.. as. Material. female; Holotype examined. 125oOO･8'E-26oO3.8'N, 125oO1.0'E, depth 1650m, Ry缶ky色Is.. ; 25. May,. DescrlptlOn. integument. The. The much depth･. Gear. Holotype is rather. carapace more. 1968.. than. It is covered. is. (Fig. 7, A, 3 times with. as. length. female,. a. moderately. B) long. splnules,. St.. Day,. 2, C,D;. T-13. (KH-68-2),. far off north. 1980:. 26oO3.8′N,. Miyako-jima. of. 239. I.,. Beam-trawl.. :. thin,. 254,fig.. is nearly as. the. and. 7.6mm. about. calci丘ed. 2/5 width,. provided. of. telson).. splnOSe.. and the. (including. total. which. is. with. nine. a. body. little palrS. length, more. Of large. and. than. the. splneS.
(13) New. Deep-Sea. Cumacean. Crustaceans. from. 13. Japa□. /-2T?-∴恵戸:. zi<ノ<,j<),.. <. D. =. 【フ,′_･. .. I. r/. -去ご'itz<). ど. .. ∠l一○. .a'一己-. 上クLi). メ. A. C. ′レ. I B. ･-ら. ノ /. H. Fig･. 』 ←.....ー+. Vemakylindrus. 7･. (including 1ast. in. is slightly. bent. 1ateral. lower. rrhe. 1/2. of. length. the. length･. has. The丘fth. The. splneS. itself, and apical Th?. 7,. body. length.. base.. furnished. spines,. 2/3. which. antennule. portion. The with is 1/7. times. palr. Of. post-anal two as. (Eig. 7, A,. D). large. for. meet. and. splneS.. (Fig. 7, A,. segments. is. B). is the. segment. almost. shortest and in length, splnOSe,. subequal. splneS. cylindrical,. four a. as rows. as. long. of. longer. as. the. is. rather. are. about. than. the. and. about. in subequal fourth. The. the丘fth. as. lateral. last. the. lateral. is about. Of small. spinose,. segments. little. long. portion. palrS long. as. horizon-. their distal portion concave. ′rhe antero-. with. slender,. longest,. than. a. F,. antenna;. almost. lobe,. ocular. The丘rst are. The丘rst. has. C,. view;. length;. thoracic. Of large. is the less. eyeless. is shallowly. is rather. A). segment. preanal. the. near. (Fig･. E,. splnOSe,. notch. segments. palrS. are. carapace. free. (Fig･ 7, C) is 2 1/2. telson. The. meat.. two. lobes. total. length.. four. 7.6mm. about. dorsal. portion D, antennule; uropods;. is furnished. the. carapace. is slightly. segment. sixth. total. posterior. total. antennal. of. length. of body,. anterior. and. the. of. all. female,. maxilla.. Of carapace. margln. The. the. The. outward･. segment each The abdomen. and. H,. 1/2. than. more. of. splnulose･. with. G, maxillule;. combined. 1/4. about. telson. ′rhe pseudorostral circles･ forward in front of the round. much. or. B,. view;. holotype. nov･,. sp･. lateral. four. projected. distance. A,. segment. mandible;. arranged. a. telson).. abdominal. right. tally. grandideniatus. splneS. 1/4. as. splneS. abdominal. and. long. and. the. as a. segdorsal. two. palr. telson Of. long. telson. large. and. cylindrical･. The. first.
(14) 14. S. GAM6. combined, as. long. bears. and. the. as. flagellum. is. segment. peduncular. three. third. longer. much stout. than. lateral. one,. and little longer. a. a. distal. two. the. second. splne. near. the. than. the. spines;. has. is 3-segmented,. the. peduncular segment. distal end.. distal. segments. is about 2/3 The main segment;. peduncular. C B. A. ////T. I/T/. I;, //. ＼/. D ノ ′ ′. 7ノ. ′ ′. I,∼ ,I////. //'二[/'T ノ. ld. 7只. // /∫. ∫/. 〟+. E. 逮.ら;. h u. cb. F. H. ♂ U. β く7b. -･b. ∠'. ＼ し+. ＼ ＼. G. Fig.. 8.. Vemakylindrus. offirst maxilliped; E, second peraeopod;. grandidentatus B,. second F-H,. nov.,. sp.. maxillipeds; third. (F). to. C, fifth. holotype. female.. third maxilliped;. (H). peraeopods.. A,. distal. D,first. portion. peraeopod;.
(15) New. 丘rst. the. the. as. 1/4. about. the. as. 1/4. about. shortest,. the. as. long. is. The. antenna. The. than. four. the. lash, and. main. the. as. the. The. aesthetascs.. third. the丘rst. is about. which. 1/2. is. segment. is. accessoryflagellum. 3-segmented;. second,. 15. Japan. second;. mandible. in Fig. 7, E. shown is 7, type, (Fig. F) normal. is the. segment. 1 1/2. times. long. as. as. has. and. 12. the. right. 7, G and H. in Fig. 8, A. and. B.. in-. on. setae. process.. The. and. maxillule The丘rst and The. basis. 1. than vided. a. with. second. times. The. merus. is cylindrical,. podus. a little slender, rather distal segments together, and. merus. The. a. stout. 1 1/2 times. are. as. has. the. the. as. outer. On. in length･ The. pro-. as. long. 3/4. and. pro-. splne. angle･. carpus,. more. projection. stout. a. subequal. about. On. splne long. distal. outer. ischium. carpus. and. have. each. its. angle;. ischium.. of the. about. is. remaining inner distal. ischium,. carpus. and. shown. (Fig. 8, C). the. portion. middle The angle.. also. the. as On. splne. in Fig.. shown are. maxillipeds. long. about. inner. as. maxilliped. as. stout. are. maxilla. of third. 1/3. reaches. as. as. from. Crustaceans. third.. cisor. the. longer. bears. and. second, long. as. Cumacean. is slightly. segment. long. as. Deep-Sea. dactylus.. the. The. basis. much. less. beset. with. of丘rst. 2/3. than. splnules･. is about The. measured. about. ischium. long. lateral. stout. (Fig. 8, D) is rather. peraeopod as. 1/2. long. as. the. as. splneS. bears. large. a. and the. as. splneS. 1/3 of the. carpus,. which. peraeopod. (Fig. 8, E) is. angle to. subequal. and The. angle･ few. a. with. inner. is about. inner. the. provided. the. near. together,. segments On. splne. and. merus,. two. merus. distal. remaining. curved,. moderately. slender,. lateral. the. propodus. is. and. its length. or. dactylus. The. second. peraeopod.. distal. mainlng. ischium. is about. angle･. The. to. the. is. 1/3. together, long. as. has. merus. of. and. the. as. wiuh. and. bears. is about. and. 1/3. large. a. of. the. 2/3. than. 2/3. about. beset. merus,. lateral. stout. merus. on On. splne. is about. which. re-. The. splneS･. spine. terminal. carpus,. the. as. lateral. stout. asthefirst. long. as. stout a. long. as. the. inner. side,. each. subequal. dactylus.. or. propodus. a. laterally,. little dilated. a. segments. length. the. and. basis. The. less. much. (Fig. 8, F) is about 6/7 as long as the second, and about peraeopod long as as distal as long fourth; basis is the remalnlng segas the nearly the but shorter (Fig･ 8, G) is much the basis of fourth ments peraeopod together, The. than. is. a. on one. also. third. the little. the. basis. splne. beset The. provided segmented,. third. of. a. with. and. and. tw･o. much. uropod. small. The. the. The丘fth is. There. fourth.. while. that. of. third. and. is about. 7/8. peraeopods,. ischium. (Fig･ 8, H). peraeopod a. spines. rowofsome. of the丘fth fourth. has. only is. peraeopods. splne.. (Fig. 7, C). spines. shorter. as. fourth. lateral. sigle of. peduncle. long. as. distal end･. the. near. with. 2/3. than. more. of distal segments.. length. combined. On. than. the the. inner exopod;. distal the. as. long. margln･. as. The. first segment. the. telson,. endopod. is about. and is 31. l/2.
(16) 16. S. CAM(ラ. β. ′′. ft ･′.〆J. cP. B. 1皿 A,B ■. l. I. A. P. q一寸1. V. ∇. 〟. i). ? ?. ?. ∀ ?. 3, ,?.. (9. >. E>. >. .,. し ー. o J. 9. 4. I.>b. 0. ●. 1v. H 9･. Vemakylindrus. lO･3mm view;. C,. antennule; mentary VleW.. texture F, pleopods. sp･. oxycanthus. (including. telson)I. of. carapace. antenna;. (p),. A,. surface;. G,丘rst ventral. two view;. holotype. nov･,. lateral. view; D,. immature B,. anterior. antennule;. abdominal. H,first. segments to. of. E, enlarged with. fifth abdominal. d■. I. t> i. J.. length. male,. portion. ユ. t>(. JL->-. 4. ∼. Fig･. >. C7. 414. body,. about dorsal. distal portion two. pairs. segments,. of. of rudidorsal.
(17) New. times. long. as. two. bears. inner. the. on. 1/2. than. two. long. the. at. furnished. with. with. than. has is. exopod. lateral. two. longer. segment. The. apex･. beset. and. distal. the. margln;. splne. peduncle,. 17. splne. little. a. setae. the. a. and. less long. a. seta.. Remarks･ nov･. the. long. Japan. combined, and is segment slightly. second. inner. the. from. segments. the. On a. Crustaceans. two. margln;. and. as. Cumacean. unequal. splneS. margln. as. terminal. inner. the. On. third, and. distal. the. as. splneS. Deep-Sea. a. at. General but. glance,. described. species. of. former. is easily. the the. of. like V･ oxycanthus species is very sp. distinguished from later and the other has characteristic large that the carapace this. appearance. by. genus. splneS. in four circles･ arranged fact that the carapace bears. The. the. the. name. speci丘c. large. characteristic. Vemakylindrus. the. of. species. splneS. refers. mentioned. above.. nov.. sp.. oxycanthus. new. (Figs. 9, 10) Makrokylindrus. (in. (Vamakylindrus). key,. V.. as. A. sp.. Material examined･ 142o46･3/E-38o52･1/N, districts. eastern. The. telson)･. length,. wide. as. 血e. hairs･. forward. The. total. The. are. plates. Each. of the nrst The丘fth segment. and. provided. of as a. telson. six long pair. The Brst. splneS as. the. of. long. prominent. (Fig･ 9, A, two. in. the. telson.. apical. the. north-. (including. mm. covered. and. with. is. notch strongly. cylindrical, pair. projected. distance concave.. serrated.. (Fig. 9, A, is the. segment. 1/2. and. sparse. a. is shallowly. segments. The丘rst. with. for. meet. and. is. thoracic. beset. than. more. horizontally. almost. antennal. carapace. thetotal. as. slightly. and. lobe,. ocular. long. as. of rudimentary. of. length.. (Fig. 9,. pleopods. a. shortest.. body. the. is. B). A,. G).. longest.. (Fig. 9, Ⅰ)is a. long,. very. free. a. 4/10. than which. The. is. 10.3. calci丘ed,. the. length.. has. segments. is the. with. H). 38o51.2'N,. (KH167-2),. about. width, is splnulose surface. Of. carapace. length. more. eyeless. all the. of. 1980.A 238. (Fig. 9, C).. are. margln. 1/3 of the. than. as. length.. carapace. length. abdomen. The. long. triangular. lower. or. combined. side. as. pseudorostral the. Day,. Beam-trawl.. moderately. is slightly. B). lobes. of. 2, A,B;. far offKamaishi,. m,. Gear:. male,. thin,. carapace. The. little less. A,. St. 4. 127011280 1967.. spinules. with. 3 times. 254,fig.. male;. immature. an. The. in front. The. is. (Fig･ 9,. depth･. about 1/2 of the The anterolateral. depth. is rather. patterns. nearly the. as. immature. 12 August,. integument. minute reticulated The carapace body. Holotype. Holotype. 197l:. 1971)).. 142o44･7/E,. of Japan;. Description･. Gama,. (A),. sp.. (Gamo,. small. about. serrated. of the. middle The. posトanal. spines,. peduncle of antennule longer is much segment. about. as. crest. on. long. twice. each. dorsum.. 1/4. the last abdominal. side. The. long. as. palrS. the. the. distal. two. base. portion Of lateral. segment,. and. a. is about splneS. row. 2/3 and. telson.. (Fig. 9, A, D, E) in robust than. the. near. pre-anal. has丘ve. portion as. as. segments. and. club-shaped;. combined,. and. the beset.
(18) S.. 18. GAM6. G. /. く//+. ∼. i. I'讃. D. 蛸. ･二毒. ど. ＼. ＼-⊥ミ. H O r■J. 詔コ. O r. 0. ∫. I(⊃. P. K. 0. 0√￣. ○. ＼. Fig.. 10.. Vemakylindrus. oxycanthus. B,. C,. mandible; G, third. maxillule;. maxilliped;. (∫) and丘fth. (K). H,丘rst peraeopods.. sp.. maxilla; peraeopod;. holotype. nov.,. DIE,first. V. immature. maxilliped; I, basis of second. F,. male. second. peraeopod;. A,. right. maxilliped;. ∫-K, third.
(19) New. four. with. the. as. spines;. second. the. distal. the. as. two. peduncular 1/2 as long. is about. is. long. tw.ice which The accessory且agellum as. 3 segments;. the. is about. second. The且agellum. 1/3. minute. of. bears. slender,. long. has. twice. about. two. short. aesthetascs.. is very. short. and. not. third,. aesthetascs. composed. of. the. first;. long. as. the. the. as. four. lash, and. main. about. segment. 4-segmented;. as. segment. the. as. longest,. and. 2/3. long. as. third. and. and. and. distal. long. as. (Fig. 9, E). antenna. combined,. second,. the. is the. segment. is very. distal segment. the. as. the. long. is rather. 2/3. about. distally;. spines. segments. 19. Japan. is short,. segment. fourth;. the. as. from. Crustaceans. and. first segment. the. Cumacean. three small provided with bell-shaped. The main且agellum. third,. long. large. lateral. is somewhat as. Deep-Sea. as. fully deveト. yet. oped. type (Fig. 10, A) is normal mandible and furnished with incisor right process. The are as in Fig. 10, B and C. maxillule and maxilla shown Thefirst are in Fig. 10, D-F. and second shown maxillipeds. 13. The the. The. basis. of third. 1 5/6 times. about a. with distal. long. as. remaining along the inner. row. splneS of several its margln; external angle. little longer merus. has. longer. than. than. splneS. a. the. merus,. outer. margln.. 2/3. about. distal margln.. the. a. as. long. The. beset. and. On splne and a large distally. The ischium. margin. long. splne. The. carpus. distal. the. near. is. a. The. end.. little dilated. a. the is. laterally,. several and beset with propodus, longer is cylindrical, than the and. the. as. curved,. togethei,. segments. little produced. a. bears. and. merus,. On. splne. the. On. the. is. moderately. slender,. distal. the. as. is. (Fig. 10, G). maxilliped. on. setae. propodus. dactylus.. The as. basis. long. lateral. The. in three. splneS. as臣1ong. (Fig. 10, peraeopod distal remalnlng segments. arranged ischium,. the. as. is about. carpus. ′1､he second The. merus. splne. 5/6. about. large. provided with is about. a. inner. the. On. and. propodus. basis. is somewhat. is. peraeopod. a. palrS. and. laterally. dilated. The. bases. and. is. distal. twice angle.. little longer. than. provided with have exopods.. of. a. distal segments.. without. little. and. the. On. outer. are. as. of the. that. fourth. long. margln.. about The. telson,. and. that. as. The. together.. segments. long. than. shorter. and about fourth peraeopods. splneS. as. nearly. and. 1 1/2 times. third. of third row. and. splnOSe,. (Fig. 10, J). peraeopod. the remalnlng little less than the. 丘fth (Fig. 10, K). in length. (Fig. 10, Ⅰ)is damaged. peraeopod. peraeopod. of the丘rst T､he basis of third ㌔ second and shoter than. as. the. subequal first four. Of peraeopods The. to. long. a. the. as. as. and. The. rows.. with. provided long. 1 1/3 times. cylindrical,. together,. longitudinal. and. elongate,. dactylus.. the. six. H) is. of丘rst. the. as. peduncle. splneS. the. segments. On. exopod. the. of. splnulose. in length;. combined;. (Fig. 9, Ⅰ)is. uropod. the. inner the second. The. margln.. first segment segment. long. as. a. is 3-segmented,. endopod. is about is. the. as. long. as. little longer. as. than. the the. beset. with. subequal distal third;. two. there.
(20) 20. S. GAM6. two. are丘ve, the. lateral. one. segments,. three long. as. and. the. as. SPlneS. Remarks.. The. margln,. two. above-mentioned is almost horizontally. type. most. related Colombia, off. of the genus, off Costa-Rica,. species. Trench,. deep,. 3718m. 2470m. deep.. However,. to. 912. m. V., gladiger deep, V.. two. the. the. front. that. of. the. (B孟cescu,. costaricans. (Jones, 1969) from have. described. three. by. (B孟cescu, 1961), the. species. new. derived. in. outer. apex.. characterized. forward. Of. 1/2. the. On. at the. splneS. V. Prolatus. and. margln. is about. exopod splneS. are. projected. On from the carapace of splneS ornamentation name is The new speci丘c of the species is covered carapace splneS. with sharp. 摘. long. species. closely. from. 1961) from Kermadec. are. new. The four. with. peduncle, provided the distal end, and two. inner. splnulose. spine.. apical. and. near. the. pseudorostrum They carapace.. one. the. On. present. and. respectively,. uropodal. two. splneS. from. di庁erent. species.. the. fact. that. the. 要. 東京大学海洋研究所の｢白鳳丸+によって行われた日本海溝および付近海域の研究航海(KH-672)で,. 1967年8月に釜石沖の水深1,270-1,280m の深海底よりビーム･トロールで採集された Vemakylindrus属(Diastylidae科)の1種と,東シナ海および隣接海域の研究航海(KH-68-2). で,. 1968年8月に宮古島北方沖,水深1,650mから得られたBathycuma属(Bodotriidae科), Vemaklindrus属(Diastylidae科)にそれぞれ属すると考えられる3種. Makrokylindrus属と. は,研究の結果,次に述べる新種として記載される｡ 1.. Bathycuma. rotunditectorum. nov.. sp.. :宮古島北方沖(1,650m深)産の未成熟雄(腹部 1匹を完模式標本として記載される｡本種に最も近縁と考 深)から記載され,地中海(1,227深) えられる種類に,カリフォルニア沖(1,174-1,218m 後部を欠く,頭胸部の長さ約5.9mm) (STEPHENSEN, longirostris 1969. (CAM(ラ,. 1915),および相模湾(約1,000m深) CALMAN,. 1912. 1967)からも記録された B.. と,インド洋(4,040m深)より記載された. ? B.. JoNES,. magnum. があるが,本新種の背甲を側面よりみると,背甲背縁が全長にわたり完全なアーチ状となっ. ていることなどで,上述の2既知種と明らかに異なっている｡ 2.. Makrokylindrus. (Coalescuma) jubatus. sp.. nov.. :宮古島北方沖(1,650m深)から採集 された｡本新種は完模式標本に東成体の雌(体長11.7mm)杏,別模式標本に未成熟雄(体長7■.4 1亜成体の雄(体後部を欠く)と. mm)を指定,他に副模式標本の4雌, よって記載される｡本新種は第3, 1961,. 4自在胸節が融合しており,. 2. manca幼生(雌)に. Coalescuma亜属(BÅcESCU,. 1962)に属する｡本種に最も近縁と考えられる種類には,スラウニシ(セレべス)のボネ湾. (Gulf. Bone)の1,158m深と,バンダ海の2,789m深より記載されたM.. of. (CALMAN, がある｡. M.. (C.) cingulatus. M. (C.) cinctus JoNES, 1905)と,バ1)島南方沖の780m深より記載された (C.) cingulatusでは,本新種同様に,背甲中程を横断し輪状に配列する棟1列を有. Ⅰ969. M.. するが,背甲前部に大きな嫌がないこと,. (C.) cinctusでは,背甲を横断し輪状に配列する妹 GAM6 (1971)および 2列を有することなどで,明らかに本新種と区別されるo本新種は以前に,. (1980)ノにより〟. (C.). DAY. も Vemakylindrus. sp.. (GAMO, sp.. grandidentatus. 1971)とされていたものである｡ nov.. :本種はDAY. V. sp.. (1980)により. (B) (GAMO,. 1971)とされていたものであり,宮古島北方沖(1,650m深)から採集された1匹の雌(体長7.6 mm)を完模式標本として記載される.本種ほ一見次に記載する種に類似しているが,本新種の背 甲には細かい棟の他に,背甲を横断する様にほづ4列に配列する大林を有するなどで,次の種と区 別される｡ 4. Vemakylindrus. oxycanthus. sp･. nov･. :本種ほDAY. (1980)により. V･. sp･. (A). (GAMO,.
(21) Deep-Sea. New. Crustaceans. Cumacean. from. 21. Japan. 1971)とされたものであり,釜石の沖,水深1,270-1,280の所より採集された1匹の未成熟雌(体長 V･. 10.3mm)を完模式標本として記載されるo上述の. grandidentatusの他に,本種に近縁と考 V. gladiator (BÅcESCU･ 1961),中米. ぇられるものに,南米のコロンビア沖(912m深)産の V･. のコスタリカ沖(3,718m深)産の. JoNES,. V. Prolatus. 演(2,470m深)産の. (BÅcESCU,. costalicans. 1961)および･ケルマデック海. 1969などがあるが,本新種の背甲上の棟の配置は上. 記3既知種の何れとも異なっており,明かに種を異にしている｡. References BÅcESCU,. M.,. lindrus. Acad.. icaine). M.,. BÅcESCU,. (crustacea Lamont. CALMAN,. :. South. Ann.. S. Air.. L.,. FACE, GAM6,. Crustacea. Seto. S., 1971.. Japan.. du. N.S.,. JoNES,. 200 STEBBING,. STEPIIENSEN,. Raport Zool.. 1969.. meters.. T.R.R., K.,. au. des. cours. Abyssal. Stebbing du. campagnes 1･. Crustacea･. Vema. on. Cumacea. order. U.S.. natl.. Family. l･. Part. Musリ66(9). :. the. (Crustacea, Malacostraca) 133-163; 15(4) : 245-274;. 16(3). Farance.. 54:. 15(2). :. sur. 80:. quatre. of. the. Vaun-. Diastylidae･. 1-136.. especes. Cumac6s. of Cumacea. Tierreich,. 39:ト210･. Danish. Oceanogr･. Rep･. de. des. Part. Japan･. of. :. 147-192･. eaux. profondes. depths. exceeding. 2511255･. systematics and distribution Rep., 10: 99-180. Galathea Cumacea･. Subfamily. Bodotriidae,. and. The. 1915･. the collection 6031676･. 177-220･. 4･ Families. Cumacea. biol, Lab‥. Cumacea･. :. Gynodiastylidae. Part. de. preliminaire Mag. Tokyo,. 1913･. in 41. Mus”. 36: i-23･. Siboga-Exped･･. Expedition.. 187-292.. Faune. Studies mar.. :. the. of Proc.. Cumacea･. 82(6). Mug.,. Siboga. ヽ. CAM(ラ,. Makrokylindrus. genre. Vol･. York･. of the. Cumacea･. S. Afr.. African. S., 1967-68. publ.. New. Museum.. Cumac占s.. 1951.. 1-3.,. The. African. South. J., 1980.. Cumacea. Ann.. thompsoniinae.. DAY,. The. National. States. du. recueillies. 209-223.. 1905.. J‥ 1975.. de. (C6t･e Amer-. Pacifique. 325-333.. :. nouvelles. Observatory. W.Tリ1912. united. DAY,. 1. W.T.,. 6(3). Vemaky-. sous-genre du. tropicales. eaux. connaissance. Esp占ces. cumacea)･. GeologlCal Ser.,. Res. CALMAN,. des. Biol.,. Revue. R.S.R.. Contributionえ1a. 1962.. Makrokylindrus. deg. nouvelles. especes. (crustac占s, Cumac6s). sg.. n.. ヽ. Deux. 1961.. Exped･･. from. 1908110･. D･L:. 29-34･.
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